
This list of informally named dinosaurs is a listing of dinosaurs (excluding
Aves
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight ...
; birds and their extinct relatives) that have never been given formally published
scientific names. This list only includes names that were not properly published ("
unavailable names") and have not since been published under a valid name (see
list of dinosaur genera for valid names). The following types of names are present on this list:
* ''
Nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
'', Latin for "naked name": A name that has appeared in print but has not yet been formally published by the standards of the
International Commission on Zoological Nomenclature
The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries.
Orga ...
. ''Nomina nuda'' (the plural form) are invalid, and are therefore not italicized as a proper generic name would be.
* ''
Nomen manuscriptum'', Latin for "manuscript name": A name that appears in manuscript but was not formally published. A ''nomen manuscriptum'' is equivalent to a ''nomen nudum'' for everything except the method of publication, and description.
* ''
Nomen ex dissertationae'', Latin for "dissertation name": A name that appears in a dissertation but was not formally published.
* Nicknames or descriptive names given to specimens or taxa by researchers or the press.
A
Alamotyrannus
"Alamotyrannus" ("Ojo Alamo tyrant") is the informal placeholder name given to an as yet undescribed genus or species of
tyrannosaurid from the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period of North America. The fossils of this animal originate from the
Ojo Alamo Formation
The Ojo Alamo Formation is a geologic Formation (geology), formation in New Mexico spanning the Mesozoic/Cenozoic boundary. Non-avian dinosaur fossils have controversially been identified in beds of this formation dating from after the Cretaceous ...
in
New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
and they were discovered during the early 2000s. The suggested binomial "Alamotyrannus brinkmani", was created when the paper describing the genus was written in 2013.
[Dalman, S.G. and S.G. Lucas. (2013). A new large Tyrannosaurid ''Alamotyrannus brinkmani'', n. gen., n. sp. (Theropoda: Tyrannosauridae), from the Upper Cretaceous Ojo Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico. New Mexico Museum of Natural History and Science Bulletin] "Alamotyrannus" lived during the early
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
.
Specimen ACM 7975, a jaw discovered in the
Ojo Alamo Formation
The Ojo Alamo Formation is a geologic Formation (geology), formation in New Mexico spanning the Mesozoic/Cenozoic boundary. Non-avian dinosaur fossils have controversially been identified in beds of this formation dating from after the Cretaceous ...
,
New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
in 1924, has been tentatively identified as ''
Gorgosaurus libratus
''Gorgosaurus'' ( ; ) is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period (geology), Period (Campanian), between about 76.5 and 75 million years ago. Fossil remains have been ...
'' but may instead belong to "Alamotyrannus" as per Dalman & Lucas (2013)
and McDavid (2022).
This specimen has been mentioned in a 2016 publication by Dalman and Lucas as an indeterminate tyrannosaurid without generic attribution, and it's noted that the specimen is under study by the senior author. Photograph taken by McDavid (2022) shows the specimen on display in the
Beneski Museum of Natural History.
Alan the Dinosaur

"Alan the Dinosaur" is the name given to a sauropod caudal vertebra (YORYM:2001.9337) found in 1995 in the
Saltwick Formation (
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
,
Aalenian) of Whitby, England. It is the oldest sauropod found in the United Kingdom, dating back 176-172 million years ago. Its name references that of its discoverer, Alan Gurr, and the fact that it is not identifiable to species level. An analysis done in 2015 found that it was a member of Eusauropoda, could be excluded from Diplodocoidea, and was most similar to ''
Cetiosaurus
''Cetiosaurus'' ( meaning 'whale lizard', from the Greek '/ meaning 'sea monster' (later, 'whale') and '/ meaning 'lizard'), is a genus of herbivorous sauropod dinosaur from the Middle Jurassic Period, living about 171 to 165 million years ago ...
''.
The fossil of "Alan" is housed in the
Yorkshire Museum
The Yorkshire Museum is a museum in York, England. It was opened in 1830, and has five permanent collections, covering biology, geology, archaeology, numismatics and astronomy.
History
The museum was founded by the Yorkshire Philosophical Soci ...
, where it forms part of the Yorkshire's Jurassic World exhibit, featuring a VR recreation.
Allosaurus robustus

"Allosaurus robustus" is an informal name used for specimen "NMV P150070", a theropod
astragalus
Astragalus may refer to:
* ''Astragalus'' (plant), a large genus of herbs and small shrubs
*Astragalus (bone)
The talus (; Latin for ankle or ankle bone; : tali), talus bone, astragalus (), or ankle bone is one of the group of foot bones known ...
known from the
Wonthaggi Formation (
Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
) of Victoria, Australia. When first studied, it was thought to have belonged to a species of ''
Allosaurus
''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
''.
Samuel Welles challenged this identification as he thought that the astragalus belonged to an ornithomimid, but the original authors defended their classification. Sometime in the early 2000s, Daniel Chure examined the bone and found that it did not represent a new species of ''Allosaurus'', but could still represent an allosauroid. At the same time, Yoichi Azuma and
Phil Currie noted that the astragalus resembled that of their new genus ''
Fukuiraptor''.
It may well represent a theropod related to ''
Australovenator'', though some argue that it could represent an abelisauroid. A 2019 study strongly supported a
megaraptoran affinity for the astragalus.
The name "Allosaurus robustus", first confined as a museum label, was first published by Chure in 2000.
Amargastegos
"Amargastegos" is an informal genus of
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
stegosaurid ornithischian dinosaur known from the
La Amarga Formation of Argentina, named by Roman Ulansky in 2014 on the basis of MACN N-43 (some dorsal
osteoderms
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of Extant taxon, extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, Temnospondyli, ...
, the cervical and caudal vertebrae, and one skull bone), and the
type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is "A. brevicollum".
[ In 2016, Peter Malcolm Galton and Kenneth Carpenter declared it a ''nomen nudum'', establishing it as an indeterminate stegosaur.]
Amphicoelias brontodiplodocus/Barackosaurus
"Barackosaurus" is the informal name created in 2010 which is used for a sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
found in Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
-aged sediments pertaining to the Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
, Wyoming
Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
. It was found in the Dana Quarry and "Barackosaurus" was supposedly 20 meters long and weighed 20 tons. In 2010, an article was made available, but not formally published, by Henry Galiano and Raimund Albersdorfer in which they dubbed the Dana Quarry specimens which had already been referred to as "Barackosaurus" as "Amphicoelias brontodiplodocus". The specific name referred to their hypothesis based on these specimens that nearly all Morrison diplodocid species are either growth stages or represent sexual dimorphism among members of the genus '' Amphicoelias'', but this analysis was met with skepticism and the publication itself has been disclaimed by its lead author, explaining that it is "obviously a drafted manuscript complete with typos, etc., and not a final paper. In fact, no printing or distribution has been attempted". As of 2015, they are now on display at the Lee Kong Chian Natural History Museum in Singapore.
Andhrasaurus
"Andhrasaurus" is an informal genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
armored ornithischian dinosaur from the Kota Formation of India. The proposed species is "A. indicus". Ulansky (2014) coined the name for skull elements, about 30 osteoderms
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of Extant taxon, extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, Temnospondyli, ...
, and the extremities of vertebrae and limbs, all preserved in the collections of the GSI and assigned to Ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs ...
by Nath et al. (2002). In 2016, Peter Malcolm Galton and Kenneth Carpenter noted that "Andhrasaurus" did not meet ICZN requirements and therefore declared it a ''nomen nudum'', listing it as Thyreophora
Thyreophora ("shield bearers", often known simply as "armored dinosaurs") is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous.
Thyreophorans are characterized by the presence of bod ...
indet., while noting that the jawbones described by Nath et al. (2002) belonging to crocodylomorphs.[Galton, Peter M. & Carpenter, Kenneth, 2016, "The plated dinosaur ''Stegosaurus longispinus'' Gilmore, 1914 (Dinosauria: Ornithischia; Upper Jurassic, western U.S.), type species of ''Alcovasaurus'' n. gen.", ''Neues Jahrbuch für Geologie und Paläontologie – Abhandlungen'' 279(2): 185–208] The dermal armor informally named "Andhrasaurus" was redescribed by Galton (2019), referring the material to Ankylosauria.
Angeac ornithomimosaur
The "Angeac ornithomimosaur" is an informal name given to an unnamed ornithomimosaur taxon known from the Early Cretaceous (previously thought to be Hauterivian
The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
-Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
in age, but now thought to be Berriasian
In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
aged[Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al]
Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
Geodiversitas, Museum National d'Histoire Naturelle Paris, In press. ffhal-03264773f) Angeac-Charente bonebed (part of the stratigraphy of the Aquitaine Basin
The Aquitaine Basin is the second largest Mesozoic and Cenozoic sedimentary basin in France after the Paris Basin, occupying a large part of the country's southwestern quadrant. Its surface area covers 66,000 km2 onshore. It formed on Varisca ...
) near Angeac-Charente in western France. The taxon is toothless and is known from numerous disarticulated remains representing at least 70 individuals covering almost all of the skeleton, some remains were described by Allain et al. (2014).
Angloposeidon
"Angloposeidon" is the informal name given to a sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from the Early Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
(Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) Wessex Formation
The Wessex Formation is a fossil-rich England, English geological formation that dates from the Berriasian to Barremian Stage (stratigraphy), stages of the Early Cretaceous. It forms part of the Wealden Group and underlies the younger Vectis Form ...
of the Isle of Wight
The Isle of Wight (Help:IPA/English, /waɪt/ Help:Pronunciation respelling key, ''WYTE'') is an island off the south coast of England which, together with its surrounding uninhabited islets and Skerry, skerries, is also a ceremonial county. T ...
in southern England. It was a possible brachiosaurid but has not been formally named. Darren Naish, a notable vertebrate palaeontologist, has worked with the specimen and has recommended that this name only be used informally and that it not be published. However, he published it himself in his book ''Tetrapod Zoology Book One'' from 2010. The remains consist of a single cervical vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
(MIWG.7306), which indicate it was a very large animal, 20 metres or greater in length.
Angustungui
"Angustungui" is an informal genus of stegosaur from the Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
Qigu Formation in China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. The intended type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is "A. qiketaiensis" and the holotype is specimen SS V16001 consisting of axial, pectoral girdle, pelvic girdle, limb and armor elements, while the paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype (biology), isotype ...
is specimen SS V16002, consisting of a right coracoid and a right scapula. Both specimens were discovered in 2016 by Li Daqing at Qiketai, Shanshan County, and the name was first announced in a ''bioRxiv'' pre-print article in October 2024, where it was recovered as sister taxon of '' Loricatosaurus''.
