2016 In Archosaur Paleontology
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This archosaur paleontology list records new
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
archosauriform Archosauriformes (Greek for 'ruling lizards', and Latin for 'form') is a clade of diapsid reptiles encompassing archosaurs and some of their close relatives. It was defined by Jacques Gauthier (1994) as the clade stemming from the last common anc ...
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
that were described during the year 2016, as well as notes other significant Archosaur paleontology discoveries and events which occurred during the year.


Basal archosauriforms


Basal archosauriform research

* A study on the
resting metabolic rate Resting metabolic rate (RMR) refers to whole-body mammal (or other vertebrate) metabolism during a time period of strict and steady ''resting conditions'' that are defined by a combination of assumptions of physiological homeostasis and List of typ ...
of 14 taxa of fossil
archosauromorph Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) than to lepidosaurs (such as tuataras, lizards, a ...
reptiles as indicated by bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
is published by Legendre ''et al.'' (2016). * A study of the phylogenetic relationships of the archosauriforms traditionally assigned to the family
Euparkeriidae Euparkeriidae is an extinct family of small carnivorous archosauriforms which lived from the Early Triassic to the Middle Triassic (Anisian). While most other early archosauriforms walked on four limbs, euparkeriids were probably facultative bi ...
is published by Sookias (2016). * A redescription of the braincase and the inner ear of ''
Euparkeria ''Euparkeria'' (; meaning "Parker's good animal", named in honor of W. K. Parker) is an Extinction, extinct genus of archosauriform reptile from the Triassic of South Africa. ''Euparkeria'' is close to the ancestry of Archosauria, the reptile gro ...
capensis'' is published by Sobral ''et al.'' (2016). * A study of the phylogenetic relationships of
archosauromorph Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) than to lepidosaurs (such as tuataras, lizards, a ...
reptiles, with an emphasis on the phylogenetic relationships of proterosuchids and
erythrosuchids Erythrosuchidae (meaning "red crocodiles" in Greek) are a family of large basal archosauriform carnivores that lived from the later Early Triassic (Olenekian) to the early Middle Triassic (Anisian). Naming The family Erythrosuchidae was named by ...
, is published by Ezcurra (2016). * A study on the patterns of morphological diversity of the skulls of late Permian to Early Jurassic archosauromorph reptiles is published by Foth ''et al.'' (2016). * A study on the braincase anatomy of the
type specimens In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes t ...
of ''
Pseudochampsa ''Pseudochampsa'' is an extinct genus of proterochampsid archosauriform known from the Late Triassic (Carnian) Cancha de Bochas Member of the Ischigualasto Formation of San Juan Province, Ischigualasto-Villa Unión Basin in northwestern Argentin ...
ischigualastensis'' and ''
Tropidosuchus ''Tropidosuchus'' is an extinct genus of carnivorous archosauriforms from the Middle Triassic period (Anisian to Ladinian stage). It is a proterochampsid which lived in what is now Argentina. It is known from the holotype PVL 4601, which consis ...
romeri'' is published by Trotteyn & Paulina-Carabajal (2016). * A reevaluation of the
neotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...
specimen of ''
Parasuchus ''Parasuchus'' is an extinct genus of basal phytosaur known from the Late Triassic (late Carnian to early Norian stage) of Andhra Pradesh and Madhya Pradesh, India. At its most restricted definition, ''Parasuchus'' contains a single species, '' ...
hislopi'' and a study of the phylogenetic relationships of the species is published by Kammerer ''et al.'' (2016), who consider the genus ''Parasuchus'' to be a senior synonym of the genera ''
Paleorhinus ''Paleorhinus'' (Greek: ''"Old Nose"'') is an extinct genus of widespread basal phytosaur known from the Late Triassic (late Carnian stage). The genus was named in 1904 based on the type species ''Paleorhinus bransoni'', which is known from Wyo ...
'' and '' Arganarhinus'', and refer the species ''Paleorhinus bransoni'' Williston (1904), ''
Francosuchus ''Francosuchus'' is a dubious genus of probably basal phytosaur known from the Late Triassic (late Carnian stage) of Bavaria, southern Germany. It was named by Oskar Kuhn in 1933 and the type species is ''Francosuchus broilii''. In the same a ...
angustifrons'' Kuhn (1936) and ''Paleorhinus magnoculus'' Dutuit (1977) to the genus ''Parasuchus''. * A study on the endocranial anatomy (including the brain, inner ear, neurovascular structures and sinus systems) of ''Parasuchus angustifrons'' and '' Ebrachosuchus neukami'' is published by Lautenschlager & Butler (2016).


