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The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period is divided in the
geologic time scale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geoch ...
.
Rock strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms wh ...
known as
chalk Chalk is a soft, white, porous, sedimentary carbonate rock. It is a form of limestone composed of the mineral calcite and originally formed deep under the sea by the compression of microscopic plankton that had settled to the sea floor. Cha ...
. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period.


Climate

During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred ...
became restricted to equatorial regions and northern
latitude In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north po ...
s experienced markedly more seasonal climatic conditions.


Geography

Due to
plate tectonics Plate tectonics (from the la, label= Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large t ...
, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient se ...
divided North America into eastern and western halves;
Appalachia Appalachia () is a cultural region in the Eastern United States that stretches from the Southern Tier of New York State to northern Alabama and Georgia. While the Appalachian Mountains stretch from Belle Isle in Newfoundland and Labrador, ...
and
Laramidia Laramidia was an island continent that existed during the Late Cretaceous period (99.6–66 Ma), when the Western Interior Seaway split the continent of North America in two. In the Mesozoic era, Laramidia was an island land mass separated from ...
. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...
seem to have remained connected and began to drift away from Africa and South America."Dinosaurs Ruled the World: Late Cretaceous Period." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. ''The Age of Dinosaurs''. Publications International, LTD. Pp. 103-104. . Europe was an island chain. Populating some of these islands were endemic dwarf dinosaur species.


Vertebrate fauna


Non-avian dinosaurs

In the Late Cretaceous, the hadrosaurs,
ankylosaurs Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
, and
ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassi ...
ns experienced success in Asiamerica (Western North America and eastern Asia). Tyrannosaurs dominated the large predator niche in North America. They were also present in Asia, although were usually smaller and more primitive than the North American varieties. Pachycephalosaurs were also present in both North America and Asia.
Dromaeosaur Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
s shared the same geographical distribution, and are well documented in both Mongolia and Western North America. Additionally therizinosaurs (known previously as segnosaurs) appear to have been in North America and Asia.
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final st ...
held a very different dinosaurian fauna, with most predators being abelisaurs and carcharodontosaurs; and
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...
s being among the dominant herbivores. Spinosaurids were also present during this time.


Birds

Birds became increasingly common, diversifying in a variety of
enantiornithe The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cl ...
and ornithurine forms. Early
Neornithes Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
such as '' Vegavis''Supporting information
/ref> co-existed with forms as bizarre as '' Yungavolucris'' and ''
Avisaurus ''Avisaurus'' (meaning "bird lizard") is a genus of Enantiornithes, enantiornithine bird from the Late Cretaceous of North America. Discovery ''Avisaurus archibaldi'' was discovered in the Late Cretaceous Hell Creek Formation of North America (M ...
''. Though mostly small, marine Hesperornithes became relatively large and flightless, adapted to life in the open sea.


Pterosaurs

Though primarily represented by
azhdarchids Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early ...
, other forms like
pteranodontids The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head ( ...
,
tapejarids Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The mo ...
('' Caiuajara'' and ''
Bakonydraco ''Bakonydraco'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period (Santonian stage) of what is now the Csehbánya Formation of the Bakony Mountains, Iharkút, Veszprém, western Hungary. Etymology ''Bakonydraco'' was name ...
''), nyctosaurids and uncertain forms ('' Piksi'', ''
Navajodactylus ''Navajodactylus'' (meaning "Navajo finger") is an extinct genus of pterodactyloid pterosaur from Late Cretaceous (late Campanian stage) deposits of the San Juan Basin in New Mexico, United States. Discovery The holotype specimen of ''Navajodac ...
'') are also present. Historically, it has been assumed that pterosaurs were in decline due to competition with birds, but it appears that neither group overlapped significantly ecologically, nor is it particularly evident that a true systematic decline was ever in place, especially with the discovery of smaller pterosaur species.


Mammals

Several old mammal groups began to disappear, with the last eutriconodonts occurring in the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
of North America. In the northern hemisphere, cimolodont,
multituberculates Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, ...
,
metatheria Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as we ...
ns and
eutherian Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic t ...
s were the dominant mammals, with the former two groups being the most common mammals in North America. In the southern hemisphere there was instead a more complex fauna of dryolestoids, gondwanatheres and other multituberculates and basal
eutherian Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic t ...
s;
monotreme Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals ( Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their bra ...
s were presumably present, as was the last of the haramiyidans, ''Avashishta''. Mammals, though generally small, ranged into a variety of ecological niches, from carnivores ( Deltatheroida), to mollusc-eater (
Stagodontidae Stagodontidae is an extinct family of carnivorous metatherian mammals that inhabited North America and Europe during the late Cretaceous, and possibly to the Eocene in South America. Description Currently, the family includes four genera, '' ...
), to herbivores (multituberculates, '' Schowalteria'', Zhelestidae and Mesungulatidae) to highly atypical cursorial forms ( Zalambdalestidae, Brandoniidae). True
placental Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguishe ...
s evolved only at the very end of the epoch; the same can be said for true
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
s. Instead, nearly all known eutherian and metatherian fossils belong to other groups.