Archaeoraptor
"Archaeoraptor" is the informal generic name for an important fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
from China that was later discovered to have been fabricated from multiple unrelated fossils. The name was created in an article published in ''National Geographic'' magazine in 1999, where the magazine claimed that the fossil was a " missing link" between birds and terrestrial theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaurs. Even prior to this publication there had been severe doubts about the fossil's authenticity. Further scientific study showed it to be a forgery constructed from rearranged pieces of real fossils from different species. Zhou et al. found that the head and upper body actually belong to a specimen of the primitive fossil bird ''Yanornis
''Yanornis'' () is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: ''Yanornis martini'', based on several fossils found in the 120-million-year-old Jiufotang Formation ...
'', and another 2002 study found that the tail belongs to a small winged dromaeosaur, ''Microraptor
''Microraptor'' (Greek language, Greek, μικρός, ''mīkros'': "small"; Latin language, Latin, ''raptor'': "one who seizes") is a genus of small, four-winged dromaeosaurid dinosaurs. Numerous well-preserved fossil specimens have been recovere ...
'', named in 2000. The legs and feet belong to an as yet unknown animal.
Archbishop
"The Archbishop" is a giant brachiosaurid sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur similar to ''Brachiosaurus
''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about . It was first Species description, described by American paleontologist Elmer S. Riggs in 1903 in paleontology, 1903 from fossi ...
'' and ''Giraffatitan
''Giraffatitan'' (name meaning "titanic giraffe") is a genus of sauropod dinosaur that lived during the late Jurassic Period (geology), Period (Kimmeridgian–Tithonian stages) in what is now Lindi Region, Tanzania. Only one species is known, ...
''. It was long considered a specimen of ''Brachiosaurus'' (now ''Giraffatitan'') ''brancai'' due to being found in the same formation in Tendaguru
The Tendaguru Formation, or Tendaguru Beds are a highly fossiliferous formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represents the oldest sedimentary unit of the Mandawa Basin, overlying Neoprote ...
, Tanzania. However, the "Archbishop" shows significant differences including a unique vertebral morphology and a proportionally longer neck, that indicates it is a different, previously unknown genus and species. It was discovered by Frederick Migeod in 1930. "The Archbishop" is a nickname that functions as a placeholder – the specimen currently has no scientific name. The specimen is currently housed in the Natural History Museum in London, and will eventually be re-described by Dr. Michael P. Taylor
Michael Paul Taylor (born 12 March 1968) is a British computer programmer with a Ph.D. in palaeontology. To date, he has published 31 paleontological papers and is co-credited with naming three genera of dinosaur (''Xenoposeidon'' in 2007 with Da ...
of Bristol University
The University of Bristol is a public research university in Bristol, England. It received its royal charter in 1909, although it can trace its roots to a Merchant Venturers' school founded in 1595 and University College, Bristol, which had ...
. In May 2018, Taylor started to work on describing the Archbishop.
Atlantohadros
"Atlantohadros", more commonly known as the "Merchantville hadrosaur", is an informally named hadosaurid dinosaur that lived in the Merchantville Formation in the northeastern United States. Brownstein (2021) found "Atlantohadros" to be more derived than ''Tethyshadros
''Tethyshadros'' ("Tethys Ocean, Tethyan hadrosauroid") is a genus of Hadrosauroidea, hadrosauroid dinosaur from the Late Cretaceous (Campanian) Calcare di Aurisina (previously thought to come from the younger Liburnia Formation) of Trieste, Ita ...
'' but less derived than Saurolophinae
Saurolophinae is a subfamily (biology), subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. How ...
and Lambeosaurinae. The name was intended to be used in that publication, but was cut for unknown reasons; initial versions of Brownstein (2021) contained the word "Atlantohadros" superimposed over "Merchantville Taxon" in a cladogram; subsequent corrections have erased the genus name entirely.[
Three specimens were discovered northwest of Freehold near Manalapan–Marlboro township line in Monmouth County during the 1970s. These are: YPM VPPU.021813, YPM VPPU.021813, and AMNH 13704, with YPM VPPU.021813 possibly belonging to the same individual as YPM VPPU.021813 due similar weathering, size and the same horizon. These specimens consist of both coracoids, both scapulae, a femur, a fragmentary proximal tibia, and a dentary from a cast of the specimen (the original likely lost in YPM's catalogue) in the adult specimen, as well as a rib, a femur and long bone portions in the juvenile. AMNH 13704, id a partial dentary of a probable perinate. Scattered bones associated with these include a quadrate, several partial maxilla portions, a partial jugal, skull roof fragments and several rib fragments.]
B
Baguasaurus
"Baguasaurus" (meaning "Bagua
The ''bagua'' ( zh, c=八卦, p=bāguà, l=eight trigrams) is a set of symbols from China intended to illustrate the nature of reality as being composed of mutually opposing forces reinforcing one another. ''Bagua'' is a group of trigrams—co ...
lizard") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' an ...
n sauropod dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
– Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-aged) Chota Formation of Peru. The proposed holotype, consisting of caudal vertebrae, was first mentioned in a review of the Chota Formation by Mourier et al. (1988), and the name "Baguasaurus" was coined by Larramendi & Molina Pérez (2020). "Baguasaurus" was estimated to be long and weighed .
Balochisaurus
"Balochisaurus" (meaning "Balochi lizard", for the Baloch tribes of Pakistan) is an informal taxon of titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
ian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Pakistan. The proposed species is "B. malkani". The discovery was made (along with other dinosaur specimens) in 2001 near Vitariki by a team of paleontologists from the Geological Survey of Pakistan. Described in 2006 by M.S. Malkani, the genus is based on seven tail vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e found in the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age Vitakri Member of the Pab Formation, with additional vertebrae and a partial skull assigned to it. ''Balochisaurus'' was assigned to the family "Balochisauridae" along with " Marisaurus". It was considered invalid by Wilson, Barrett and Carrano (2011).
Barnes High Sauropod
The "Barnes High sauropod" is the informal name given to MIWG-BP001, an undescribed sauropod dinosaur specimen from the Wessex Formation
The Wessex Formation is a fossil-rich England, English geological formation that dates from the Berriasian to Barremian Stage (stratigraphy), stages of the Early Cretaceous. It forms part of the Wealden Group and underlies the younger Vectis Form ...
on the Isle of Wight. It was discovered in the cliffs around Barnes High in 1992 and is currently owned by the privately run unaccredited Dinosaur Farm Museum near Brighstone, the ownership situation was described as "complex" and the specimen is currently inaccessible to researchers. It is roughly 40% complete and consists of a "Partial postcranial skeleton, including presacral vertebrae, anterior caudal vertebrae, girdle and limb elements" including a largely complete forelimb. It has been suggested to be a brachiosaurid and is possibly synonymous with the earlier named '' Eucamerotus'' due to similarities with the vertebrae.
Bayosaurus
"Bayosaurus" is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of theropod dinosaur. The name was coined by paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
s Rodolfo Coria, Philip J. Currie, and Paulina Carabajal in 2006
2006 was designated as the International Year of Deserts and Desertification.
Events
January
* January 1– 4 – Russia temporarily cuts shipment of natural gas to Ukraine during a price dispute.
* January 12 – A stampede during t ...
. It apparently was an abelisauroid from the Turonian
The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
Cerro Lisandro Formation of Neuquén
Neuquén (; ) is the capital city of the Argentine province of Neuquén and of the Confluencia Department, located in the east of the province. It occupies a strip of land west of the confluence of the Limay and Neuquén rivers which form t ...
, Argentina, around long. The specimen is MCF-PVPH-237, including dorsal and sacral vertebrae
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
, a fragmentary pelvis
The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...
, and other partial bones, which were discovered in 2000. The name was used in a phylogenetic analysis to indicate the position of MCF-PVPH-237.
Beelemodon
"Beelemodon" is the informal name given to an undescribed theropod genus from the Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
, possibly belonging to a coelurosaur. The fossils include two teeth found in Wyoming
Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
, United States. The name appeared in print in 1997
Events January
* January 1 – The Emergency Alert System is introduced in the United States.
* January 11 – Turkey threatens Cyprus on account of a deal to buy Russian S-300 missiles, prompting the Cypriot Missile Crisis.
* January 1 ...
, when paleontologist Robert T. Bakker mentioned it in a symposium for the Academy of Natural Sciences.[Bakker, R. ''Raptor family values: Allosaur parents brought great carcasses into their lair to feed their young.'' In "Dinofest International", Proceedings of a Symposium, Academy of Natural Sciences, eds Wolberg, Sump and Rosenberg, 51 – 63 (1997).] The teeth are most similar to ''Compsognathus
''Compsognathus'' (; Ancient Greek, Greek ''kompsos''/κομψός; "elegant", "refined" or "dainty", and ''gnathos''/γνάθος; "jaw") is a genus of small, bipedalism, bipedal, carnivore, carnivorous theropoda, theropod dinosaur. Members o ...
'', but have no unique features and also share similarities with '' Protarchaeopteryx'' and dromaeosaurids.
Biconcavoposeidon
"Biconcavoposeidon" is the placeholder name for AMNH FARB 291, five consecutive posterior dorsal vertebrae of a brachiosaurid sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
, from the Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
, Wyoming
Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
. Not much else is currently known about "Biconcaveoposeidon", except that it was discovered in the Bone Cabin quarry in 1898.
Bihariosaurus
"Bihariosaurus" (meaning "Bihor lizard") is an invalid genus of iguanodontian dinosaur from Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Bauxite of Cornet, Romania. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, "Bihariosaurus bauxiticus", was named but not described by Marinescu in 1989. It was similar to ''Camptosaurus
''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic Period (geology), period of western North America and possibly also Europe. The name means 'flexible lizard' (Ancient Greek, Greek (') meaning ...
'', and was an iguanodont
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivore, herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and rel ...
. The original publication of the taxon did not include sufficient description, and the illustrations cannot distinguish it from any other ornithopod.
Biscoveosaurus
"Biscoveosaurus" is the informal name of an ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
dinosaur specimen from the Early Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
age Snow Hill Island Formation of James Ross Island
James Ross Island () is a large island off the southeast side and near the northeastern extremity of the Antarctic Peninsula, from which it is separated by Prince Gustav Channel.
Rising to , it is irregularly shaped and extends in a north–so ...
, Antarctica. It comes from the Cape Lamb Member of the formation, the same member as '' Morrosaurus'', another basal ornithopod. As such, it's been suggested it may be a secondary specimen of that species, but as the holotype of ''Morrosaurus'' is fragmentary and doesn't overlap with the material of "Biscoveosaurus", this can't as yet be tested. The specimen consists of dentaries, teeth, a braincase, parts of the maxillae, forelimb elements, assorted vertebrae, and the pectoral girdle; this makes it unique compared to the other James Ross Island ornithopods, which do not have both cranial and postcranial remains. It has been estimated the animal would have been about in length.