Pseudosuchians


Research

* A study of the skull anatomy of the ornithosuchid ''
Riojasuchus ''Riojasuchus'' is an extinct genus of ornithosuchid archosaur from the Late Triassic (Norian) of Argentina. Ornithosuchidae was a widespread family of facultatively bipedal pseudosuchians (crocodilian-line archosaurs) with adaptations for scav ...
tenuisceps'' is published by von Baczko & Desojo (2016). * A restudy of ''
Dasygnathoides ''Dasygnathoides'' is an extinct genus of pseudosuchian from the Late Triassic (Carnian) Lossiemouth Sandstone of Scotland. Classification "''Dasygnathus''" ''longidens'' was erected by Thomas Huxley for a maxilla from the Lossiemouth Sandstone ...
longidens'' and ''
Ornithosuchus ''Ornithosuchus'' (from , "bird" and , "crocodile") is an extinct genus of pseudosuchians from the Late Triassic (Carnian) Lossiemouth Sandstone of Scotland. It was originally thought to be the ancestor to the carnosaurian dinosaurs (such as ''Al ...
woodwardi'', rejecting their synonymy, is published by von Baczko & Ezcurra (2016). * A study on the
cranial Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...
anatomy and phylogenetic relationships of the
aetosaur Aetosaurs () are heavily armored reptiles belonging to the extinct order (biology), order Aetosauria (; from Ancient Greek, Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized Omnivore, omnivorous or Herbivore, herbivor ...
''
Paratypothorax ''Paratypothorax'' is an extinct genus of aetosaur, known from a single species, ''Paratypothorax andressorum''. It was a broadly distributed member of the group found in Germany, North America, and possibly parts of Gondwana. The best specimens ...
andressorum'' is published by Schoch & Desojo (2016). * New fossil material from the
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
(
Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic ...
or earliest
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...
) Pinheiros-Chiniquá Sequence of the Santa Maria Supersequence in
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
attributed to the
rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...
n species ''
Prestosuchus ''Prestosuchus'' (meaning "Prestes crocodile") is an extinct genus of pseudosuchian in the group Loricata, which also includes ''Saurosuchus'' and ''Postosuchus''. It has historically been referred to as a "rauisuchian", and was the defining mem ...
chiniquensis'' is described by Lacerda ''et al.'' (2016). * A study on the presence, size, shape, and position of the subnarial
foramen In anatomy and osteology, a foramen (; : foramina, or foramens ; ) is an opening or enclosed gap within the dense connective tissue (bones and deep fasciae) of extant and extinct amniote animals, typically to allow passage of nerves, artery, ...
(an opening located between
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals h ...
and
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
) in ''Prestosuchus chiniquensis'' and its implication for archosaurian phylogeny is published by Roberto-da-Silva ''et al.'' (2016). * A redescription of the fossil material assignable to the species ''
Trialestes ''Trialestes'' is an extinct genus of Late Triassic (Carnian) crocodylomorphs that lived in South America. It has been classified as a dinosaur in the past due it being adapted as a terrestrial, running carnivore. It is classified in Sphenosuchi ...
romeri'' and a study of the phylogenetic relationships of the species is published by Lecuona, Ezcurra & Irmis (2016). * The
osteological Osteology () is the scientific study of bones, practiced by osteologists . A subdiscipline of anatomy, anthropology, archaeology and paleontology, osteology is the detailed study of the structure of bones, skeletal elements, teeth, microbone morp ...
description of ''
Carnufex ''Carnufex'' is an extinct genus of crocodylomorph suchian from the Late Triassic of North America. The genus was first described in 2015 by Zanno ''et al.'', who named the Binomial nomenclature, binomial ''Carnufex carolinensis'', meaning "Carol ...
carolinensis'' and a study of its phylogenetic position is published by Drymala & Zanno (2016). * A study on the changes of
crocodyliform Crocodyliformes is a clade of Crurotarsi, crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the ...
biodiversity through the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
/
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
transition and on probable causes of the decline of some crocodyliform lineages at this time is published by Tennant, Mannion & Upchurch (2016). * Description of postcranial skeletons of three specimens of the
sphagesaurid Sphagesauridae is a Gondwanan family of mesoeucrocodylians that lived during the Late Cretaceous. It was a clade of terrestrial crocodilians that evolved very mammal-like teeth and jaws.MARINHO, T.S. & CARVALHO, I.S. Revision of the Sphagesaurida ...
''
Caipirasuchus ''Caipirasuchus'' is an extinct genus of sphagesaurid notosuchians known from the Late Cretaceous of northern São Paulo State and western Minas Gerais State, southeastern Brazil. The type species, ''C. paulistanus'', was named in 2011. A seco ...
'' (representing ''Caipirasuchus montealtensis'', ''Caipirasuchus paulistanus'' and ''Caipirasuchus'' sp.) is published by Iori, Carvalho & Marinho (2016). * Description of the postcranial elements of the skeleton of ''
Pissarrachampsa ''Pissarrachampsa'' (meaning "piçarra he local name for the sandstones it was recovered fromcrocodile") is an extinct genus of baurusuchidae, baurusuchid mesoeucrocodylian from the Late Cretaceous of Brazil. It is based on a nearly complete ...
sera'' is published by Godoy ''et al.'' (2016). * Description of new cranial remains of ''
Pholidosaurus ''Pholidosaurus'' is an extinct genus of neosuchian crocodylomorph. It is the type genus of the family Pholidosauridae. Fossils have been found in northwestern Germany. The genus is known to have existed during the Berriasian-Albian stages of the ...
purbeckensis'' from the Early Cretaceous (
Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
) of France and a study of phylogenetic relationships of the species is published by Martin, Raslan-Loubatié & Mazin (2016). * Fossils of the
dyrosaurid Dyrosauridae is a family (biology), family of extinct neosuchian crocodyliforms that lived from the Campanian to the Eocene. Dyrosaurid fossils are globally distributed, having been found in Africa, Asia, Europe, North America and South America. ...
crocodylomorph Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. Extinct crocodylomorphs were considerably mor ...
''
Hyposaurus ''Hyposaurus'' is a genus of extinct marine Dyrosauridae, dyrosaurid crocodyliform. Fossils have been found in Paleocene aged rocks of the Iullemmeden Basin in West Africa, Campanian–Maastrichtian (Late Cretaceous, Late Cretaceous) Shendi Forma ...
'' are described from the Late Cretaceous
Shendi Formation The Shendi Formation is a Late Cretaceous geologic Formation (geology), formation of the Atbara-Shendi Basin in northern Sudan.Eltayib et al., 2018, p.2 Indeterminate theropod remains have been recovered from it.Weishampel, et al. (2004). "Dinosa ...
of
Sudan Sudan, officially the Republic of the Sudan, is a country in Northeast Africa. It borders the Central African Republic to the southwest, Chad to the west, Libya to the northwest, Egypt to the north, the Red Sea to the east, Eritrea and Ethiopi ...
by Salih ''et al.'' (2016). * A description of the
endocranial The endocranium in comparative anatomy is a part of the skull base in vertebrates and it represents the basal, inner part of the cranium. The term is also applied to the outer layer of the dura mater in human anatomy. Structure Structurally, t ...
anatomy of ''
Steneosaurus ''Steneosaurus'' (from , 'narrow' and , 'lizard') is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic (Callovian or early Oxfordian) of France. The genus has been used as a wastebasket taxon for thalattosuchian ...
'' is published by Brusatte ''et al.'' (2016). * A study on the body proportions and body size of teleosaurids is published by Young ''et al.'' (2016). * Fossils of
teleosauroid Teleosauroidea is an extinct superfamily of thalattosuchian crocodyliforms living from the Early Jurassic to the Early Cretaceous. It is phylogenetically defined by Mark T. Young and colleagues in 2024 in the ''PhyloCode'' as "the largest clade w ...
thalattosuchia Thalattosuchia is a clade of mostly marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not member ...
ns, including a close relative of '' Lemmysuchus'' and ''
Machimosaurus ''Machimosaurus'' is an extinct genus of machimosaurid crocodyliform from the Late Jurassic (Kimmeridgian and Tithonian) and Early Cretaceous. The type species, ''Machimosaurus hugii'', was found in Switzerland. Other fossils have been found in ...
'', are described from the
Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
(
Bathonian In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
) of
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
by Jouve ''et al.'' (2016), who name a new
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Machimosaurini Machimosauridae is an extinct family of teleosauroid thalattosuchian crocodyliforms. Text was copied from this source, which is available under Creative Commons Attribution 4.0 International License The family was first identified in 2016, w ...
. * A redescription of the holotype specimen of the
metriorhynchid Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period (Bajocian to early Aptian) of Europe, North America and South America. The name Metriorhynchidae ...
species ''"Plesiosaurus" mexicanus'' Wieland (1910) is published by Barrientos-Lara ''et al.'' (2016), who transfer the species to the genus '' Torvoneustes''. * New
goniopholidid Goniopholididae is an extinct family of moderate-sized semi-aquatic neosuchian crocodyliformes. Their bodyplan and morphology are convergent on living crocodilians. They lived across Laurasia (Asia, Europe and North America) between the Middle J ...
specimen belonging or related to the species ''
Goniopholis ''Goniopholis'' (meaning "angled scale") is an extinct genus of goniopholidid crocodyliform that lived in Europe and North America during the Late Jurassic and Early Cretaceous. Like other goniopholidids, it resembled living crocodilians, and pro ...
lucasii'' / ''
Amphicotylus ''Amphicotylus'' is an extinct genus of goniopholidid mesoeucrocodylian from the Tithonian of Colorado, Wyoming, and Oklahoma. It was described in 1878.E. D. Cope. 1878. Descriptions of new extinct Vertebrata from the Upper Tertiary and Dakota Fo ...
lucasii'', representing a single individual rather than a composite of unassociated elements, is described from the
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
(
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
,
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) by
Erickson Erickson may refer to: Places Canada * Erickson, British Columbia, an unincorporated area *Erickson, Manitoba Erickson is an unincorporated urban community in the Municipality of Clanwilliam – Erickson within the Canadian province of Manitoba ...
(2016). * A Middle Jurassic
dentary In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone ...
from the Isle of
Skye The Isle of Skye, or simply Skye, is the largest and northernmost of the major islands in the Inner Hebrides of Scotland. The island's peninsulas radiate from a mountainous hub dominated by the Cuillin, the rocky slopes of which provide some o ...
,
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
, United Kingdom, referred to ''
Theriosuchus ''Theriosuchus'' is an extinct genus of atoposaurid neosuchian from Late Jurassic to Early Cretaceous of Europe (Hungary & southern England), Southeast Asia (Thailand) and western North America (Wyoming), with fragmentary records from Middle Jur ...
'' sp., is described by Young ''et al.'' (2016). * A
histological Histology, also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
study of a specimen of ''
Susisuchus ''Susisuchus'' is an extinct genus of neosuchian mesoeucrocodylian crocodyliform from the Early Cretaceous of Brazil. Fossils have been found from the Nova Olinda Member of the Aptian-age Crato Formation in the Araripe and Lima Campos Basins of ...
anatoceps'' is published by Sayão ''et al.'' (2016). * New fossil material of ''
Allodaposuchus ''Allodaposuchus'' is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages, and possibly the Santonian stage, of the Late Cretaceous. Although generally classified as a non-cro ...
precedens'' is described from the Late Cretaceous of France by Martin ''et al.'' (2016). * Fossil
mekosuchine Mekosuchinae is an extinct clade of crocodilians from the Cenozoic of Australasia. They represented the dominant group of crocodilians in the region during most of the Cenozoic, first appearing in the fossil record in the Eocene of Australia, and ...
vertebrae, tentatively assigned to ''
Mekosuchus ''Mekosuchus'' is a genus of extinct Australasian mekosuchine crocodilian. Species of ''Mekosuchus'' were generally small-sized (less than long), terrestrial animals with short, blunt-snouted heads and strong limbs. Four species are currently ...
whitehunterensis'', are described from Riversleigh (Australia) by Stein,
Archer Archery is the sport, practice, or skill of using a bow to shoot arrows.Paterson ''Encyclopaedia of Archery'' p. 17 The word comes from the Latin ''arcus'', meaning bow. Historically, archery has been used for hunting and combat. In modern ...
& Hand (2016), who interpret them as confirming that even adult specimens of this species were smaller in snout-vent length than adults of extant small
crocodilia Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchia ...
n species belonging to the genera ''
Paleosuchus ''Paleosuchus'' is a South American genus of reptiles in the subfamily Caimaninae of the family Alligatoridae. They are the smallest members of the order Crocodilia in the Americas. Classification The genus name ''Paleosuchus'' is derived from t ...
'' and ''
Osteolaemus ''Osteolaemus'' is a genus of crocodiles. They are small, secretive crocodiles that occur in wetlands of West and Middle Africa. They are commonly known as the African dwarf crocodiles. Unlike other crocodiles, ''Osteolaemus'' are strictly noctur ...
'', and indicating that this species employed feeding behaviours that were unusual for crocodilians. * Partial skeleton of the
Chinese alligator The Chinese alligator (; ), also known as the Yangtze alligator (), China alligator, or historically the muddy dragon, is a crocodilian endemic to China. It and the American alligator (''A. mississippiensis'') are the only living species in th ...
is described from the late Pliocene of western Japan by Iijima, Takahashi & Kobayashi (2016). * A study on the
osteology Osteology () is the scientific study of bones, practiced by osteologists . A subdiscipline of anatomy, anthropology, archaeology and paleontology, osteology is the detailed study of the structure of bones, skeletal elements, teeth, microbone mo ...
of
alligator An alligator, or colloquially gator, is a large reptile in the genus ''Alligator'' of the Family (biology), family Alligatoridae in the Order (biology), order Crocodilia. The two Extant taxon, extant species are the American alligator (''A. mis ...
fossils from the late Miocene Moss Acres Racetrack locality in
Marion County, Florida Marion County is a county located in the North Central region of the U.S. state of Florida. As of the 2020 United States census, the population was 375,908. Its county seat is Ocala. Marion County comprises the Ocala, Florida Metropolitan S ...
and the phylogenetic placement of the alligators these fossils belonged to within the genus ''Alligator'' is published by Whiting, Steadman & Vliet (2016). * New information on the anatomy of ''
Globidentosuchus ''Globidentosuchus'' is an extinct genus of basal caimanine crocodylian known from the late Middle to Late Miocene of the Middle and the Upper Members of the Urumaco Formation at Urumaco, Venezuela. Its skull was very short and robust, with l ...
brachyrostris'' and ''
Centenariosuchus ''Centenariosuchus'' is an extinct genus of caimanine crocodylian known from the Miocene of the Panama Canal Zone of Panama.
gilmorei'' and a study of the phylogenetic relationships of these species is published by Hastings, Reisser & Scheyer (2016).