Marine life

In the seas,
mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Ma ...
s suddenly appeared and underwent a spectacular evolutionary radiation. Modern sharks also appeared and giant-penguin-like
polycotylid Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. ...
plesiosaurs (3 meters long) and huge long-necked elasmosaurs (13 meters long) also diversified. These predators fed on the numerous
teleost Teleostei (; Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts ), is, by far, the largest infraclass in the class Actinopterygii, the ray-finned fishes, containing 96% of all extant species of fish. Tel ...
fishes, which in turn evolved into new advanced and modern forms ( Neoteleostei).
Ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...
s and
pliosaur Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive t ...
s, on the other hand, became extinct during the Cenomanian-Turonian anoxic event.


Flora

Near the end of the Cretaceous Period,
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s diversified. In temperate regions, familiar plants like
magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendro ...
s,
sassafras ''Sassafras'' is a genus of three extant and one extinct species of deciduous trees in the family Lauraceae, native to eastern North America and eastern Asia.Wolfe, Jack A. & Wehr, Wesley C. 1987. The sassafras is an ornamental tree. "Middle E ...
,
rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can b ...
s,
redwood Sequoioideae, popularly known as redwoods, is a subfamily of coniferous trees within the family Cupressaceae. It includes the largest and tallest trees in the world. Description The three redwood subfamily genera are '' Sequoia'' from coas ...
s, and
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist ...
s could be found in abundance.


Cretaceous–Paleogene mass extinction discovery

The Cretaceous–Paleogene extinction event was a large-scale
mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. I ...
of animal and plant species in a geologically short period of time, approximately (Ma). It is widely known as the ''K–T extinction event'' and is associated with a geological signature, usually a thin band dated to that time and found in various parts of the world, known as the
Cretaceous–Paleogene boundary The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end ...
(K–T boundary). ''K'' is the traditional abbreviation for the Cretaceous Period derived from the German name ''Kreidezeit'', and ''T'' is the abbreviation for the
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non- avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
Period (a historical term for the period of time now covered by the
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
and
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago (Mya) to the beginning of the present Quaternary Period Mya. ...
periods). The event marks the end of the
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ...
Era and the beginning of the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...
Era. "Tertiary" being no longer recognized as a formal time or rock unit by the
International Commission on Stratigraphy The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigr ...
, the K-T event is now called the Cretaceous—Paleogene (or K-Pg) extinction event by many researchers. Non- avian
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
are found only below the Cretaceous–Paleogene boundary and became extinct immediately before or during the event. A very small number of dinosaur fossils have been found above the Cretaceous–Paleogene boundary, but they have been explained as reworked fossils, that is, fossils that have been eroded from their original locations then preserved in later
sedimentary Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particles ...
layers.
Mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Ma ...
s, plesiosaurs,
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the Order (biology), order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cre ...
s and many
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of plants and
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s also became extinct.
Mammalian Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
and bird clades passed through the boundary with few extinctions, and
evolutionary radiation An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. Radiations may affect one clade or many, and be rap ...
from those
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
clades occurred well past the boundary. Rates of extinction and radiation varied across different clades of organisms. Many scientists hypothesize that the Cretaceous–Paleogene extinctions were caused by catastrophic events such as the massive
asteroid impact An impact event is a collision between astronomical objects causing measurable effects. Impact events have physical consequences and have been found to regularly occur in planetary systems, though the most frequent involve asteroids, comets or me ...
that caused the
Chicxulub crater The Chicxulub crater () is an impact crater buried underneath the Yucatán Peninsula in Mexico. Its center is offshore near the community of Chicxulub, after which it is named. It was formed slightly over 66 million years ago when a large ast ...
, in combination with increased
volcanic activity Volcanism, vulcanism or volcanicity is the phenomenon of eruption of molten rock (magma) onto the surface of the Earth or a solid-surface planet or moon, where lava, pyroclastics, and volcanic gases erupt through a break in the surface called a ...
, such as that recorded in the
Deccan Traps The Deccan Traps is a large igneous province of west-central India (17–24°N, 73–74°E). It is one of the largest volcanic features on Earth, taking the form of a large shield volcano. It consists of numerous layers of solidified flood ...
, both of which have been firmly dated to the time of the extinction event. In theory, these events reduced sunlight and hindered
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
, leading to a massive disruption in Earth's
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...
. A much smaller number of researchers believe the extinction was more gradual, resulting from slower changes in
sea level Mean sea level (MSL, often shortened to sea level) is an average surface level of one or more among Earth's coastal bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical datuma standardise ...
or
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologica ...
.


See also

* Flora and fauna of the Maastrichtian stage * Hațeg Island


References

{{Geological history, p, m Late Geological epochs