C
Capitalsaurus
"Capitalsaurus" is the informal genus name given to a tailbone
The coccyx (: coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other ...
belonging to a large theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur that lived during the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
. It was discovered on 28 January 1898, by construction workers excavating a sewer at the intersection of Washington, D.C.'s First and F Streets SE. The only known specimen, it was assigned two different species designations – ''Creosaurus potens'' and ''Dryptosaurus potens'' – and eventually overturned each time. In the 1990s, the paleontologist Peter Kranz asserted that it represented a unique type of dinosaur and assigned it the name "Capitalsaurus". He successfully campaigned through local schools to make "Capitalsaurus" the official dinosaur of Washington, D.C., which became law in 1998. A year later, the district further recognized F Street at the discovery site as Capitalsaurus Court. It designated 28 January 2001, as Capitalsaurus Day.
Changdusaurus
"Changdusaurus" (also known as "Changtusaurus") is the informal name given to a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
Period. It lived in what is now China. "Changdusaurus" is classified as a stegosaurid. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was named "Changdusaurus laminoplacodus" by Zhao in 1983, but it has never been formally described, and remains a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. One source indicates the fossils have been lost.
Cinizasaurus
"Cinizasaurus" is a ''nomen ex dissertationae'' for fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ized remains from the Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
of New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
that were initially interpreted as belonging to a theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur. The intended type species "Cinizasaurus hunti" was named in an unpublished 1997 thesis based on NMMNH P-18400, which consists of a tibia, vertebrae, and other fragments. In 2007, Nesbitt, Irmis and Parker classified both NMMNH P-18400 and NMMNH P-18401 as specimens of an indeterminate archosauriform.
Comanchesaurus
"Comanchesaurus" is a ''nomen ex dissertationae'' for fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ized remains from the Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
of New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
that were initially interpreted as belonging to a theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur. The remains, NMMNH P-4569, consist of a partial skeleton including vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
l centra and hindlimb bones, and came from the Norian
The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
-age Upper Triassic
The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...
Bull Canyon Formation of Guadalupe County. Adrian Hunt, in his unpublished dissertation, proposed the name "Comanchesaurus kuesi" for the specimen, but the name was never adopted, and was first referred to in the scientific literature in a 2007 redescription of Late Triassic North American material thought to belong to dinosaurs (Nesbitt, Irmis, and Parker, 2007). In the redescription, the authors found the material to belong to a "possible indeterminate saurischia
Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithi ...
n".
Cryptoraptor
"Cryptoraptor" is a ''nomen ex dissertationae'' for fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ized remains from the Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
of New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
that were initially interpreted as belonging to a theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur. The remains, NMMNH P-17375, consist of a partial skeleton including partial hindlimb and pelvic bones, and came from the Norian
The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
-age Upper Triassic
The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...
Bull Canyon Formation of Quay County. Adrian Hunt, in his unpublished dissertation, proposed the name "Cryptoraptor lockleyi" for the specimen, but the name was never adopted, and was first referred to in the scientific literature in a 2007 redescription of Late Triassic North American material thought to belong to dinosaurs. In the redescription, the authors found the material to belong to an intermediate archosaur, as no features exclusive to dinosaurs could be identified.[
]
Cryptotyrannus
"Cryptotyrannus" (meaning "secret/hidden tyrant"), more commonly known as the "Merchantville tyrannosauroid", is an informally named tyrannosauroid dinosaur that lived in the Merchantville Formation. It was informally named by Brownstein (2021), who found it to be the sister taxon of '' Dryptosaurus'', reinstating Dryptosauridae. The name appeared in the initial version of Brown's paper, superimposed over "Merchantville Taxon" in a cladogram; a subsequent correction has erased the name entirely.
"Cryptotyrannus" is known from two specimens discovered during the 1970s, the holotype YPM VPPU.021795 and the paratype YPM VPPU.022416. Similar coloration and weathering indicate that these are probably the same individual. These are a partial foot bone and one caudal vertebrae. However, a skeletal produced for the paper depicts a hand claw. The foot morphology is consistent with tyrannosaurs, being extremely similar to the '' Dryptosaurus aquilunguis''. Autapomorphies include a metatarsal IV that is far more gracile and IV in proximal view also has a triangular, rather than subrectangular in outline. The holotype was once tentatively assigned to '' "Coelosaurus" antiquus''. Shark bites present on the holotype suggest that the specimen's fragmentary nature is due to predation or scavenging by marine predators.
D
Dachongosaurus
"Dachongosaurus" is the informal name given to an undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from the Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
of China. It is known from fossils including at least a partial articulated skeleton from the Dark Red Beds of the Lower Lufeng Series (Sinemurian
In the geologic timescale, the Sinemurian is an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between 199.5 ±0.3 annu ...
stage) in Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
.[Zhao, 1985. The reptilian fauna of the Jurassic in China. Pages 286–289, 347 in Wang, Cheng and Wang (eds.). The Jurassic System of China. Geological Publishing House, Beijing.] Possibly a cetiosaur, the "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
" is "Dachongosaurus yunnanensis", coined by Zhao in 1985. An alternate spelling is "Dachungosaurus". As with other informal names coined by Zhao in 1985 and 1983, nothing has since been published, and the remains may have been redescribed under another name.
Damalasaurus
"Damalasaurus" (meaning "Damala lizard") is the informal name given to a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of herbivorous
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
dinosaur from the Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
. It was a sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
, though its exact classification within the clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
is unknown. Fossils of "Damalasaurus", including a rib, have been found in the Middle Daye Group of Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
. Species attributed to this genus include "Damalasaurus laticostalis" and "D. magnus", although it is possible that both names refer to the same species.
Dongshengosaurus
"Dongshengosaurus" is the informal name given to an undescribed genus of iguanodontian dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
of Liaoning
)
, image_skyline =
, image_alt =
, image_caption = Clockwise: Mukden Palace in Shenyang, Xinghai Square in Dalian, Dalian coast, Yalu River at Dandong
, image_map = Liaoning in China (+all claims hatched).svg
, ...
, China. the "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
", "D. sinensis", was named by Pan Rui in his 2009 thesis. It is known from a partial juvenile skeleton discovered from the Yixian Formation
The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
.
Dubeynarainsaurus
"Dubeynarainsaurus" is an informal genus of purported theropod dinosaurs from the Late Cretaceous Lameta Formation of India described by Malkani (2025) in Scientific Research Publishing
Scientific Research Publishing (SCIRP) is a predatory academic publisher of open-access electronic journals, conference proceedings, and scientific anthologies that are of questionable quality. , it offered 244 English-language open-access journ ...
, a known predatory publisher. The proposed holotype is allegedly a partial dentary with associated teeth, collected in January 1944 by V. S. Dubey and Kedar Narain (who are referenced in the proposed genus name). The intended type species is "Dubeynarainsaurus sahni." The specimen was initially identified as a pterosaur mandible in 1946, although Malkani reclassified it as belonging to a noasaurid theropod.
Duranteceratops
"Duranteceratops" is a purported new taxon of chasmosaurine ceratopsid
Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', '' Centrosaurus'', and '' Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are ...
from the Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
.[ In 2012, a ceratopsid skull supposedly distinguishable from '']Triceratops
''Triceratops'' ( ; ) is a genus of Chasmosaurinae, chasmosaurine Ceratopsia, ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous Period (geology), period, about 68 to 66 million years ago on the island ...
'' was unearthed in South Dakota
South Dakota (; Sioux language, Sioux: , ) is a U.S. state, state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Dakota people, Dakota Sioux ...
by a fossil poacher named John Carter. Though it has yet to be published, according to the ''Prehistoric Times'' issue no. 121 from Spring 2017, the specimen is to be named "Duranteceratops".
E
Echizensaurus
"Echizensaurus" is an informal genus of ornithischian dinosaur from Echizen, Japan which lived during the Late Cretaceous. It has been assigned to Ceratopsia ''incertae sedis'' in paleontologist Lida Xing's homepage.
EK troodontid
The "EK troodontid" (specimen SPS 100/44) is an unnamed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of troodontid
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
dinosaur discovered in Mongolia. In the scientific literature it is referred to as the "EK troodontid", after the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
sediments in which it was found. SPS 100/44 was discovered by Sergei Mikhailovich Kurzanov during the 1979 Soviet-Mongolian Paleontological Expedition. It was found in deposits of the Barunbayaskaya Svita at the Khamareen Us locality, Dornogov (southeastern Gobi Desert), in the Mongolian People's Republic. SPS 100/44 was described by Rinchen Barsbold and colleagues in 1987.[Barsbold, Rhinchen, Osmolska, Halszka, Kurzanov, Sergei Mikhailovich. (1987). "On a new troodontid (Dinosauria. Theropoda) from the Early Cretaceous of Mongolia". ''Acta Palaeontologica Polonica'' 32(1–2): 121–132]
Its fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
remains include an incomplete skeleton consisting of the braincase, posterior parts of the lower mandibles, a maxillary fragment with teeth, parts of five cervical vertebrae (cervicals ?2-?6), an articulated right manus with partial semilunate, left manus phalanx
The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
I-1, distal end of the left femur, and fragmentary left and right pedes. Barsbold pointed out that the specimen was smaller and from older sediments than other known troodontids, but it had some features of the skull that could have made it a juvenile. Barsbold also indicated the high degree of fusion of the bones of the skull and the unusual foot morphology to indicate that it might be an adult of an unknown taxon. Barsbold took the conservative position and did not name this specimen because it was not complete enough to rule out the possibility that it was a juvenile of a known genus of troodontid. Barsbold also noted that the naturally articulated manus of SPS 100/44 showed no signs of an opposable third digit, as was suggested for '' Troodon'' by Russell and Seguin in 1982. Turner and colleagues, in 2007, found the EK troodontid to be a distinct basal genus of troodontid, in a polytomy with '' Jinfengopteryx'' and a clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
of more derived troodontids.
Eoplophysis
"Eoplophysis" is a genus of stegosaur known from the Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
Cornbrash Formation, Sharp's Hill Formation, and Chipping Norton Formation of England. It was originally named '' Omosaurus vetustus'' by the renowned German paleontologist Friedrich von Huene. The holotype, OUM J.14000, is a right femur of a juvenile individual from the Middle Jurassic (upper Bathonian
In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
) Cornbrash Formation of Oxfordshire
Oxfordshire ( ; abbreviated ''Oxon'') is a ceremonial county in South East England. The county is bordered by Northamptonshire and Warwickshire to the north, Buckinghamshire to the east, Berkshire to the south, and Wiltshire and Glouceste ...