New taxa


Basal dinosauromorphs


Research

* Marsicano ''et al.'' (2016) date the
Chañares Formation The Chañares Formation is a Carnian-age geologic formation of the Ischigualasto-Villa Unión Basin, located in La Rioja Province, Argentina. It is characterized by drab-colored fine-grained volcaniclastic claystones, siltstones, and sandstones ...
, containing fossils of non-dinosaurian
dinosauromorphs Dinosauromorpha is a clade of avemetatarsalians (archosaurs closer to birds than to crocodilians) that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, ...
''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely k ...
'', ''
Lewisuchus ''Lewisuchus'' is a genus of archosaur that lived during the Late Triassic (early Carnian). As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs (possibly tru ...
'', ''
Marasuchus ''Marasuchus'' (meaning "Mara (mammal), Mara crocodile") is a genus of basal Dinosauriformes, dinosauriform archosaur which is possibly synonymous with ''Lagosuchus''. Both genera lived during the Late Triassic in what is now La Rioja Province, A ...
'' and '' Pseudolagosuchus'', to early
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...
(236–234 Ma), 5–10 million years younger than previously thought. On this basis the authors postulate that the origin of dinosaurs was a relatively rapid event, as the transition from vertebrate communities containing only non-dinosaurian dinosauromorphs to communities containing the first dinosaurs occurred in less than a 5-million year interval. * A study on the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
of the
femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...
and the
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of
long bone The long bones are those that are longer than they are wide. They are one of five types of bones: long, short, flat, irregular and sesamoid. Long bones, especially the femur and tibia, are subjected to most of the load during daily activities ...
s of the
silesaurid Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids compr ...
''
Asilisaurus ''Asilisaurus'' ( ); from Swahili, ''asili'' ("ancestor" or "foundation"), and Greek, (, "lizard") is an extinct genus of silesaurid archosaur. The type species is ''Asilisaurus kongwe''. ''Asilisaurus'' fossils were uncovered in the Manda Bed ...
kongwe'' is published by Griffin & Nesbitt (2016). * Basal dinosauromorph fossils including fossils of both ''
Dromomeron ''Dromomeron'' (meaning "running femur") is a genus of Lagerpetidae, lagerpetid Avemetatarsalia, avemetatarsalian which lived around 220 to 211.9 ± 0.7 million years ago. The genus contains species known from Late Triassic-age rocks of the South ...
romeri'' and ''D. gregorii'', as well as a
dinosauriform Dinosauromorpha is a clade of avemetatarsalians (archosaurs closer to birds than to crocodilians) that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, ...
fibula The fibula (: fibulae or fibulas) or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. ...
resembling the fibula of ''Marasuchus lilloensis'' but with much larger size, are described from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Color ...
of
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
, USA by Sarıgül (2016).