, England, although it was probably reworked from the slightly older Forest Marble Formation in view of its eroded nature. Because of the renaming of ''Omosaurus'', an occupied name, as ''Dacentrurus'', ''O. vetustus'' was renamed into a ''Dacentrurus vetustus'' in 1964. In the 1980s, researcher Peter Malcolm Galton reviewed all known stegosaur material from the Bathonian of England and concluded that ''Omosaurus vetustus'' was valid and should be tentatively referred to '' Lexovisaurus''. However, the species was later considered a ''nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
'' in both reviews of Stegosauria. In their alpha-taxonomic review of stegosaurs, Susannah Maidment and her colleagues noted that OUM J.14000 shares characters present in both sauropods and stegosaurs, but that it lacks synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ...
exclusive to Stegosauria
Stegosauria is a group of Herbivore, herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous Period (geology), periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere (North America, Europe a ...
and assigned it as a Dinosauria indet. Nevertheless, the amateur paleontologist Roman Ulansky coined the new genus "Eoplophysis" ("Dawn Armed Form") for ''O. vetustus'', noting differences with the femora of other stegosaurs.[
]
Eugongbusaurus
"Eugongbusaurus" is the informal name (''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
'') proposed for a neornithischian found in the Oxfordian-age Shishugou Formation of Xinjiang
Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...
, China. The intended type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''"Gongbusaurus" wucaiwanensis'', was described by Dong Zhiming
Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
in 1989 for two partial skeletons as a second species of the poorly known tooth taxon
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
'' Gongbusaurus''. Fragmentary skeleton IVPP 8302, the type specimen
In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
for the new species, included a partial lower jaw, three tail vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e, and a partial forelimb. Second specimen IVPP 8303 consisted of two hip vertebrae, eight tail vertebrae, and two complete hind limbs. Dong estimated it as around long, and considered it to be a strong runner. He assigned the genus ''Gongbusaurus'' to the Hypsilophodontidae
Hypsilophodontidae (or Hypsilophodontia) is a traditionally used family (biology), family of ornithopod dinosaurs, generally considered invalid today. It historically included many small bodied bipedal neornithischian taxa from around the world, ...
, a paraphyletic grade of small herbivorous bipedal
Bipedalism is a form of terrestrial locomotion where an animal moves by means of its two rear (or lower) limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' ...
dinosaurs. Because dinosaur teeth are generally not distinctive enough to hold a name, it is unsurprising that other paleontologists have suggested removing "G." ''wucaiwanensis'' from ''Gongbusaurus'' and giving it its own genus. The possible replacement name "Eugongbusaurus" leaked out accidentally and remains informal.
F
Fendusaurus
"Fendusaurus" is a ''nomen ex dissertatione'' proposed by Fedak (2006) for FGM 998GF13-II, which includes a skull
The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate.
In the human, the skull comprises two prominent ...
. Other specimens referred to "Fendusaurus" are FGM998GF13-I, FGM998GF13-III, FGM998GF69, FGM998GF9, and FGM998GF18, all found by a crew from the Princeton University
Princeton University is a private university, private Ivy League research university in Princeton, New Jersey, United States. Founded in 1746 in Elizabeth, New Jersey, Elizabeth as the College of New Jersey, Princeton is the List of Colonial ...
. All the specimens include femora and coracoid
A coracoid is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is present as part of the scapula, but this is n ...
s, and although they each share slightly different features, the differences are credited to intra-specific variation. Known specimens of "Fendusaurus" were previously classified as cf. '' Ammosaurus''. The femora and coracoids also help identify different individuals, and Timothy J. Fedak, the describer of the specimens, found that each block represented about one individual. "Fendusaurus" is known from the Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
(Hettangian
The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triass ...
) McCoy Brook Formation of Wasson Bluff, Nova Scotia. It is the first non-avian dinosaur from Nova Scotia. As five specimens of "Fendusaurus" are from the McCoy Brook Formation, the formation is the richest prosauropod site in North America. The formation is also similar to other formations of North America and Asia, as it lacks any remains presently assigned to ''Anchisaurus''. Fedak places "Fendusaurus" as a genus of the family Massospondylidae
Massospondylidae is a family (biology), family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarc ...
.[Fedak TJ. Description and evolutionary significance of the sauropodomorph dinosaurs from the Early Jurassic (Hettangian) McCoy Brook Formation. PhD Dissertation, Dalhousie University. 2006.]
The specimens of "Fendusaurus" include mostly crushed vertebrae
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
, along with appendicular elements. They are distinguishable from ''Anchisaurus'' by the morphology of both the ilium and sacral vertebrae. However, in some specimens, the morphology of the femora and coracoids are quite different, which led Fedak to speculate that more than one species may have been present. "Fendusaurus", according to Fedak, can be distinguished from all closely related sauropodomorphs by the extreme elongation of the cervical vertebrae; a four vertebrae sacrum that includes a dorsosacral and caudosacral; the elongate postacetabular process of the ilium; and an expanded anterior distal process of the tibia.[
]
Ferganastegos
"Ferganastegos" is a dubious genus of stegosaur from the Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
(Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
) Balabansai Formation of Fergana Valley
The Fergana Valley (also commonly spelled the Ferghana Valley) in Central Asia crosses eastern Uzbekistan, southern Kyrgyzstan and northern Tajikistan.
Encompassing three former Republics of the Soviet Union, Soviet republics, the valley is e ...
, Kyrgyzstan. The holotype of "Ferganastegos callovicus", IGB 001, consists of four posterior dorsal vertebrae. Although Averianov et al. did not consider the vertebrae diagnostic to genus, the freelance Russian dinosaur enthusiast and amateur paleontologist Roman Ulansky decided that the differences between IGB 001 and other stegosaurs were sufficient to warrant a binomial for IGB 001, "Ferganastegos callovicus" (Callovian roof from Fergana Valley), despite the fact he did not examine the material himself. Other researchers still contend that the material is not diagnostic and that the genus is a nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
.
Ferropectis
"Ferropectis" is a nodosaurid
Nodosauridae is a family of ankylosaurian dinosaurs known from the Late Jurassic to the Late Cretaceous periods in what is now Asia, Europe, North America, and possibly South America. While traditionally regarded as a monophyletic clade as the s ...
ankylosaur
Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) Eagle Ford Group
The Eagle Ford Group (also called the Eagle Ford Shale) is a Sedimentary rock, sedimentary rock formation deposited during the Cenomanian and Turonian ages of the Late Cretaceous over much of the modern-day state of Texas. The Eagle Ford is pr ...
in Texas
Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
that was named in a 2018 dissertation by Matt Clemens. The intended type species is "Ferropectis brysorum", and in the phylogenetic analysis it was placed as the sister taxon to '' Borealopelta'' in a clade including '' Hungarosaurus'', '' Europelta'', and '' Pawpawsaurus''.
Francoposeidon
"Francoposeidon" (meaning "French earthquake god") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of turiasauria
Turiasauria is an unranked clade of eusauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa.
Description
Turiasauria was originally erected by Royo-Torres et al. (2006) to include '' Tur ...
n sauropod dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
(Hauterivian
The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
)-aged Angeac-Charente bonebed of France. The proposed type species is "F. charantensis", and the remains consist of a braincase, some skull bones, teeth, cervical, dorsal and caudal vertebrae, chevrons, pelvic girdle and all the limb bones" alongside isolated teeth, belonging to at least 7 individuals. The length of the femur was measured to be around , (± ), making "Francoposeidon" one of the largest known sauropods discovered in Europe.
Futabasaurus
"Futabasaurus" is an informal name for a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Japan, known only from a partial shin bone of ~ wide that was discovered in the Coniacian-age Ashizawa Formation of the Futaba Group; it was likely around when fully grown. It was first mentioned as "Futaba-ryu" by Hasegawa et al. (1987), and the name was coined by David Lambert in 1990 as a conversion from the Japanese nickname "Futaba-ryu", for an undescribed theropod. Dong Zhiming
Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
and coauthors briefly discussed the fossil shin bone it was based on that same year, publishing a photograph. They considered the bone to belong to an indeterminate tyrannosaurid. If the specimen is eventually described and named, it will require a different name, because the name '' Futabasaurus'' has since been used for a genus of plesiosaur
The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia.
Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
.
G
Gadolosaurus
"Gadolosaurus" is an informal name given to PIN, no. 3458/5 an unnamed juvenile hadrosauroid dinosaur specimen from the Bayan Shireh Formation of Baishan Tsav, Mongolia. The name "Gadolosaurus" was first used in a 1979 book by Japanese paleontologist Tsunemasa Saito, in a caption for a photo of the specimen. This specimen represents an individual that was only about a meter long (39 inches). The specimen was part of a Soviet exhibition of fossils in Japan. Apparently, the name comes from a Japanese phonetic translation of the Cyrillic
The Cyrillic script ( ) is a writing system used for various languages across Eurasia. It is the designated national script in various Slavic, Turkic, Mongolic, Uralic, Caucasian and Iranic-speaking countries in Southeastern Europe, Ea ...
word ''gadrosavr'', or hadrosaur
Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
, and was never meant by the Russians to establish a new generic name.
Despite the only name ever applied to it being merely a mistranslation of ''gadrosavr'', this specimen has appeared in many popular dinosaur books, with varying identifications. Donald F. Glut in 1982 reported it as either an iguanodont
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivore, herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and rel ...
or hadrosaur, with no crest or boot on the (the lack of which are both characteristics of the crested lambeosaurine duckbills), and suggested it could be the juvenile of a previously named genus like '' Tanius'' or '' Shantungosaurus''. David Lambert in 1983 classified it as an iguanodont, but changed his mind by 1990, when it was listed as a synonym
A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
of '' Arstanosaurus'' without comment. What may be the same animal is mentioned but not named by David B. Norman and Hans-Dieter Sues
Hans-Dieter Sues (born 1956) is a German-born American palaeontologist who is a Senior Research Geologist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC.
Career ...
in a 2000 book on Mesozoic reptiles from Mongolia and the former USSR; this material, from the Soviet-Mongolian expeditions of the 1970s, had been listed as ''Arstanosaurus'' in the Russian Academy of Sciences
The Russian Academy of Sciences (RAS; ''Rossíyskaya akadémiya naúk'') consists of the national academy of Russia; a network of scientific research institutes from across the Russian Federation; and additional scientific and social units such ...
, and was found in the Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
-age Bayan Shireh Formation of Baishin Tsav.
Averianov, Lopatin, and Tsogtbaatar in 2022 provided a preliminary description of this specimen and its taxonomic position, finding that the specimen may represent a juvenile of a novel taxon that was closely related to but more derived than the contemporary hadrosauroid '' Gobihadros''.
Gallimimus mongoliensis
"Gallimimus mongoliensis" is an informal name Rinchen Barsbold used for a nearly complete skeleton (IGM 100/14) known from the Bayan Shireh Formation, but since it differs from '' Gallimimus'' in some details, Yoshitsugu Kobayashi and Barsbold proposed in 2006 that it probably belongs to a different genus. It was recently included in a phylogenetic analysis, which recovered it as closely related to '' Tototlmimus.''
Gspsaurus
"Gspsaurus" (a ''nomen manuscriptum'') is a titanosaurian sauropod dinosaur from the Late Cretaceous Vitakri Member of the Pab Formation of Sulaiman Basin of Pakistan. It has been suggested to be synonymous with the also invalid taxon " Maojandino", also proposed by Malkani. The intended holotype, MSM-79-19 and MSM-80-19, consisting of parts of the skull, including a rostrum, was discovered in 2001, and parts of the holotype were initially referred to "Marisaurus jeffi".[Malkani, M.S., (2015). Dinosaurs, Mesoeucrocodiles, Pterosaurs, New Fauna and Flora from Pakistan. ''Geological Survey of Pakistan'', Information Release 823: 1–32.]
Grusimimus
"Grusimimus" (or "Tsurumimus") is an informal name for an undescribed genus of ornithomimid from the Early Cretaceous (Hauterivian
The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
–Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) aged Shinekhudag Formation of Mongolia. Known from a skeleton including all regions except the skull, "Grusimimus" was given an invalid name in 1997
Events January
* January 1 – The Emergency Alert System is introduced in the United States.
* January 11 – Turkey threatens Cyprus on account of a deal to buy Russian S-300 missiles, prompting the Cypriot Missile Crisis.
* January 1 ...
by Rinchen Barsbold, who also suggested the species name "tsuru". The specimen (GIN 960910KD) was found in 1996 and examined by Barsbold before he suggested the informal name, a ''nomen nudum''. An abstract and poster were presented on the taxon by Kobayashi & Barsbold in 2002, and the former published a thesis paper on the specimen (referred to as "Ornithomimosauria indet.") which found the taxon to be close to '' Harpymimus'' phylogenetically but possible more derived. A recent phylogenetic analysis recovered "Grusimimus" closely related to '' Beishanlong'' and '' Garudimimus.''
H
Hanwulosaurus
"Hanwulosaurus" is the informal
Formal, formality, informal or informality imply the complying with, or not complying with, some set of requirements ( forms, in Ancient Greek). They may refer to:
Dress code and events
* Formal wear, attire for formal events
* Semi-formal att ...
name given to an as-yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
. It was an ankylosaur
Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
around long, which is long for an ankylosaur. Its fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s were found in Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...
, China. Much of a skeleton, including a complete skull
The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate.
In the human, the skull comprises two prominent ...
, vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e, ribs, a scapula
The scapula (: scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side ...
, an ulna
The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
, femora, bones from the shin, and armor
Armour (Commonwealth English) or armor (American English; see American and British English spelling differences#-our, -or, spelling differences) is a covering used to protect an object, individual, or vehicle from physical injury or damage, e ...
, was discovered; this may be the most complete ankylosaurian skeleton
A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...
yet found in Asia, according to early reports. Zhao Xijin, who has studied it, suggests that it may belong to its own subgroup within the Ankylosauria. The name first surfaced in news reports in 2001.
Haute Moulouya Sauropod
The "Haute Moulouya Sauropod", also known as NHMUK PV R36834, consisted originally of two complete cervical vertebrae recovered from the Lower Jurassic sediments of the Haute Moulouya Basin, likely the Aganane Formation, on central Morocco. This material was initially identified as belonging to an early member of Eusauropoda, if so, the oldest member of the group. Additional material was previously recovered, SNSB-BSPG 2014 I 106 that consists of dorsal vertebrae and a pubis fragment. A recent revision suggest both specimens belong to the same taxon, that likely comes from a higher stratigraphic level (Likely Late Pliensbachian) and that represents a valid more basal taxon, related with ''Amygdalodon''. Tought other analisis still recover it alternatively as an Eusauropod, in a polytomy with ''Barapasaurus''.
Heilongjiangosaurus
"Heilongjiangosaurus" (meaning "Heilongjiang lizard") is the informal name given to an as-yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of hadrosaurid, duckbilled dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
. It possibly was a lambeosaurine, and may in fact be the same animal as ''Charonosaurus''. The fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s were found in Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age rocks in Heilongjiang, China. As a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
'', it is unclear what material it was intended to be based on, but might be connected to the ''nomen nudum'' ''"Mandschurosaurus" jiainensis'', informally named in a 1983 publication.
The "type species" is "H. jiayinensis", and it was coined in 2001 in a faunal list by Li and Jin.
Hironosaurus
"Hironosaurus" (meaning "Hirono lizard") is the informal name given to an as-yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
. Found in Hirono, Fukushima, Japan, it was probably a type of hadrosaur
Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
, although no subfamily identification has been made. The fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s are quite fragmentary, and consist of teeth and a vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
, possibly from the tail. Since the fossils have never been fully described in a scientific paper, "Hironosaurus" is considered a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. It was first mentioned by Hisa in an obscure 1988 publication and was later (1990) brought to a wider audience by David Lambert.[ ]Dong Zhiming
Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
, Y. Hasegawa, and Y. Azuma regarded the material as belonging to a hadrosaurid, but lacking any characteristics to allow more precise identification (thus indeterminate).
Hisanohamasaurus
"Hisanohamasaurus" (meaning "Hisano-hama lizard") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
. It is a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
'' known only from teeth that first appeared in a general-audience dinosaur book by David Lambert (author), David Lambert in 1990. Although initially identified a diplodocid, it later re-identified as a nemegtosaurid similar to ''Nemegtosaurus''. As its name suggests, its fossils were found in Japan. The location is part of Iwaki, Fukushima.
Hughenden sauropod
The "Hughenden sauropod" is an informal name given to a titanosauriform sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
specimen (QM F6142) discovered from the Early Cretaceous (Albian) Toolebuc Formation of Australia. It is uncertain whether the specimen can be referred to ''Austrosaurus'' from the Allaru Formation, which overlies the Toolebuc Formation.
I
Ikqaumishan
"Ikqaumishan" is an informal genus of titanosaurian dinosaurs from the Late Cretaceous (Maastrichtian) Vitakri Formation of Pakistan described by Malkani (2023) in Scientific Research Publishing
Scientific Research Publishing (SCIRP) is a predatory academic publisher of open-access electronic journals, conference proceedings, and scientific anthologies that are of questionable quality. , it offered 244 English-language open-access journ ...
, a known predatory publisher. The assigned fossil material includes multiple humeri. Caudal vertebrae and osteoderms found nearby may also be referrable to "Ikqaumishan". The intended type species is "Ikqaumishan smqureshi."
Imrankhanhero
"Imrankhanhero" is an informal genus of titanosaurian dinosaurs from the Late Cretaceous (Maastrichtian) Vitakri Formation of Pakistan described by Malkani (2023) in Scientific Research Publishing
Scientific Research Publishing (SCIRP) is a predatory academic publisher of open-access electronic journals, conference proceedings, and scientific anthologies that are of questionable quality. , it offered 244 English-language open-access journ ...
, a known predatory publisher. The assigned fossil material includes a humerus, a femur, fibulae, a tibia, and a metatarsal. Caudal vertebrae found nearby may also be referrable to "Imrankhanhero". The intended type species is "Imrankhanhero zilefatmi."[
]
Imrankhanshaheen
"Imrankhanshaheen" is an informal genus of titanosaurian dinosaurs from the Late Cretaceous (Maastrichtian) Vitakri Formation of Pakistan described by Malkani (2024) in Scientific Research Publishing
Scientific Research Publishing (SCIRP) is a predatory academic publisher of open-access electronic journals, conference proceedings, and scientific anthologies that are of questionable quality. , it offered 244 English-language open-access journ ...
, a known predatory publisher. The proposed holotype includes a braincase, vertebrae, a humerus, ulnae, a radius, metacarpals, a tibia, fibulae, ribs, girdle bones, and osteoderms. The intended type species is "Imrankhanshaheen masoombushrai."
J
Jeholraptor
"Jeholraptor" is the informal replacement genus name given to the microraptorine ''Sinornithosaurus haoiana''—resulting in the Combinatio nova, new combination "Jeholraptor" ''haoiana''—by Gregory S. Paul in the third edition of The Princeton Field Guide to Dinosaurs in 2024. The ''S. haoiana'' fossil is known from the Early Cretaceous (Barremian) upper Yixian Formation
The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
of China. The specimen, which is nearly complete, is about long and was probably close to in weight. Paul suggested that, due to similarities in the quadratojugal, "Jeholraptor" may have been a close relative of ''Wulong bohaiensis, Wulong''.
Jiangjunmiaosaurus
"Jiangjunmiaosaurus" (meaning "temple of the general lizard") is an informal name created by an anonymous author in 1987 for a possible chimaera of ''Monolophosaurus'' and ''Sinraptor''. Gregory S. Paul, Paul (1988) tentatively placed "Jiangjunmiaosaurus" within Allosauridae and commented on the nasal ridges and orbital horn combining to form low, rugose-surfaced crests, and mentioned that "other excellent bones" may also be referable to "Jiangjunmiaosaurus".
Jindipelta
"Jindipelta" (Lei et al., 2019; in press) is the currently informal name given to an Ankylosauridae, ankylosaur from the Zhumapu Formation in China. It is known from a partial skeleton found in Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
rocks and the intended type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is "J. zouyunensis". The name was first announced in the 2019 SVP abstract book, alongside the megalosauroidea, megalosauroid ''Yunyangosaurus''.
Julieraptor
"Julieraptor" is the nickname of a dromaeosaurid fossil found in the Judith River Formation, Montana in 2002. Parts of the same skeleton were illegally excavated and nicknamed Sid Vicious in 2006, and the poacher responsible subsequently served jail time for the theft. Bob Bakker therefore also nicknamed the specimen "Kleptoraptor". The skeleton was arranged to be sold to Royal Ontario Museum. It is known from a skeleton consisting of an almost complete skeleton missing most of its skull, most tail vertebra, part of the femur, some spinal and neck vertebra, one claw but it has a well preserved braincase.
K
Kagasaurus
"Kagasaurus" (meaning "Kaga lizard") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
of what is now Japan. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was named by Hisa in 1988, but is known from only two teeth with no formal description, so it is considered a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. It may belong to either a non-maniraptoriform coelurosaur or a dromaeosaurid.