New taxa


Non-avian dinosaurs


Research

* An assessment of methods used to the determine the
ontogenetic Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
status of non-avian dinosaur specimens is published by Hone, Farke &
Wedel Wedel () is a town in the Pinneberg (district), district of Pinneberg, in Schleswig-Holstein, in northern Germany. It is situated on the right bank of the Elbe, approximately south of Elmshorn, and west of Hamburg. History Foundation and Mi ...
(2016). * A study of the evolutionary dynamics of
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
and extinction through time in Mesozoic dinosaurs is published by Sakamoto, Benton & Venditti (2016). * A study on the dinosaur metabolism, re-evaluating earlier studies of Werner & Griebeler (2014) and Grady ''et al.'' (2014), is published by Myhrvold (2016). * A study on the morphological similarities of the skulls of ''
Plateosaurus ''Plateosaurus'' (probably meaning "broad lizard", often mistranslated as "flat lizard") is a genus of plateosaurid dinosaur that lived during the Late Triassic period, around 214 to 204 million years ago, in what is now Central and Northern Eu ...
engelhardti'', ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fo ...
stenops'' and ''
Erlikosaurus ''Erlikosaurus'' (meaning "Erlik's lizard") is a genus of therizinosaurid that lived in Asia during the Late Cretaceous period. The fossils, a skull and some post-cranial fragments, were found in the Bayan Shireh Formation of Mongolia in 1972, da ...
andrewsi'', their feeding mechanics and behaviour is published by Lautenschlager ''et al.'' (2016). * A study testing for a correlation between the presence of bony
cranial Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...
ornaments and large body size in non-avian
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaurs is published by Gates, Organ & Zanno (2016). * A description of
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
teeth from the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
of Northern
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
and a study of their phylogenetic relationships is published by Gerke & Wings (2016). * A study on the tooth attachment tissues in ''
Coelophysis ''Coelophysis'' ( Traditional English pronunciation of Latin, traditionally; or , as heard more commonly in recent decades) is a genus of coelophysid Theropoda, theropod dinosaur that lived Approximation, approximately 215 to 201.4 million y ...
bauri'' is published by Fong ''et al.'' (2016). * A study on the variation in morphological changes during
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
among members of the same species in early dinosaurs ''Coelophysis bauri'' and ''Megapnosaurus rhodesiensis'' as compared to the variation among living
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s and
crocodilia Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchia ...
ns is published by Griffin & Nesbitt (2016). * Senter & Juengst (2016) identify pathological features in eight
pectoral girdle The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans, it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists o ...
and forelimb bones of the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''
Dilophosaurus ''Dilophosaurus'' ( ) is a genus of theropod dinosaurs that lived in what is now North America during the Early Jurassic, about 186 million years ago. Three skeletons were discovered in northern Arizona in 1940, and the two best preserv ...
wetherilli''. * A study of
osteology Osteology () is the scientific study of bones, practiced by osteologists . A subdiscipline of anatomy, anthropology, archaeology and paleontology, osteology is the detailed study of the structure of bones, skeletal elements, teeth, microbone mo ...
and phylogenetic relationships of ''
Elaphrosaurus ''Elaphrosaurus'' ( ) is a genus of ceratosaurian Theropoda, theropod dinosaur that lived approximately 154 to 150 million years ago during the Late Jurassic, Late Jurassic Period in what is now Tanzania in Africa. ''Elaphrosaurus'' was a medium- ...
bambergi'' is published by Rauhut & Carrano (2016). * A new specimen of ''
Velocisaurus ''Velocisaurus'' ("swift lizard") is a genus of noasaurid theropod dinosaur from the Late Cretaceous period of Argentina. Description ''Velocisaurus'' was probably around long, based on a tibia length of . This makes it the smallest noasau ...
unicus'' is described by Brissón Egli, Agnolín & Novas (2016). * Footprints attributed to large
megalosaurid Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively ...
theropods are described from the Middle Jurassic (
Bathonian In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
) Serra de Aire Formation (
Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
) by Razzolini ''et al.'' (2016), who interpret the tracks as left by dinosaurs crossing the
tidal flat Mudflats or mud flats, also known as tidal flats or, in Ireland, slob or slobs, are coastal wetlands that form in intertidal areas where sediments have been deposited by tides or rivers. A global analysis published in 2019 suggested that tidal ...
during low tide periods. * A study on the validity of the theropod genus ''
Altispinax ''Altispinax'' (; "with high spines") is a genus of large predatory theropod dinosaur from the Early Cretaceous period (Valanginian, 140 to 133 million years ago) of what is now the Wadhurst Clay Formation of East Sussex, England. History Proba ...
'' is published by Maisch (2016). * Six isolated
spinosaurid Spinosauridae (or spinosaurids) is a clade or Family (taxonomy), family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America, and Asia. ...
quadrates, most likely coming from the
Kem Kem Beds The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations ...
, are described by Hendrickx, Mateus & Buffetaut (2016), who interpret the differences in their anatomy as confirming the presence of two
spinosaurine Spinosauridae (or spinosaurids) is a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America, and Asia. Their remains have ...
taxa in the
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
of North Africa, rather than only one (''
Spinosaurus ''Spinosaurus'' (; ) is a genus of large spinosaurid theropod dinosaurs that lived in what now is North Africa during the Cenomanian faunal stage, stage of the Late Cretaceous Period (geology), period, about 100 to 94 annum, million year ...
aegyptiacus''). * The description of a new large
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
femur (
Dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
ia:
Theropoda Theropoda (; from ancient Greek iktionary:θηρίον, , (''therion'') "wild beast"; wiktionary:πούς, , wiktionary:ποδός, (''pous, podos'') "foot" is one of the three major groups (Clade, clades) of Dinosaur, dinosaurs, alon ...
) from the
Kem Kem Beds The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations ...
, b
Alfio Alessandro Chiarenza & Andrea Cau (2016)
demonstrates the presence of large bodied individuals of this clade sympatric with other giant theropod dinosaurs from this area. This study includes also an overview on the
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
(Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
) theropod assemblage from Morocco. * Fossils of a large Early Cretaceous (
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) megaraptorid theropod are described from the
Griman Creek Formation The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian stages of the mid-Cretaceous. It is most notable being a major source of opal, ...
(
New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
,
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
) by Bell ''et al.'' (2016), who consider the theropod to be the largest predatory dinosaur yet identified from Australia. * A study on the manual anatomy of ''
Megaraptor ''Megaraptor'' () is a genus of large Theropoda, theropod dinosaur that lived during the Turonian and Coniacian ages of the Late Cretaceous. Its fossils have been discovered in the Patagonian Portezuelo Formation of Argentina, South America. Th ...
'' and ''
Australovenator ''Australovenator'' (meaning "southern hunter") is a genus of megaraptoran theropod dinosaur from Cenomanian (Late Cretaceous)-age Winton Formation (dated to 95 million years ago) of Australia. Some specimens from the Albian-aged Eumeralla Forma ...
'', as well as its implications for the phylogenetic relationships of these taxa, is published by
Novas A nova ( novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", Latin for "new") that slowly fades over weeks or months. All observed novae involve white d ...
, Aranciaga Rolando & Agnolín (2016). * A study of the phylogenetic relationships of
tyrannosauroid Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinen ...
theropods Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
is published by Brusatte and Carr (2016). *
Medullary bone The medullary cavity (''medulla'', innermost part) is the central cavity of bone shafts where red bone marrow and/or yellow bone marrow (adipose tissue) is stored; hence, the medullary cavity is also known as the marrow cavity. Located in the ma ...
homologous with one present in living birds is identified in a specimen of ''
Tyrannosaurus ''Tyrannosaurus'' () is a genus of large theropod dinosaur. The type species ''Tyrannosaurus rex'' ( meaning 'king' in Latin), often shortened to ''T. rex'' or colloquially t-rex, is one of the best represented theropods. It lived througho ...
rex'' by
Schweitzer Schweitzer is a surname. Notable people with the surname include: * Albert Schweitzer (1875–1965), German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize * Anton Schweitzer (1735–1787), German op ...
''et al.'' (2016). * Three fossil feathers from the Crato Member of the Early Cretaceous
Santana Formation The Santana Group is a geologic group, formerly included as the middle part of the Araripe Group, in the Araripe Basin of northeastern Brazil. The group comprises the Crato, Ipubi and Romualdo Formations and is dated to the Aptian to Albian s ...
(
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
) are described by Prado ''et al.'' (2016), who attribute them to
coelurosauria Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
n theropods of uncertain phylogenetic placement. * Feathered tail of a theropod dinosaur, probably of a juvenile non-avian
coelurosaur Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
, preserved in
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
(
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
-
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
)
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
is described by Xing ''et al.'' (2016) * A study of the effectiveness of proposed pathways for the evolution of the flight stroke in non-avian coelurosaurian theropods and early birds using biomechanical mathematical models is published by Dececchi, Larsson & Habib (2016). * A detailed description of the
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of the
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
and teeth of ''
Segnosaurus ''Segnosaurus'' is a genus of therizinosaurid dinosaur that lived in what is now southeastern Mongolia during the Late Cretaceous, about 102–86 million years ago. Multiple incomplete but well-preserved specimens were discovered in the Go ...
galbinensis'' is published by Zanno ''et al.'' (2016). * The first known
oviraptorosaur Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or wi ...
(''
Avimimus ''Avimimus'' ( ), meaning "bird mimic" (Latin ''avis'' = bird + ''mimus'' = mimic), is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 t ...
'')
bone bed A bone bed is any Geology, geological stratum or deposition (geology), deposit that contains bones of whatever kind. Inevitably, such deposits are Sedimentary rock, sedimentary in nature. Not a formal term, it tends to be used more to describe esp ...
is described from the
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
(
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
) by Funston ''et al.'' (2016). * New specimens of ''
Elmisaurus ''Elmisaurus'' (meaning "foot sole lizard") is an extinct genus of caenagnathid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was a theropod belonging to the Oviraptorosauria. Discovery In 1970, a paleontological Polish-Mo ...
rarus'' are described from the Late Cretaceous of
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
by
Currie Currie is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edinburgh Council. It is situated be ...
, Funston & Osmólska (2016). * New specimens of '' Leptorhynchos elegans'' and ''Leptorhynchos'' sp. are described from the Late Cretaceous of
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
by Funston, Currie & Burns (2016). * A study on the
micro- ''Micro'' (Greek letter μ, Mu (letter), mu, non-Italic type, italic) is a metric prefix, unit prefix in the metric system denoting a factor of one millionth (10−6). It comes from the Ancient Greek, Greek word (), meaning "small". It is the ...
and
ultrastructure Ultrastructure (or ultra-structure) is the architecture of cells and biomaterials that is visible at higher magnifications than found on a standard optical light microscope. This traditionally meant the resolution and magnification range of a c ...
of the fossil claw sheath of a specimen of ''
Citipati ''Citipati'' (; meaning "funeral pyre lord") is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous Period (geology), period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality ...
osmolskae'', indicating the preservation of original
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. It is the key structural material making up Scale (anatomy), scales, hair, Nail (anatomy), nails, feathers, horn (anatomy), horns, claws, Hoof, hoove ...
ous claw material, is published by Moyer, Zheng &
Schweitzer Schweitzer is a surname. Notable people with the surname include: * Albert Schweitzer (1875–1965), German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize * Anton Schweitzer (1735–1787), German op ...
(2016). * A study of the morphological disparity of teeth of
maniraptora Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, ...
n
theropods Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
living during the last 18 million years of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
is published by Larson, Brown and Evans (2016). * A robust ilium of a basal
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...
dinosaur is described from the
Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, so ...
(
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
) by McPhee & Choiniere (2016). * A new complete
femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...
assigned to ''
Pampadromaeus ''Pampadromaeus'' is an extinct genus of basal sauropodomorph dinosaurs known from the Late Triassic (Carnian) Santa Maria Formation of the Paraná Basin in Rio Grande do Sul, southern Brazil.Cabreira et al., 2011 Discovery ''Pampadromaeus'' ...
barberenai'' is described by Müller ''et al.'' (2016). * A study on the jaw adductor musculature and bite forces in ''
Plateosaurus ''Plateosaurus'' (probably meaning "broad lizard", often mistranslated as "flat lizard") is a genus of plateosaurid dinosaur that lived during the Late Triassic period, around 214 to 204 million years ago, in what is now Central and Northern Eu ...
'' and ''
Camarasaurus ''Camarasaurus'' ( ) is a genus of sauropod dinosaur that lived in North America during the Late Jurassic period. Its fossil remains have been found in the Morrison Formation, dating to the Kimmeridgian and Tithonian ages of the Jurassic, betwe ...
'' is published by Button, Barrett & Rayfield (2016). * A study of the evolution of whole-body shape and body segment properties of
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaurs is published by Bates ''et al.'' (2016). * A study on the intervertebral
joint A joint or articulation (or articular surface) is the connection made between bones, ossicles, or other hard structures in the body which link an animal's skeletal system into a functional whole.Saladin, Ken. Anatomy & Physiology. 7th ed. McGraw- ...
s in the necks and tails of sauropod dinosaurs, characterized by having the convex articular face directed away from the body and the concave articular face directed toward the body, is published by Fronimos, Wilson & Baumiller (2016), who argue that these joints evolved to prevent possible joint failure caused by rotation, providing stability with greater mobility and facilitating the evolution of elongated necks and tails in sauropods. * A restudy of '' Sanpasaurus yaoi'', originally classified as an
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
dinosaur, is published by McPhee ''et al.'' (2016), who consider this species to be an early sauropod instead. * Description of several sauropod vertebrae collected from the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Kirkwood Formation The Kirkwood Formation is a geological formation found in the Eastern and Western Cape provinces in South Africa. It is one of the four formations found within the Uitenhage Group of the Algoa Basin – its type locality – and in the neighbour ...
(
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
) and a study on the diversity of the sauropods known from the Kirkwood Formation is published by McPhee ''et al.'' (2016). * Gallina (2016) argues that ''
Amargatitanis ''Amargatitanis'' (meaning "La Amarga Formation, Amarga giant") is a genus of dicraeosaurid sauropod dinosaur (a type of large, long-necked quadrupedal herbivore, herbivorous dinosaur) from the Barremian-age (Lower Cretaceous) La Amarga Formati ...
macni'', initially considered to be a
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
, is actually a
dicraeosaurid Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as ''Amargasaurus'', ''Suuwassea'', '' ...
. * A reassessment of the systematics, paleoenvironment, life history and geologic age of ''
Sonorasaurus ''Sonorasaurus'' is a genus of brachiosaurid dinosaur from the Early to Late Cretaceous (Albian to Cenomanian stages, around 112 to 93 million years ago). It was a herbivorous sauropod whose fossils have been found in southern Arizona in th ...