Katsuyamasaurus
"Katsuyamasaurus" is an informal name for a genus of intermediate theropod known from the Early Cretaceous (Barremian) of the Kitadani Formation, Japan. Known from a single middle caudal vertebra and an ulna, the taxon was informally called "Katsuyama-ryu", until Lambert (1990) made it into an invalid genus name, "Katsuyamasaurus". The caudal vertebra was suggested to belong to an ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
by Chure (2000), and Olshevsky (2000) suggested the material was a synonym of '' Fukuiraptor''. However, the ulna differs from ''Fukuiraptor'', and the large olecranon suggests the taxon falls outside Maniraptoriformes.
Khanazeem
"Khanazeem" is an informal genus of titanosaurian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
Vitakri Formation of Pakistan. The holotype is a partial skeleton and consists of a dentary with teeth, caudal vertebrae, femora, humeri, and tibiae. The intended type species is "Khanazeem saraikistani" and was first mentioned by Malkani (2022).
Khetranisaurus
"Khetranisaurus" (meaning "Khetran lizard", for the Khetran people of Pakistan) is an informal taxon of titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
ian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Balochistan (Pakistan), Balochistan, western Pakistan (also spelled "Khateranisaurus" in some early reports). The proposed species is "K. barkhani", described by M. Sadiq Malkani in 2006, and it is based on a tail vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
, found in the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age Vitakri Member of the Pab Formation. It was assigned to "Pakisauridae" (used as a synonym of Titanosauridae), along with "#Pakisaurus, Pakisaurus" and "#Sulaimanisaurus, Sulaimanisaurus". It was considered invalid by Wilson, Barrett and Carrano (2011).
Koreanosaurus
"Koreanosaurus" (meaning "Korean lizard") is the informal name given to an as-yet unnamed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
(Aptian-Albian). It was a possible dromaeosaur (or similar theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
) which was discovered in the Gugyedong Formation of South Korea, although at times it has been referred to the Tyrannosauridae, Hypsilophodontidae and Hadrosauridae. Based solely on DGBU-78(=DGBU-1978B), a femur, the name was coined by Kim in 1979, but by 1993 Kim decided that it was a species of ''Deinonychus'', and created the informal name "D." "koreanensis".[Zhen, Li, Han, & Kim. (1993). Comparative studies of dinosaur remains in China and Korea. Journal of Natural History and Environment, 1(1), 1–21.] Kim et al. (2005) referred the specimen to Eumaniraptora based on a proximolateral ridge, shelf-like posterior trochanter, and absence of an accessory trochanter and mediodistal crest. The presence of a large fourth trochanter was noted to be similar to ''Adasaurus'' and ''Velociraptor''.
Kunmingosaurus
"Kunmingosaurus" is an informally named primitive sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
which lived during the Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
. Its fossils were found in Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
, China in 1954. The type species, type and only species is "Kunmingosaurus wudingensis", invalidly coined by Zhao in 1985. It is known from fossils found in the Fengjiahe Formation (or the Lower Lufeng Series), including pelvis, pelvic, hind limb, and vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
l material."Kungmingosaurus" at the Fukui Prefectural Dinosaur Museum (contains photos of the mounted skeleton)
->
L
Lancanjiangosaurus
"Lancanjiangosaurus" (alternative spelling "Lanchanjiangosaurus"; meaning "Lancangjiang lizard", named after the Lancangjiang River of China) is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from the Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
. The "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
", "L. cachuensis", was coined by Zhou in 1983, but remains a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. It is known from the Dapuka Group of Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
.
Lijiagousaurus
"Lijiagousaurus" (meaning "Lijiagou lizard") is the nomen nudum, informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of herbivorous
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
iguanodontian dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of what is now Sichuan, China. It has not been formally described yet, but the formal publication is forthcoming, from Chinese paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Ouyang Hui. "Lijiagousaurus" was only briefly mentioned in the Chongqing Natural History Museum guidebook (2001) and is thus a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''.The holotype consists of hindlimb bones, a scapula, an ischium and other fragments.[Anonymous (2001). ''Chongqing Natural History Museum guidebook.'']
Likhoelesaurus
"Likhoelesaurus" (meaning "Li Khole lizard") is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of archosauriform, either a dinosaur or rauisuchian, from the Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
of what is now South Africa. The name was coined by Ellenberger in 1970, and the "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
" is "Likhoelesaurus ingens". It is named after the town in Lesotho where the fossils were found. The only fossils recovered have been teeth, from the Carnian, late Carnian–Norian, early Norian-age Lower Elliot Formation. Ellenberger (1972) regarded the genus as a giant carnosaur, and Kitching and Raath (1984) treated it as possibly referable to ''Basutodon''. Knoll listed "Likhoelesaurus" as a rauisuchian, also he noted that could also be a rauisuchian.
Lopasaurus
"Lopasaurus" (meaning "Alberto Lopa's lizard") is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dromaeosauridae, dromaeosaurid theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
, possibly belonging to Unenlagiinae due to its similarity to ''Buitreraptor'', ''Neuquenraptor'' and ''Pamparaptor'', from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
)-aged Serra da Galga Formation in the Ponto 1 do Price site of Brazil. The intended holotype, a partial right metatarsus showing metatarsals II, III and IV, was discovered by Alberto Lopa during the 1950s but the fossil was lost shortly after the death of Llewellyn Ivor Price in 1980 and it has not been located since. "Lopasaurus" was briefly mentioned by Brum et al. in their description of ''Ypupiara, Ypupiara lopai'', where it was tentatively referred to Unenlagiinae. Brum et al. (2021) also did not refer "Lopasaurus" to ''Ypupiara'', which was found in the same formation as "Lopasaurus".
M
Magulodon
"Magulodon" is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
(Aptian to Albian faunal stage, stages, approximately 112 million years ago). It was a possible ornithischian, either an ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
or basal ceratopsian, which was discovered in the Arundel Formation of Maryland, United States. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, "Magulodon muirkirkensis", was coined by Kranz in 1996.[Kranz, P. (1996). Notes on the sedimentary iron ores of Maryland and their dinosaurian faunas. Maryland Geological Survey Special Publications 3:87–115.] It is a tooth taxon, based solely on a single tooth. Since it has not been formally described, it is also a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. It was considered to be an indeterminate specimen in a paper which cited the intended type specimen but avoided using the name to prevent taxonomic clutter.[Chinnery, B.J., Lipka, T.R., Kirkland, J.I., Parrish, J.M., and Brett-Surman, M.K. (1998). Neoceratopsian teeth from the Lower Cretaceous of North America. In: Kirkland, J.I., and Estep, J.W. (eds.). ''Lower Cretaceous Terrestrial Ecosystems''. New Mexico Museum of Natural History and Science Bulletin 14:297–302.]
Maltaceratops
"Maltaceratops" is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of Centrosaurinae, centrosaurine ceratopsian from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
-aged) Judith River Formation of Montana. The proposed type species is "M. hammondorum", and the proposed holotype is a possible skull. It had been previously nicknamed the "Malta new taxon".
Mangahouanga
"Mangahouanga" (named after Te Hoe River, the stream of the same name), or the "Joan Wiffen's theropod" is an informal name given to the theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
discovered in the Tahora Formation, New Zealand by Joan Wiffen, who considered it to be a possible Megalosauridae, megalosaurid in 1975. The vertebra was described by Molnar 1981, and it was ruled as an indeterminate theropod in 2010 by Agnolin et al. The name "Mangahouanga" was coined by Molina-Pérez & Larramendi (2016) and no species name was given. They estimated it to reach up to long and weigh up to and is represented by of a single vertebra.
Maojandino
"Maojandino" is an informally named taxon of titanosaurid sauropod dinosaur from the Late Cretaceous Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
stage of Pakistan. The intended type species is "Maojandino alami."
Marisaurus
"Marisaurus" (meaning "Mari lizard", for the Mari tribe of Pakistan) is an informal taxon of titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
ian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Balochistan (Pakistan), Balochistan, western Pakistan. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is "M. jeffi", described by Muhammad Sadiq Malkani in 2004, and it is based on tail vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e, found in the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age Vitakri Member of the Pab Formation. Much additional material, including a partial skull, many vertebrae, and a few hindlimb bones, was referred to this genus. "Marisaurus" was assigned to "Balochisauridae" with "#Sulaimanisaurus, Sulaimanisaurus", although the family was used as a synonym of Saltasauridae. It was considered invalid by Wilson, Barrett and Carrano (2011).
Maroccanoraptor
"Maroccanoraptor" is an informal name suggested for a supposed unenlagiine theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
from the Kem Kem Formation of Morocco, however, it lacks the requirements to become a valid taxon, thus leaving it as a naked name. The intended type species is "M. elbegiensis", first described by Singer (2015) on the basis of a single coracoid. The fossil was later suggested to belong to a non-dinosaurian crocodyliform.
Megacervixosaurus
"Megacervixosaurus" (meaning "big neck lizard") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of herbivorous
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
Zonggo Formation of Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
. It was a titanosaur sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
which lived in what is now China The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, "Megacervixosaurus tibetensis", was coined by Chinese paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Zhao Xijin in 1983. "Megacervixosaurus" has never been formally described, and remains a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''.[Zhao X. (1985). "Phylogeny and evolutionary stages of Dinosauria." ''Acta Palaeontologica Polonica'' 28(1–2); 295–306.]
Megapleurocoelus
"Megapleurocoelus" is an informally named sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
belonging to Flagellicaudata, from the Kem Kem Formation of Morocco, however, it lacks the requirements to become a valid taxon, thus leaving it as a naked name. The intended type species is "M. menduckii", first described by Singer (2015) and the holotype is JP Cr376, a single centrum from a dorsal vertebra.
Microcephale
"Microcephale", also known as "Mycocephale", (meaning "tiny head") is the nomen nudum, informal name of a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of very small pachycephalosaurid dinosaur, otherwise known as the "North American dwarf species", which lived during the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
. Its fossils were found in the Campanian, late Campanian-age Dinosaur Park Formation, in Alberta, Canada. Not much is known about this dinosaur, as it has not yet been fully described; it is therefore a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. The fossils of "Microcephale", including tiny skull caps, were first mentioned by paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Paul Sereno in 1997, in a list of pachycephalosaurids. These skull caps measure less than 5 cm (2 in) each. No potential species name was given.
Microdontosaurus
"Microdontosaurus" (meaning "tiny-toothed lizard") is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from China. It was named from fossils from the Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
-age Dapuka Group of Xinjiang
Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...
. The intended type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is "M. dayensis." As with other informal names created by Zhao in 1985 or 1983, it has not been used since then, and may have been redescribed under another name.