thompsoni'' is published by D’Emic, Foreman & Jud (2016). * A study on divergence dates and ancestral ranges of Titanosauria is published by Gorscak & O‘Connor (2016). *
Osteoma An osteoma (plural ''osteomas'' or less commonly ''osteomata'') is a new piece of bone usually growing as a benign tumour on another piece of bone, typically the skull. When grown on other bone it is known as "homoplastic osteoma"; on other tissue ...
and
hemangioma A hemangioma or haemangioma is a usually benign vascular tumor derived from blood vessel cell types. The most common form, seen in infants, is an infantile hemangioma, known colloquially as a "strawberry mark", most commonly presenting on the sk ...
are documented for the first time in a
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
of a
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
from the Late Cretaceous of
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
by de Souza Barbosa ''et al.'' (2016). * Sauropod fossils, including a caudal vertebra attributed to a large-bodied
lithostrotia Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' an ...
n titanosaur, are reported from the Cretaceous
Kem Kem Beds The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations ...
(
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
) by Ibrahim ''et al.'' (2016). * A study on the anatomy of the
appendicular skeleton The appendicular skeleton is the portion of the vertebrate endoskeleton consisting of the bones, cartilages and ligaments that support the paired appendages ( fins, flippers or limbs). In most terrestrial vertebrates (except snakes, legless li ...
of ''
Dreadnoughtus ''Dreadnoughtus'' is a genus of titanosaurian sauropod dinosaur containing a single species, ''Dreadnoughtus schrani''. ''D. schrani'' is known from two partial skeletons discovered in Upper Cretaceous (Campanian to Maastrichtian; approximately ...
schrani'' is published by Ullmann & Lacovara (2016). * A study of the skull anatomy and phylogenetic relationships of ''
Tapuiasaurus ''Tapuiasaurus'' (meaning " Tapuia lizard") is a genus of titanosaur which lived during the Lower Cretaceous period (Aptian age) in what is now Minas Gerais, Brazil. Discovery Its fossils, including a partial skeleton with a nearly complet ...
macedoi'' is published by Wilson ''et al.'' (2016). * A juvenile specimen of ''
Rapetosaurus ''Rapetosaurus'' ( ) is a genus of titanosaurian sauropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period. Only one species, ''Rapetosaurus krausei'', has been identified. Like other sauropod ...
krausei'' is described by Curry Rogers ''et al.'' (2016). * Well-vascularised endosteally formed bone tissue is reported in the
saltasaurine Saltasauridae (named after the Salta region of Argentina where they were first found) is a family of armored herbivorous sauropods from the Upper Cretaceous. They are known from fossils found in South America, Africa, Asia, North America, and Eur ...
titanosaurs by Chinsamy, Cerda & Powell (2016), who argue that additional evidence is required to determine whether vascularised endosteal bone tissues reported in extinct archosaurs are
medullary bone The medullary cavity (''medulla'', innermost part) is the central cavity of bone shafts where red bone marrow and/or yellow bone marrow (adipose tissue) is stored; hence, the medullary cavity is also known as the marrow cavity. Located in the ma ...
or just a pathological bone. * A study on the effect of jaw shape and jaw adductor musculature on the relative bite force in members of 52
ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n genera is published by Nabavizadeh (2016). * A study on the anatomical diversity of the
predentary Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Ancient ...
in ornithischian dinosaurs is published by Nabavizadeh & Weishampel (2016). *
Heterodontosaurid Heterodontosauridae is a family of ornithischian dinosaurs that were likely among the most basal (primitive) members of the group. Their phylogenetic placement is uncertain but they are most commonly found outside of the group Genasauria. Alth ...
metatarsi,
phalanges The phalanges (: phalanx ) are digit (anatomy), digital bones in the hands and foot, feet of most vertebrates. In primates, the Thumb, thumbs and Hallux, big toes have two phalanges while the other Digit (anatomy), digits have three phalanges. ...
and tail
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e are described from the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic ...
(late
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
) Cañadon Asfalto Formation (
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
) by Becerra ''et al.'' (2016), who note the similarities in anatomy of the digits of this heterodontosaurid and the digits of arboreal
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s and argue that the heterodontosaurid might have had grasping feet with long digits. * New specimens of ''
Lesothosaurus ''Lesothosaurus'' is a Monotypic taxon, monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from L ...
diagnosticus'' are described by Barrett ''et al.'' (2016). * A description of the braincase anatomy of ''
Pawpawsaurus ''Pawpawsaurus'', meaning "Pawpaw Lizard", is a nodosaurid ankylosaur from the Cretaceous (late Albian) of Tarrant County, Texas, discovered in May 1992. The only species yet assigned to this taxon, ''Pawpawsaurus campbelli,'' is based on a compl ...
campbelli'' based on CT scans is published by Paulina-Carabajal, Lee & Jacobs (2016). * A new specimen of '' Haya griva'' is described from the Late Cretaceous of Mongolia by
Norell Norell is a surname. Notable people with the surname include: * Arne Norell (1917–1971), Swedish furniture designer *Ingeborg Norell (born 1727), the first Finnish female to have received an official decoration *Mark Norell (born 1957), American ...
& Barta (2016). * A reassessment of the holotype locality of ''
Leaellynasaura ''Leaellynasaura'' (meaning "Leaellyn's lizard") is a genus of small herbivorous ornithischian dinosaurs from the late Aptian to early Albian stage of the Early Cretaceous, around 118-110 million years ago.Holtz, Thomas R. Jr. (2012) ''Dinosaurs ...
amicagraphica'' is published by Herne, Tait & Salisbury (2016), who argue that several fossils traditionally referred to ''L. amicagraphica'' cannot be confidently attributed to this species. * A study on the evolution of the teeth morphologies of the
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
dinosaurs is published by Strickson ''et al.'' (2016), who argue that major increases of rates of dental character evolution among ornithopods did not correspond to times of plant diversification, including the radiation of the
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s. * Fossils of a diminutive
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
dinosaur, probably a member of
Rhabdodontidae Rhabdodontidae is a family of herbivorous iguanodontian ornithopod dinosaurs whose earliest stem members appeared in the middle of the Lower Cretaceous. The oldest dated fossils of these stem members were found in the Barremian Castrillo de l ...
, are described from the upper
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
-lower
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
Castrillo de la Reina Formation The Castrillo de la Reina Formation is a geological formation in Spain. It is late Barremian to early Aptian in age. It interpreted as a fluvial deposit. It primarily consists of red clay, with ribbon shaped sandstone channel fills. The rebbachisa ...
( Cameros Basin,
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) by Dieudonné ''et al.'' (2016). * A new specimen of ''
Valdosaurus ''Valdosaurus'' ("Weald Lizard") is a genus of bipedal herbivorous iguanodont ornithopod dinosaur found on the Isle of Wight and elsewhere in England, and possibly also Spain and Romania. It lived during the Early Cretaceous. Discovery and namin ...
canaliculatus'', the most complete yet found, is described by Barrett (2016). *
Tibia The tibia (; : tibiae or tibias), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two Leg bones, bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outsi ...
and tail vertebrae of
iguanodontia Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
n dinosaurs are described from the
Cleaver Bank The Cleaver Bank ( Dutch: ''Klaverbank)'' is a sandbank in the North Sea about off the west coast of the Netherlands and south of the Dogger Bank. The size of the bank is about 1235 km2. The bank is 30-40 metres below sea level. The Cleaver ...
(
North Sea The North Sea lies between Great Britain, Denmark, Norway, Germany, the Netherlands, Belgium, and France. A sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian Se ...
) by Mulder & Fraaije (2016). * Isolated teeth of large-bodied
iguanodontia Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
ns are described from the Early Cretaceous (
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) of
Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
by Fanti ''et al.'' (2016). * Parallel trackways of medium-sized and robust
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
s similar to ''
Draconyx ''Draconyx'' (meaning "dragon claw") is a genus of dinosaur from the Late Jurassic. It was an Ornithopoda, ornithopod which lived in what is now Portugal and was a herbivore. It was found in the Lourinhã Formation in 1991, and described by Oct ...
'' or ''
Cumnoria ''Cumnoria'' is a genus of herbivorous iguanodontian dinosaur. It was a basal iguanodontian that lived during the Late Jurassic period (Kimmeridgian age) in what is now Oxfordshire, United Kingdom. Description The holotype of ''Cumnoria'' is a ...
'', providing evidence of gregarious behavior, are described from the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
of
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
by Piñuela ''et al.'' (2016). * A
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
of ''
Telmatosaurus ''Telmatosaurus'' (meaning "marsh lizard") is a genus of basal (phylogeny), basal hadrosauromorph dinosaur from the Late Cretaceous of Romania. It was a relatively small hadrosaur, measuring approximately in length and in body mass, which has b ...
transsylvanicus'' exhibiting
ameloblastoma Ameloblastoma is a rare, benign or cancerous tumor of odontogenic epithelium ( ameloblasts, or outside portion, of the teeth during development) much more commonly appearing in the lower jaw than the upper jaw. It was recognized in 1827 by Cusa ...
is described from the Late Cretaceous Sînpetru Formation (Hațeg Basin,
Romania Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
) by Dumbravă ''et al.'' (2016). * A revision of the original diagnosis of '' Willinakaqe salitralensis'' and of fossil material attributed to this species is published by Cruzado Caballero and Coria (2016), who argue that the fossils attributed to ''Willinakaqe salitralensis'' might represent more than a single taxon of
hadrosaurid Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
and that all characters of the original diagnosis are invalid. * Large
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
(probably
hadrosaurid Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
) tracks, assigned to the
ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxon'' comes from the Ancient Greek (''íchnos'') meaning "track" and English , itself derived from ...
'' Hadrosauropodus'', are described from the
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-
Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretac ...
Yacoraite Formation The Yacoraite Formation is a largely Mesozoic geologic formation. The deposits of this formation mainly date from the Maastrichtian of the Upper Cretaceous, but the Cretaceous–Paleogene boundary (K–T boundary) runs right through this formati ...
of
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
by Díaz-Martínez, de Valais & Cónsole-Gonella (2016). * A hadrosaurid
radius In classical geometry, a radius (: radii or radiuses) of a circle or sphere is any of the line segments from its Centre (geometry), center to its perimeter, and in more modern usage, it is also their length. The radius of a regular polygon is th ...
and
ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
affected by a severe
septic arthritis Acute septic arthritis, infectious arthritis, suppurative arthritis, pyogenic arthritis, osteomyelitis, or joint infection is the invasion of a joint by an infectious agent resulting in joint inflammation. Generally speaking, symptoms typica ...
are described from the Late Cretaceous
Navesink Formation The Navesink Formation is a 66 to 70 mya greensand glauconitic marl and sand geological formation in New Jersey. It is known for its Cretaceous period fossil shell beds and dinosaur bones. Description The Navesink Formation, named after Navesin ...
(
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
, USA) by Anné, Hedrick & Schein (2016). * A study on the development of the dental battery of the hadrosaurid dinosaurs through their
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
and on the evolution of the hadrosaurid dental battery is published by LeBlanc ''et al.'' (2016). * Chondroid bone (a tissue intermediate between
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
and
cartilage Cartilage is a resilient and smooth type of connective tissue. Semi-transparent and non-porous, it is usually covered by a tough and fibrous membrane called perichondrium. In tetrapods, it covers and protects the ends of long bones at the joints ...
) is reported in embryos and nestlings of ''
Hypacrosaurus ''Hypacrosaurus'' (meaning "near the highest lizard"
reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high Reek may refer to: Places * Reek, Netherlands, a village in the Dutch province of North Brabant * Croagh Patrick, a mountain in the west of Ireland nicknamed "The Reek" People * Nikolai Reek (1890–1942), Estonian military commander * Salme Ree ...
because it was almost but not quite as large as ''Tyrannosaurus'') is an extinct genus of hadrosaurid, duckbill dinosaur simila ...
'' by Bailleul ''et al.'' (2016). * Restudies of the fossil material attributed to ''
Stegoceras ''Stegoceras'' is a genus of Pachycephalosauria, pachycephalosaurid (dome-headed) dinosaur that lived in what is now North America during the Late Cretaceous Period (geology), period, about 77.5 to 74 million years ago (mya). The first specim ...
novomexicanum'' are published by Williamson & Brusatte (2016) and Jasinski & Sullivan (2016). * A study on the skull anatomy of ''
Yinlong ''Yinlong'' (, meaning "hidden dragon") is a genus of basal ceratopsian dinosaur from the Late Jurassic Period of China. By far the earliest known ceratopsian, it was a small, primarily bipedal herbivore. Discovery and species A coalition of ...
downsi'' is published by Han ''et al.'' (2016). * A study of the bristle-like appendages on the tail of ''
Psittacosaurus ''Psittacosaurus'' ( ; "parrot lizard") is a genus of extinct ceratopsian dinosaur from the Early Cretaceous of what is now Asia, existing between 125 and 105 million years ago. It is notable for being the most species-rich non-avian dinosaur ...
'' is published by
Mayr Mayr is a German surname. Notable people with the surname include: * Andrea Mayr (born 1979), Austrian female long-distance runner * Ernst Mayr (1904–2005), German American evolutionary biologist * Franz Xaver Mayr (1875–1965), Austrian gastro ...
''et al.'' (2016). * A study on the color patterns of a well-preserved specimen of ''Psittacosaurus'' sp. as indicated by the distribution of organic residues is published by Vinther ''et al.'' (2016). * A study on the dental microwear in ''
Leptoceratops ''Leptoceratops'' (meaning 'small horn face') is a genus of ceratopsian dinosaur from the Late Cretaceous of North America. First found in Alberta in 1910, the type species ''Leptoceratops gracilis'' was named in 1914 by Barnum Brown for a parti ...
gracilis'' is published by Varriale (2016). * A study of the frill bones of ''
Protoceratops ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
andrewsi'', indicating that its frill increased in length and width during the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
of the animal and that the growth of the frill was greater than the overall growth of the animal, is published by Hone, Wood & Knell (2016), who interpret these findings as indicating that ''Protoceratops'' most likely used its frill for sexual and social dominance signaling. * Partial skull of a
ceratopsid Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', '' Centrosaurus'', and '' Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are ...
related to ''
Nasutoceratops ''Nasutoceratops'' is a genus of ceratopsid dinosaur that lived in North America during the Late Cretaceous period, about 76.0–75.5 million years ago. The first known specimens were discovered in Utah in the Kaiparowits Formation of the Grand ...
titusi'' is described from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...
(
Alberta Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) by Ryan ''et al.'' (2016), who also name new ceratopsid tribes Centrosaurini and
Nasutoceratopsini Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid, a group of large quadrupedal ornithischian dinosaur. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alber ...
. * A revision of the species assigned to the genus ''
Chasmosaurus ''Chasmosaurus'' ( ) is a genus of ceratopsid dinosaur from the Late Cretaceous Period in North America. Its given name means 'opening lizard', referring to the large openings ( fenestrae) in its frill (Greek ''chasma'', meaning 'opening', 'hol ...
'' is published by Campbell ''et al.'' (2016). * Forelimb studies show ''Oryctodromeus'' was extremely adapted for an underground lifestyle (2016). * A group of paleontologists discovered the remains of the smallest specimen of ''Pachycephalosaurus'' to date. The specimen also casts doubt on the validity of ''Dracorex'' and ''Stygimoloch'' (2016). * A study was done on the skulls of ''Majungasaurus'' and revealed changes throughout the life cycle of this dinosaur (2016). * A study was conducted on the skeleton of ''Nasutoceratops'', revealing that it and ''Avaceratops'' belonged to a completely new group of centrosaurines (2016).


New taxa


Birds


Research

* A study on the rates of morphological evolution in Early Cretaceous birds is published by Wang and Lloyd (2016). * A study on the microbodies associated with feathers of a new specimen of '' Eoconfuciusornis'' from the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Huajiying Formation The Huajiying Formation is a geological formation in Hebei, People's Republic of China. Known for its fossils including feathered dinosaurs, the age of the formation is uncertain. It may represent an early portion of the Jehol Biota, dating to som ...
(
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) and on the matrix in which the microbodies were embedded is published by Pan ''et al.'' (2016), who interpret the microbodies as
melanosome A melanosome is an organelle found in animal cells and is the site for synthesis, storage and transport of melanin, the most common light-absorbing pigment found in the animal kingdom. Melanosomes are responsible for color and photoprotectio ...
s. * Remains of non-
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
soft tissues, including scales, toe pads, skin and muscle, are identified in two specimens of ''
Confuciusornis ''Confuciusornis'' is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, ''Confuciusornis'' had a toothless beak, bu ...
'' by Falk ''et al.'' (2016). * A skeleton of an
enantiornithine The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and c ...
bird preserving a gastric pellet that includes fish bones is described from the Early Cretaceous Jehol Biota of China by Wang, Zhou & Sullivan (2016). * Two partial wings with vestiges of soft tissues, probably belonging to precocial hatchlings of enantiornithine birds, are described from the Late Cretaceous (
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
)
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
by Xing ''et al.'' (2016). * A revised diagnosis of ''
Cerebavis ''Cerebavis'' (from Latin ''cerebrum'', "brain", and ''avis'', "bird") is an extinct genus of ornithuran dinosaurs that lived during the middle Cenomanian of the Late Cretaceous period, and is known from a single partial skull (PIN 5028/2) found ...
cenomanica'', a study on the braincase anatomy of the species and a study on its phylogenetic relationships is published by Walsh, Milner & Bourdon (2016). * A study on the shape, growth, attachment, implantation, replacement, and tissue microstructures of the teeth of ''
Hesperornis ''Hesperornis'' (meaning "western bird") is a genus of cormorant-like Ornithuran that spanned throughout the Campanian age, and possibly even up to the early Maastrichtian age, of the Late Cretaceous period. One of the lesser-known discoverie ...
'' and ''
Ichthyornis ''Ichthyornis'' (meaning "fish bird", after its fish-like vertebrae) is an extinct genus of toothy seabird-like ornithuran from the late Cretaceous period of North America. Its fossil remains are known from the chalks of Alberta, Alabama, Kansa ...
'' is published by Dumont ''et al.'' (2016). * A phylogenetic analysis of
Hesperornithiformes Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as ''Hesperorn ...
is published by Bell & Chiappe (2016). * A specimen of ''
Hesperornis ''Hesperornis'' (meaning "western bird") is a genus of cormorant-like Ornithuran that spanned throughout the Campanian age, and possibly even up to the early Maastrichtian age, of the Late Cretaceous period. One of the lesser-known discoverie ...
'' with a healed wound is described from the Late Cretaceous
Pierre Shale The Pierre Shale is a geologic formation or series in the Upper Cretaceous which occurs east of the Rocky Mountains in the Great Plains, from Pembina Valley in Canada to New Mexico. The Pierre Shale was described by Fielding Bradford Meek, Meek a ...
(
South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state, state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Dakota people, Dakota Sioux ...
,
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) by
Martin Martin may refer to: Places Antarctica * Martin Peninsula, Marie Byrd Land * Port Martin, Adelie Land * Point Martin, South Orkney Islands Europe * Martin, Croatia, a village * Martin, Slovakia, a city * Martín del Río, Aragón, Spain * M ...
, Rothschild & Burnham (2016), who interpret the wound as caused by an unsuccessful attack of a
polycotylid Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. They are known as false pliosaurs. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widesprea ...
plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
. *
Pelvic The pelvis (: pelves or pelvises) is the lower part of an anatomical trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis or pelvic skeleton). ...
elements of ''
Gargantuavis ''Gargantuavis'' (meaning 'gargantuan bird') is an extinct genus of large, primitive bird containing the single species ''Gargantuavis philoinos''. It is the only member of the monotypic family Gargantuaviidae. Its fossils were discovered in seve ...
philoinos'', providing new information about the pelvic
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of the species, are described from the Late Cretaceous (late
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
/early
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
) of southern
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
by Buffetaut & Angst (2016). * A specimen of ''
Vegavis ''Vegavis'' is a genus of extinct bird that lived in Antarctica during the Maastrichtian stage of the Late Cretaceous. The type and only species is ''Vegavis iaai'', representing one of the earliest known crown group birds. Initially described ...
iaai'' with a fossilized
syrinx In classical Greek mythology, Syrinx () was an Arcadian nymph and a follower of Artemis, known for her chastity. Being pursued by Pan, she fled into the river Ladon, and at her own request was metamorphosed into a reed from which Pan then mad ...
is described from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
by Clarke ''et al.'' (2016). * A study on the feeding mechanics and behaviour of five
moa Moa are extinct giant flightless birds native to New Zealand. Moa or MOA may also refer to: Arts and media * Metal Open Air, a Brazilian heavy metal festival * MOA Museum of Art in Japan * The Moas, New Zealand film awards People * Moa ...
species is published by Attard ''et al.'' (2016). * Mariana B.J. Picasso & María Clelia Mosto, 2016: ''Hinasuri nehuensis'' Tambussi was a robust, extinct rheid bird from the early Pliocene of Buenos Aires province, Argentina. This paper revisits the femoral morphology of ''H. nehuensis'' and provides an updated osteological description together with new insights into its palaeobiology. * Restudies of the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
species ''Rhea pampeana'' and ''Rhea anchorenensis'' are published by Picasso (2016) and Picasso and Mosto (2016), respectively, who consider these species to be junior synonyms of the extant
greater rhea The greater rhea (''Rhea americana'') is a species of flightless bird native to eastern South America. Other names for the greater rhea include the grey, common, or American rhea; ema (Portuguese (language), Portuguese); or ñandú (Guaraní lang ...
(''Rhea americana''). * Demarchi ''et al.'' (2016) report the recovery of mineral-bound protein sequences from ostrich eggshells from the paleontological sites of
Laetoli Laetoli is a pre-historic site located in Enduleni ward of Ngorongoro District in Arusha Region, Tanzania. The site is dated to the Plio-Pleistocene and famous for its Hominina footprints, preserved in volcanic ash. The site of the Laetoli footp ...
and
Olduvai Gorge The Olduvai Gorge or Oldupai Gorge in Tanzania is one of the most important paleoanthropology, paleoanthropological localities in the world; the many sites exposed by the gorge have proven invaluable in furthering understanding of early human evo ...
(
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
). * Worthy ''et al.'' (2016) argue that ''
Sylviornis ''Sylviornis'' is an extinct genus of large, flightless bird that was endemic to the islands of New Caledonia in the Western Pacific. It is considered to constitute one of two genera in the extinct family Sylviornithidae, alongside '' Megaviti ...
neocaledoniae'' is a
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
-
galliform Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...
related to ''
Megavitiornis altirostris ''Megavitiornis altirostris'' is an extinct, flightless, giant stem-galliform bird that was endemic to Fiji, it is the only known species in the genus ''Megavitiornis''. Originally thought to be a megapode, more recent morphological studies indic ...
'' and both are placed in the
Sylviornithidae Sylviornithidae is an extinct family of flightless birds, known from subfossil bones found in Holocene aged deposits on the Melanesian islands of New Caledonia and Fiji. For many years it was considered a monotypic family consisting of the New ...
Mourer-Chauviré et Balouet, 2005. * A revision of the systematics of the early Eocene North American members of
Geranoididae Geranoididae is a clade of extinct birds from the early to late Eocene and possibly early Oligocene of North America and Europe. These were mid-sized, long-legged flightless birds.Gerald Mayr (2009). Paleogene Fossil Birds Recent research shows t ...
is published by Mayr (2016), who argues that geranoidids might be
stem group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...
representatives of the Gruoidea (the clade including
trumpeters The trumpet is a brass instrument commonly used in classical and jazz ensembles. The trumpet group ranges from the piccolo trumpet—with the highest register in the brass family—to the bass trumpet, pitched one octave below the standard B ...
, cranes and related birds). * Zelenkov, Boev & Lazaridis (2016) reinterpret ''Otis hellenica'' from the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
of
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
, originally thought to be a
bustard Bustards, including floricans and korhaans, are large, terrestrial birds living mainly in dry grassland areas and in steppe regions. They range in length from . They make up the family Otididae (, formerly known as Otidae). Bustards are ...
, as a member of
Gruiformes The Gruiformes ( ) are an order containing a considerable number of living and extinct bird families, with a widespread geographical diversity. Gruiform means "crane-like". Traditionally, a number of wading and terrestrial bird families that ...
belonging to the family
Eogruidae Eogruidae (also spelled Eogruiidae in some publications) is an extinct family of large, flightless birds that inhabited Asia from the Eocene to Pliocene epochs. Related to modern ostriches, it was formerly thought to be related to cranes, limpki ...
and the subfamily Ergilornithinae; the authors classify it as a possible member of the genus '' Amphipelargus'' of uncertain specific assignment ("?''Amphipelargus'' sp."). * A restudy of the holotype specimen of ''
Bathornis ''Bathornis'' ("tall bird") is an extinct genus of birds related to modern day seriemas, that lived in North America about 37–20 million years ago. Like the closely related and also extinct phorusrhacids, it was a flightless predator, occupying ...
grallator'' and a study on the taxonomic composition and phylogenetic affinities of
bathornithids Bathornithidae is an extinct family of birds from the Eocene to Miocene of North America. Part of Cariamiformes, they are related to the still extant seriemas and the extinct Phorusrhacidae. They were likely similar in habits, being terrestria ...
is published by Mayr (2016). * Zelenkov, Volkova and Gorobets (2016) describe
buttonquail Buttonquail or hemipodes are members of a small family of birds, Turnicidae, which resemble, but are not closely related to, the quails of Phasianidae. They inhabit warm grasslands in Asia, Africa, Europe, and Australia. There are 18 species in ...
fossils from the late Miocene of
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
, southern
Ukraine Ukraine is a country in Eastern Europe. It is the List of European countries by area, second-largest country in Europe after Russia, which Russia–Ukraine border, borders it to the east and northeast. Ukraine also borders Belarus to the nor ...
and northern
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a landlocked country primarily in Central Asia, with a European Kazakhstan, small portion in Eastern Europe. It borders Russia to the Kazakhstan–Russia border, north and west, China to th ...
, and transfer the species ''
Calidris ''Calidris'' is a genus of Arctic-breeding, strongly migratory wading birds in the family Scolopacidae. These birds form huge mixed flocks on coasts and estuaries in winter. Migratory shorebirds are shown to have decline in reproductive traits ...
janossyi'' Kessler (2009) to the genus '' Ortyxelos''. * Gerald Mayr and Zbigniew M. Bochenski,(2016) describe a disarticulated postcranial skeleton of a Ralloidea from the Early Oligocene (Rupelian) Jamna Dolna Site 2 in Poland as Gen. et Sp. indet. * Agnolin, Tomassini and Contreras (2016) describe a distal end of
tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...
from the late Miocene levels of the Loma de Las Tapias Formation ( San Juan Province,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
), identified as the oldest
seedsnipe The seedsnipes are a small family, Thinocoridae, of small gregarious waders which have adapted to a herbivorous diet. The family is divided into two genera, '' Attagis'' and '' Thinocorus'', each containing two species. The family has a South Am ...
fossil discovered so far. * Body mass estimates for 25 extinct pan-alcids and a study of body mass evolution in Pan-Alcidae are published by Smith (2016). * The earliest known
cranial Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
of a
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
-
penguin Penguins are a group of aquatic flightless birds from the family Spheniscidae () of the order Sphenisciformes (). They live almost exclusively in the Southern Hemisphere. Only one species, the Galápagos penguin, is equatorial, with a sm ...
(a member of the genus ''
Waimanu ''Waimanu'' is a genus of early penguin which lived during the Paleocene, soon after the Cretaceous–Paleogene extinction event, around 62–60 million years ago. It was about the size of an emperor penguin (1 metre (3.3 ft)). It is one of the ...
'') is described from the Paleocene Waipara Greensand (
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
) by Proffitt, Clarke & Scofield (2016). * Thomas & Ksepka (2016) classify a Whaingaroan penguin from the Glen Massey Formation (
North Island The North Island ( , 'the fish of Māui', historically New Ulster) is one of the two main islands of New Zealand, islands of New Zealand, separated from the larger but less populous South Island by Cook Strait. With an area of , it is the List ...