Microvenator chagyabi
"Microvenator chagyabi" is the informal name given to an as yet undescribed species of theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur, likely belonging to Coelurosauria, from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Lura Formation of Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, China. It was coined by Zhao (1985) and the proposed holotype consists of a specimen including teeth.
Mifunesaurus
"Mifunesaurus" (meaning 'Mifune lizard') is a nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
given to an extinction, extinct non-avian dinosaur, non-avian non-maniraptoriform tetanuran theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
; ~96 Ma) Kabu Formation of Japan. The intended holotype, stored at the Mifune Dinosaur Museum, with the tooth on display, of "Mifunesaurus" consists only of a few bones, among which are a tibia, a phalanx
The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
, a metatarsus and a single tooth (tooth catalogued as YNUGI 10003; rest of the skeleton catalogued as MDM 341), discovered by N. & K. Wasada in 1979. The genus was informally coined by Hisa in 1985 and no epithet was given. The known tooth was too thick to be the tooth of a ceratosauridae, ceratosaurid, and too tall to belong to an abelisauridae, abelisaurid, which means that "Mifunesaurus" was probably a megalosauroidea, megalosauroid or a carnosauria, carnosaur based on the shape of the known tooth.
Mitchell ornithopod
The "Mitchell ornithopod" is the informal nickname of an ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
dinosaur discovered near Mitchell, Oregon, being the first described dinosaur from Oregon but not the first discovered; a hadrosauridae, hadrosaurid sacrum was discovered in the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
)-aged Cape Sebastian Sandstone near Cape Sebastian, Oregon, Cape Sebastian during the 1960s and excavated in 1994 by Dave Taylor, but the remains of the Cape Sebastian ornithopod were not prepared for peer review and described until 2019, merely weeks after the Mitchell ornithopod was described. The single known bone, F118B00, was a toe bone, specifically the third phalanx
The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
of the central digit of the right hindlimb foot, and was discovered by Gregory Retallack in 2015 while on an annual field trip with his students, in a layer of the Albian-aged Hudspeth Formation, Hudspeth Shale Formation; in 2021, Gloria Carr discovered another bone, this time a vertebra, that likely belonged to the same species of ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
. No excavation was required – the bone was found resting on the ground and Retallack immediately knew it was different from the various marine fossils scattered nearby. The bone was described in 2018 by Gregory Retallack, Jessica Theodor, Edward Davis, Samantha Hopkins and Paul Barrett.[Retallack, Gregory J.; Jessica M. Theodor; Edward B. Davis; Samantha S. Hopkins, and Paul Z. Barrett. (2018). First Oregon dinosaur (Ornithopoda) from Early Cretaceous (Albian) of Oregon, U.S.A. ''Journal of Vertebrate Paleontology.'' 1–5.] It was part of a bloated carcass swept out into the ocean, likely originating from Idaho, although further discoveries, such as Strommer (2021), dispute this claim and suggest it may have been deposited by a mudflow.
The bone was later compared to more complete remains of other ornithopods and the "Mitchell ornithopod" bone most closely matched those of hadrosaur
Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
s and iguanodontia, iguanodonts, although it was likely a basal ornithopod. Rettalack believes that the bone belonged to a new genus, although there is not enough sufficient remains to base this claim on.
Moshisaurus
Hisa (1985) used "Moshisaurus" (or "Moshi-ryu") for the incomplete sauropod humerus NSM PV17656, from the Early Cretaceous Miyako Group of Japan. Dong et al. (1990) and Hasegawa et al. (1991) referred them to ''Mamenchisaurus'', but Azuma & Tomida (1998) and Barrett et al. (2002) assigned them to Sauropoda indet.
N
Newtonsaurus
"Newtonsaurus" is an informally named genus erected for the theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur species ''Zanclodon cambrensis.'' The species is based on the specimen BMNH R2912, an Fossil#Casts and molds, external mold of a dentary, which was discovered in the Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
(Rhaetian) aged beds of the Lilstock Formation near Bridgend, Wales in 1898 and described by Edwin Tulley Newton in 1899.[Newton, E.T. (1899). On a megalosaurid jaw from Rhaetic beds near Bridgend (Glamorganshire). ''Quarterly Journal of the Geological Society of London'' 55:89–96.] The taxon was reassigned to ?''Megalosaurus'' by Molnar in 1990, which was followed by Peter Galton in publications in 1998 and 2005. The species is considered to be a ''nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
'', as it has no diagnostic features, and is considered to be a Coelophysoidea, coelophysoid grade theropod outside Averostra based on the low interdental plates and possession of only a single Meckelian groove, meckelian foramen. It has alternatively been suggested to possibly represent another indeterminate predatory archosaur.
The name "Newtonsaurus" was coined in 1999 by Stephan Pickering, in reference the describer. Paleontologists have avoided using the name "Newtonsaurus" since its appearance in 1999 in private publications, although "''Zanclodon''" ''cambrensis'' or ''Megalosaurus cambrensis'' have both been used for this taxon.
Ngexisaurus
"Ngexisaurus" is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur, likely belonging to Avetheropoda, from the Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
Dapuka Group of Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, China. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, "Ngexisaurus dapukaensis", was coined by Zhao in 1983.[Zhao, X. (1985). [The Jurassic Reptilia]. [The Jurassic System of China. ''Stratigraphy of China'', No. 11] 286–290.] A synonym of "Ngexisaurus" coined by Zhao (1985) is "''Megalosaurus''" dapukaensis and Fossilworks lists "''M.''" dapukaensis as a Megalosauridae, megalosaurid tetanurae, tetanuran separate from "Ngexisaurus" proper.
Nicksaurus
"Nicksaurus" is an informally named Titanosaurian sauropod dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
red muds of the Vitakri Formation of the Sulaiman Basin, Pakistan. The dinosaur shared a habitat with other sauropod dinosaurs including #Khetranisaurus, Khetranisaurus, #Sulaimanisaurus, Sulaimanisaurus, #Pakisaurus, Pakisaurus, #Gspsaurus, Gspsaurus, #Saraikimasoom, Saraikimasoom, and #Maojandino, Maojandino.
The intended type species is "Nicksaurus razashahi" and was first used by Malkani (2019).
Nurosaurus
"Nurosaurus" (Nur-o-saw-rus, meaning "Nur lizard") is the informal
Formal, formality, informal or informality imply the complying with, or not complying with, some set of requirements ( forms, in Ancient Greek). They may refer to:
Dress code and events
* Formal wear, attire for formal events
* Semi-formal att ...
name for a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur. It is known from a partial, large skeleton, that was presented as soon-to-be-described by Zhiming Dong in 1992 in paleontology, 1992, where he gave the proposed binomial "Nurosaurus qaganensis". It was discovered in the Qagannur Formation of Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...
, southeast of Erenhot. The deposit is younger than the ''Psittacosaurus''-bearing Guyang Group, but is still Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
. It was found alongside the plates and scapula of a stegosaur.
The foot of "Nurosaurus" is notable for a stress fracture present on the first of the fourth digit of the left foot, which was the first identified fracture of its kind, and have since been identified on the phalanges and metatarsals of ''Apatosaurus'', ''Barosaurus'', ''Brachiosaurus
''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about . It was first Species description, described by American paleontologist Elmer S. Riggs in 1903 in paleontology, 1903 from fossi ...
'', ''Camarasaurus'', and ''Diplodocus''.
O
Oharasisaurus
"Oharasisaurus" is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of somphospondylian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
, possibly belonging to the Euhelopodidae, from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Kuwajima Formation (Facies III layer) of Japan. The name "Oharasisaurus" was coined by Larramendi & Molina Pérez (2020) and the holotype, a tooth, was first mentioned by Matsuoka (2000).
Orcomimus
"Orcomimus" (Pronounced or-coh-mEYEm-us) is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of dinosaur from the Late Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
period 66 million years ago. The dinosaur was an ornithomimid which lived in what is now South Dakota
South Dakota (; Sioux language, Sioux: , ) is a U.S. state, state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Dakota people, Dakota Sioux ...
, in the United States. The type (biology), type was coined by Michael Triebold in 1997, but has never been formally described and is currently a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. "Orcomimus" was a bipedal theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
, but the dinosaur is known from only a pelvis
The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...
and a hindlimb. "Orcomimus" is thought to be relatively advanced for other ornithomimids at the time, although this is hard to tell from the limited amount of specimens found of the dinosaur. It may be referable to one of the ornithomimosaur species currently known from the Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
, where the holotype of "Orcomimus" was found.
Oshanosaurus
"Oshanosaurus" (meaning "Eshan Yi Autonomous County, Oshan lizard") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from the Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
period of Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
, China. Its fossils were found in the Lower Lufeng Series. The intended "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
", "Oshanosaurus youngi", was coined by Zhao in 1985. It has sometimes been associated with heterodontosauridae, heterodontosaurids, which appears to be due to the juxtaposition of a species of ''Dianchungosaurus'' (formerly thought to be a heterodontosaurid) in the text of Zhao (1985).
In 1971 Zhao Xijin discovered a dinosaur fossil at Dianchung in Eshan Yi Autonomous County, Eshan county, giving it the informal name "Oshanosaurus youngi". In their 2019 popular book ''Dinosaur Facts and Figures: The Theropods'', Molina-Perez and Larramendi suggested that it belonged to the theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
''Eshanosaurus'', but without elaboration.
Osteoporosia
"Osteoporosia gigantea" is an informally named theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
, either belonging to Carcharodontosauridae or Megaraptora, from the Kem Kem Group of Morocco. It lacks the requirements to become a valid taxon, thus leaving it as a Nomen nudum, naked name. It was informally described by Singer (2015) based on a tooth and a fragment of neural arch probably belonging to a posterior dorsal vertebra (JP Cr340).[Singer (2015). JURAPARK NA TROPIE NOWYCH DINOZAURÓW Z MAROKA (online).https://web.archive.org/web/20151206224352/https://jurapark.pl/jurapark-na-tropie-nowych-dinozaurow-z-maroka/] The material is from a private collection.
In 2019 Molina-Pérez & Larramendi suggested that "Osteoporosia" could be a possible synonym of ''Sauroniops pachytholus''. They also estimated it in 7.9 m long and 1.55 tons, with a hip height of 2.25 m.
Otogosaurus
"Otogosaurus" is an informally named sauropod from Inner Mongolia, China. The supposed type species is "Otogosaurus sarulai". It is known from partial postcranial remains, including a tibia long and several footprints. It is named after Otog Banner in Inner Mongolia where it was discovered, and Sarula, the girl who discovered the fossils. Despite sometimes being presented as a valid taxon, sometimes accompanied by citations to Zhao (2004) or Zhao & Tan (2004), scholars have not been able to locate such a source, so it remains informal until a paper is discovered.