, New Zealand), first described in 1973, as a member of the genus '' Kairuku'' of uncertain specific assignment, extending the geographic range of the genus. * Park ''et al.'', 2016 The description of recently collected penguin fossils from the re-dated upper Miocene Port Campbell Limestone of Portland (Victoria), in addition to reanalysis of previously described material, has allowed the Cenozoic history of penguins in Australia to be placed into a global context for the first time. Australian pre-Quaternary fossil penguins represent stem taxa phylogenetically disparate from each other and ''
Eudyptula minor The little penguin (''Eudyptula minor'') is the smallest species of penguin. It originates from New Zealand. It is commonly known as the fairy penguin, little blue penguin, or blue penguin, owing to its slate-blue plumage and is also known by ...
'', implying multiple dispersals and extinctions. * Carolina Acosta Hospitaleche, Leandro M. Pérez, Sergio Marenssi, Marcelo Reguero (2016). The purpose of this paper is to provide a taphonomic analysis of the holotype of '' Crossvallia unienwillia'', in order to improve the knowledge of the vertebrate record of the Cross Valley Formation, a unit exposed in the central area of Marambio (Seymour) Island, Antarctic Peninsula. * A new skeleton of the Eocene penguin ''
Palaeeudyptes klekowskii ''Palaeeudyptes klekowskii'' is an extinct species of the penguin genus ''Palaeeudyptes''. It was previously thought to have been approximately the size of its congener ''Palaeeudyptes antarcticus'', somewhat larger than the modern emperor pengu ...
'' is described from the Submeseta Formation (
Seymour Island Seymour Island or Marambio Island, is an island in the chain of 16 major islands around the tip of the Graham Land on the Antarctic Peninsula. Graham Land is the closest part of Antarctica to South America. It lies within the section of the isl ...
,
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
) by Acosta Hospitaleche (2016). * Carolina Acosta Hospitaleche & Eduardo Olivero, 2016: Eocene penguins are known mostly from Antarctic specimens. A previously documented partial skeleton consisting of a pelvis, femur, tibiotarsus and fibula, from the middle Eocene Leticia Formation, Tierra del Fuego Province, Argentina, has been prepared and re-described. Re-analysis favours assignment to ''Palaeeudyptes gunnari'', a species widely recorded in the Eocene of Antarctica. * Fossils of a
stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons and ibise ...
and a
heron Herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 75 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genus ''Botaurus'' are referred to as bi ...
belonging or related to the tribe Nycticoracini are described from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
by Stidham ''et al.'' (2016). * A restudy of the fossils attributed to the species ''Liornis floweri'' and ''Callornis giganteus'' from the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
Santa Cruz Formation The Santa Cruz Formation is a geological formation in the Magallanes Basin, Magallanes/Austral Basin in southern Patagonia in Argentina and adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with the eponym ...
(
Patagonia Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
) is published by Buffetaut (2016), who considers ''L. floweri'' to be a
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
of ''
Brontornis ''Brontornis'' is an extinct genus of giant bird that inhabited Argentina during the Early Miocene, Early to Middle Miocene. Its taxonomic position is highly controversial, with authors alternatively considering it to be a Cariamiformes, cariami ...
burmeisteri'' and considers ''C. giganteus'' to be a
chimera Chimera, Chimaera, or Chimaira (Greek for " she-goat") originally referred to: * Chimera (mythology), a fire-breathing monster of ancient Lycia said to combine parts from multiple animals * Mount Chimaera, a fire-spewing region of Lycia or Cilicia ...
based on a
phorusrhacid Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous, mostly flightless birds that were among the largest apex predators in South America during the Cenozoic era. Their definitive fossil records range from ...
tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...
and a brontornithid
tibiotarsus The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These ...
. * A study of eggshell fragments from the Pleistocene of Australia putatively referred to ''
Genyornis ''Genyornis newtoni'' is an extinct species of large, flightless bird that lived in Australia during the Pleistocene Epoch until around 50,000 years ago. Over two metres in height, they were likely herbivorous. Many other species of Austral ...
newtoni'' is published by Grellet-Tinner, Spooner & Worthy (2016), who argue that these fossils are more likely to be remains of eggs laid by
megapode The megapodes, also known as incubator birds or mound-builders, are stocky, medium-large, chicken-like birds with small heads and large feet in the family Megapodiidae. Their name literally means "large foot" and is a reference to the heavy leg ...
s. Based on the similarities in the structure of eggshells of megapodes and dromornithids, the authors also hypothesize that dromornithids might be a
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to
galliforms Galliformes is an order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed disperser ...
rather than to or within anseriforms. * A study of burnt putative ''
Genyornis ''Genyornis newtoni'' is an extinct species of large, flightless bird that lived in Australia during the Pleistocene Epoch until around 50,000 years ago. Over two metres in height, they were likely herbivorous. Many other species of Austral ...
'' eggshell fragments from the Pleistocene of Australia is published by Miller ''et al.'' (2016), who interpret them as confirming that eggs of ''Genyornis newtoni'' were harvested by humans. * A study on the possible presence, form, and extent of sexual dimorphism in ''
Dromornis ''Dromornis'' is a genus of large to enormous prehistoric birds native to Australia during the Oligocene to Pliocene epochs. The species were flightless, possessing greatly reduced wing structures but with large legs, similar to the modern ostric ...
stirtoni'' is published by Handley ''et al.'' (2016). * Gastornithid and presbyornithid fossils are described from the early Eocene of
Ellesmere Island Ellesmere Island (; ) is Canada's northernmost and List of Canadian islands by area, third largest island, and the List of islands by area, tenth largest in the world. It comprises an area of , slightly smaller than Great Britain, and the total ...
(
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) by Stidham & Eberle (2016). * The genus ''
Wilaru ''Wilaru'' is an extinct genus of presbyornithid bird from Australia during the Late Oligocene to Early Miocene, around 26-22 million years ago. The type species is ''Wilaru tedfordi'', and the second species is ''Wilaru prideauxi''. Discovery ...
'', initially considered to be of a
stone-curlew The stone-curlews, also known as dikkops or thick-knees, consist of 10 species within the family Burhinidae, and are found throughout the tropical and temperate parts of the world, with two or more species occurring in some areas of Africa, Asia, ...
, is reinterpreted as a member of
Presbyornithidae Presbyornithidae is an extinct group of birds found in North America, South America, East Asia, Australia and possibly North Africa. They had evolved by the Maastrichtian age of the Late Cretaceous and became extinct during the Aquitanian age of ...
by De Pietri ''et al.'' (2016); the authors also reassess the Cretaceous species ''
Teviornis ''Teviornis'' is an extinct genus of presbyornithid which lived during the Maastrichtian stage, around 70 million years ago. One species has been described, ''T. gobiensis''. It is the oldest known neognath along with '' Vegavis'', and its fos ...
gobiensis'' and confirm it as a member of Presbyornithidae. * A revision of
anseriform Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which inc ...
birds known from the late
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
localities in central
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
is published by Zelenkov (2016), who transfers the species ''
Anas ''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
denesi'' Kessler (2013) to the genus ''
Aythya ''Aythya'' is a genus of diving ducks, with twelve species currently accepted. The genus was described in 1822 by the German zoologist Friedrich Boie, with the type species being greater scaup. The name ''Aythya'' comes from the Ancient Greek ...
'' and classifies the species ''Anas albae'' Janossy (1979) as a member of tribe
Mergini The sea ducks (Mergini) are a tribe of the duck subfamily of birds, the Anatinae. The taxonomy of this group is incomplete. Some authorities separate the group as a subfamily, while others remove some genera. Most species within the group spend ...
of uncertain generic assignment. * A revision of
galliform Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...
birds known from the late Miocene localities in central Hungary is published by Zelenkov (2016), who transfers the subspecies '' Pavo aesculapi phasianoides'' Janossy (1991) to the genus ''
Syrmaticus __NOTOC__ The genus ''Syrmaticus'' contains the five species of long-tailed pheasants. The males have short spurs and usually red facial wattles, but otherwise differ wildly in appearance. The hens (females) and chicks of all the species have a ...
'' and raises it to the rank of a separate species '' Syrmaticus phasianoides''. * New fossil remains of the Eocene
cuckoo Cuckoos are birds in the Cuculidae ( ) family, the sole taxon in the order Cuculiformes ( ). The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals, and anis. The coucals and anis are somet ...
''
Chambicuculus ''Chambicuculus'' is an extinct genus of cuculid that lived during the Eocene epoch. Distribution ''Chambicuculus pusillus'' is known from the Chambi Formation of Tunisia Tunisia, officially the Republic of Tunisia, is a country in the M ...
pusillus'' are described from
Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
by Mourer-Chauviré ''et al.'' (2016). * Virtual cranial
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
of the
dodo The dodo (''Raphus cucullatus'') is an extinction, extinct flightless bird that was endemism, endemic to the island of Mauritius, which is east of Madagascar in the Indian Ocean. The dodo's closest relative was the also-extinct and flightles ...
is described by Gold, Bourdon &
Norell Norell is a surname. Notable people with the surname include: * Arne Norell (1917–1971), Swedish furniture designer *Ingeborg Norell (born 1727), the first Finnish female to have received an official decoration *Mark Norell (born 1957), American ...
(2016). * An
ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropod Sauropoda (), whose members are known as sauropods (; ...
phalanx The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
of a large member of
Accipitridae The Accipitridae () is one of the four families within the order Accipitriformes, and is a family of small to large birds of prey with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects ...
belonging to an unknown genus and species is described from the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
of
Panama Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
by Steadman & MacFadden (2016). * Partial
tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...
of a small parrot is described from the Early Miocene Khalagay Formation (Baikal region, Russia) by Zelenkov (2016). * A study on the phylogenetic relationships of extant and extinct
New Zealand wren The New Zealand wrens are a family (Acanthisittidae) of tiny passerines endemic to New Zealand. They were represented by seven Holocene species in four or five genera, although only two species in two genera survive today. They are understood to ...
s, as indicated by data from novel mitochondrial genome sequences, is published by Mitchell ''et al.'' (2016). * Fossil avian feet from the Early Eocene of Messel, Germany are described by Gerald Mayr * A new tracksite with bird footprints (attributed to the ichnospecies '' Uvaichnites riojana''), preserved in the early Miocene Lerín Formation (
Bardenas Reales The (sometimes referred as ' in Spanish; in Basque: ''Errege Bardeak'') is a semi-desert natural region, or badlands, of some in southeast Navarre (Spain). The soils are made up of clay, chalk, and sandstone and have been eroded by water and ...
de Navarra Natural Park,
Navarre Navarre ( ; ; ), officially the Chartered Community of Navarre, is a landlocked foral autonomous community and province in northern Spain, bordering the Basque Autonomous Community, La Rioja, and Aragon in Spain and New Aquitaine in France. ...
,
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
), is described by Díaz-Martínez ''et al.'' (2016). * A new ichnospecies, ''Koreananornis lii'', from the Lower Cretaceous avian track locality in the Guanshan area, Yongjing County, Gansu Province, northwest China, is described by Xing, Buckley, Lockley, Zhang, Marty, Wang, Li, McCrea et Peng, 2016. (2016). * An avian egg from the Lower Cretaceous (Albian) Liangtoutang Formation is described by Lawver ''et al.'' (2016) and named ''Pachycorioolithus jinyunensis'' oogen. et oosp. nov. within Pachycorioolithidae oofam. nov. * Three pellets with bird remains are described from the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
Messel pit The Messel Formation is a Formation (geology), geologic formation in Hesse, central Germany, dating back to the Eocene Epoch (geology), epoch (about 47 Ma). Its geographic range is restricted to the Messel pit. There it unconformably overlie ...
(
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
) by
Mayr Mayr is a German surname. Notable people with the surname include: * Andrea Mayr (born 1979), Austrian female long-distance runner * Ernst Mayr (1904–2005), German American evolutionary biologist * Franz Xaver Mayr (1875–1965), Austrian gastro ...
& Schaal (2016), who interpret two of the pellets as probably produced by snakes or other
squamates Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles; most members of which are commonly known as lizards, with the group also including snakes. With over 11,991 species, it is also the second-largest order ...
, and one as probable owl pellet (which, if confirmed, would make it the oldest owl pellet identified so far), possibly produced by the owl ''
Palaeoglaux ''Palaeoglaux'' is a genus of fossil owls from the Eocene epoch. The two known species are ''P. perrierensis'' from the Upper Eocene of Quercy, France, and ''P. artophoron'' from the Middle Eocene Messel shales, Germany.Mourer-Chauviré, Céci ...
artophoron''.


New taxa


Pterosaurs


Research

* A new
wukongopterid Wukongopteridae is a group of basal pterosaurs, found in China and the UK. It contains eight species in five genera, all dated to the Middle to Late Jurassic period, The Wukongopteridae were first named by Wang ''et al.'' in 2009, not yet givin ...
specimen is described from the Late Jurassic Daohugou Bed or Tiaojishan Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) by Cheng ''et al.'' (2016). * Description of a new specimen of ''
Gladocephaloideus ''Gladocephaloideus'' is a genus of ctenochasmatid Ctenochasmatoidea, ctenochasmatoid pterosaur from the Early Cretaceous period (early Aptian stage) of what is now western Liaoning, China. Discovery ''Gladocephaloideus'' is known from a compl ...
jingangshanensis'' and a study of the phylogenetic relationships of this species is published by Lü, Kundrát & Shen (2016). * New information on the braincase anatomy of ''
Pterodaustro ''Pterodaustro'' (from Greek language, Greek , and Latin , ) is a genus of ctenochasmatid Pterodactyloidea, pterodactyloid pterosaur from South America. Its fossil remains dated back to the Early Cretaceous period, about 105 million years ago. ...
guinazui'' is published by Codorniú, Paulina-Carabajal & Gianechini (2016). * A small
azhdarchoid Azhdarchoidea (International Phonetic Alphabet, , meaning "azhdarchid-like forms") is a group of pterosaurs within the suborder Pterodactyloidea. Pterosaurs belonging to this group lived throughout the Early Cretaceous, Early and Late Cretaceous ...
, possibly an
azhdarchid Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cre ...
, is described from the Late Cretaceous (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
)
Northumberland Formation The Northumberland Formation is a Late Cretaceous (?Campanian-?Maastrichtian)-aged geologic Formation (geology), formation in Canada. It belongs to the larger Nanaimo Group. Indeterminate bird and pterosaur fossils have been recovered from the for ...
(
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...
,
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) by Martin-Silverstone ''et al.'' (2016). * Kellner ''et al.'' (2016) redescribe the first pterosaur remains from Japan, referring it to a
pteranodontid The Pteranodontidae are a family of large pterosaurs from the Late Cretaceous of North America and possibly other continents including Europe and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elon ...
-like pterosaur and indicating that it is the largest pterosaur recorded from Asia so far.


New taxa


References

{{Reflist, 30em 2010s in paleontology P