P
Pakisaurus
"Pakisaurus" (meaning "Pakistan lizard") is an informal taxon of titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
ian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Balochistan (Pakistan), Balochistan, western Pakistan, and also Gujarat, India. The proposed species is "P. balochistani", and it was named by M. Sadiq Malkani in 2006, based on isolated tail vertebrae found in the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age Vitakri Member of the Pab Formation. In 2023, a femur discovered in the Lameta Formation of India was assigned to "Pakisaurus".
It was considered invalid by Wilson, Barrett and Carrano (2011) during their description of a ''Jainosaurus, Jainosaurus cf. septentrionalis'' skeleton.
"Anokhadino mirliaquati" was synonymised with "Pakisaurus balochistani" by Malkani (2019).
Paw Paw scuteling
The "Paw Paw scuteling" is the name used for a juvenile nodosaurid
Nodosauridae is a family of ankylosaurian dinosaurs known from the Late Jurassic to the Late Cretaceous periods in what is now Asia, Europe, North America, and possibly South America. While traditionally regarded as a monophyletic clade as the s ...
discovered in 1990 from the Paw Paw Formation of northern Fort Worth, Texas. It was discovered by John C. Maurice, the 12-year-old son of fossil collector John M. Maurice. The specimen consists of a partial skeleton including a third of the backbone, part of the skull, and both leg and arm elements. It is one of two or three nodosaurs known from the formation alongside '' Pawpawsaurus'' and ''Texasetes'', and one of the very few known specimens of a baby nodosaur. Some phylogenetic analyses have recovered it as sister to ''Niobrarasaurus''. Although taxonomically indeterminate due to its life stage and fragmentary nature, it is often used in phylogenetic analyses for determining the taxonomic affinity of other nodosaur genera.
Podischion
"Podischion" is an informal genus of hadrosaurid dinosaur known from a skeleton discovered in 1911 on the Red Deer River in Alberta by a crew led by Barnum Brown. The remains were tentatively named "Podischion", which was not mentioned in published literature until Dingus & Norell (2010). It is possible that the skeleton represents an individual of ''Hypacrosaurus''.
Q
Qaikshaheen
"Qaikshaheen" is an informal genus of titanosaurian dinosaurs from the Late Cretaceous (Maastrichtian) Vitakri Formation of Pakistan described by Malkani (2023) in Scientific Research Publishing
Scientific Research Publishing (SCIRP) is a predatory academic publisher of open-access electronic journals, conference proceedings, and scientific anthologies that are of questionable quality. , it offered 244 English-language open-access journ ...
, a known predatory publisher. The proposed holotype specimen includes fragmentary cervical and dorsal vertebrae, partial pectoral and pelvic girdles, humeri, femora, a tibia, and fibulae. Other bones, including several vertebrae, ribs, a humerus, ulnae, metacarpals, metatarsals, a femur, and a partial pelvic girdle, were also referred. The intended type species is "Qaikshaheen masoomniazi."[
]
R
Ronaldoraptor
"Ronaldoraptor", also known as the "Mitrata" Oviraptorid, is an undescribed oviraptoridae, oviraptorid from Mongolia[Gee, H. & Rey, L. V. 2003. A Field Guide to Dinosaurs: The Essential Handbook for Travelers in the Mesozoic. Barron's Educational Series, Inc.] and has been listed as "''Oviraptor sp.''" The name was first used by Luis Rey in 2003, in his book ''A Field Guide to Dinosaurs: The Essential Handbook for Travelers in the Mesozoic'', where he drew an illustration, captioning it "Ronaldoraptor". "Ronaldoraptor" may have been closely related to ''Citipati osmolskae''.
Rutellum
"Rutellum" is the pre-Linnaean taxonomy, Linnaean name given to a dinosaur specimen from the Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
(Oxfordian (stage), Oxfordian)-aged Coralline Oolite Formation. It was a sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
, possibly a Cetiosauridae, cetiosaurid,[Delair, J.B., and Sarjeant, W.A.S. (2002)]
The earliest discoveries of dinosaurs: the records re-examined
''Proceedings of the Geologists' Association'' 113:185–197. which lived in what is now England. The specimen (University of Oxford, OU 1352), called "Rutellum impicatum", was described in 1699 by Edward Lhuyd alongside specimen OU 1358, what is now believed to be a ''Megalosaurus'' tooth crown,[Lhuyd, E. (1699). ]
Lithophylacii Britannici Ichnographia, sive lapidium aliorumque fossilium Britannicorum singulari figura insignium
'. Gleditsch and Weidmann:London. and is notable as the earliest named entity that is recognizable as a dinosaur.[ It was based on a tooth collected from Caswell, near Witney, Oxfordshire.][Gunther, R.T. (1945). ''Early Science in Oxford: Life and Letters of Edward Lhuyd'', volume 14. Author:Oxford.]
Because "Rutellum impicatum" was named before 1758 (the official starting date for zoological nomenclature according to the International Code of Zoological Nomenclature, ICZN), it is not considered a part of modern biological nomenclature.
S
Sabinosaurus
"Sabinosaurus" or "Sabinosaurio" is a name used for PASAC-1, a partial skeleton of a hadrosaur
Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
that was discovered in the Sabinas Basin in Mexico in 2001. It was initially described as ''Kritosaurus'' sp. by Jim Kirkland and colleagues (2006), but considered an indeterminate saurolophine by Prieto-Márquez (2014). This skeleton is about 20% larger than other known specimens, around long, and with a distinctively curved , and represents the largest known well-documented North American saurolophine. Unfortunately, the nasal bones are also incomplete in the skull remains from this material.[
]
Saldamosaurus
"Saldamosaurus" is an informal genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of stegosauridae, stegosaurid dinosaur known from a complete braincase discovered in the Early Cretaceous Saldam Formation of Siberia, Russia. The type species, "Saldamosaurus tuvensis", was named in 2014[ but according to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature and is hence a ''nomen nudum''.]
Saltillomimus
"Saltillomimus" is an informal name for an ornithomimid theropod from the Late Cretaceous (late Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
) of the Cerro del Pueblo Formation in Mexico. It is known from SEPCP 16/237, a partial tail, most of a hindlimb, and forelimb bones, discovered in 1998, and the possible juvenile specimen SEPCP 16/221, a partial leg and hip bone, that was given the name "Saltillomimus rapidus" by Martha Carolina Aguillón Martinez in 2010 in paleontology, 2010. A skeletal reconstruction was put on display in 2014 at the Museo del Desierto, which served to highlight its robust thighs and unusual hips that combine primitive and advanced features seen in ornithomimosaurs from both Asia and North America. Named in Martinez' 2010 thesis, the taxon name is an invalid ''nomen ex dissertatione''.[
]
Sanchusaurus
"Sanchusaurus" (meaning "Lizard from Sanchu") or "Sanchu-ryu" is an nomen nudum, informal name for possible ornithomimosaur dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Period (geology), period of Asia. It is only known by a partial Caudal vertebrae, tail vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
, found in Nakasato, Japan. Dong (1990)[ considered it synonymous with '' Gallimimus'' but the large discrepancy in both age and location between the two species renders this opinion untenable. The genus has not been formally described and is considered a '']nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. It was first mentioned by Hisa in 1985. In 2006, it is shown that this animal is not fully grown, and characters of tail vertebra is not unique to that of ornithomimosaur.
Saraikimasoom
"Saraikimasoom" (meaning 'Innocent one') is an invalid species of titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
dinosaur from the Vitakri Formation in Pakistan. The type species, ''Saraikimasoom vitakri'', was described by Sadiq Malkhani in 2015, in a paper describing multiple Pakistani dinosaurs, such as ''Gspsaurus'', "Nicksaurus" and " Maojandino". ''Saraikimasoom'' is currently recognised as a ''nomen manuscriptum''.
Saraikisaurus
"Saraikisaurus" (meaning "Saraiki people, Saraiki lizard") is an invalid genus name proposed for a putative reptile found in the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
)-aged Vitakri Formation of Pakistan and possibly also the Lameta Formation of India. The intended type species is "Saraikisaurus minhui", known from the proposed holotype—a fragmentary dentary (GSP/MSM-157-16)—and a referred specimen—an incomplete vertebra (GSP/MSM-64-15). Malkani initially interpreted the dentary as belonging to a basal pterodactyloid and created the further monotypic family "Saraikisauridae" and subfamily "Saraikisaurinae" to house it. The name "Saraikisaurus" was first proposed by M. Sadiq Malkani in a 2013 conference. A later endeavor to describe it in 2015 was not peer-reviewed. In 2021, Malkani attempted to formally describe "Saraikisaurus" and other taxa in Scientific Research Publishing
Scientific Research Publishing (SCIRP) is a predatory academic publisher of open-access electronic journals, conference proceedings, and scientific anthologies that are of questionable quality. , it offered 244 English-language open-access journ ...
, a known predatory publisher. In 2024, he reinterpreted the specimen as instead belonging to a noasaurid theropod, redescribing it as a novel taxon in another unreviewed paper on ResearchGate. He assigned the fragmentary vertebra to this genus on the basis of compatible size and preservation styles, and apparent similarities to the corresponding bones in ''Laevisuchus''.
Shake-N-Bake theropod
The "Shake-N-Bake theropod" is an undescribed species of coelophysoid from the Kayenta Formation, known from partial skeleton MCZ 8817 within the collection of Harvard Museum of Natural History.
Shansaraiki
"Shansaraiki" (meaning "respected Saraiki peoples") is an informal genus of Theropoda, theropod that was probably an abelisaur. The holotype was found in the Shalghara locality of the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
Vitakri Formation of Pakistan and consists of GSP/ MSM-140-3 (symphysis), GSP/MSM-5-3 (mid-ramus with partial teeth bases) and GSP/MSM-57-3 (dorsal vertebrae), although they may belong to separate specimens as they were found apart from each other. The intended type species is "Shansaraiki insafi" and was first mentioned by Malkani (2022).
Siamodracon
"Siamodracon" is an extinct genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of nomen nudum, invalid stegosauridae, stegosaurid dinosaur known from a single dorsal vertebra found in Thailand's Phu Kradung Formation. The type species, "Siamodracon altispinax", was named by Ulansky in 2014.[ According to Galton and Carpenter (2016) it did not meet the requirements of the International Code of Zoological Nomenclature.][ "Siamodracon" was the first thyreophoran dinosaur discovered in South East Asia.
]
Sidormimus
"Sidormimus" is an informal genus of noasauridae, noasaurid discovered in the Elrhaz Formation in Niger. It was discovered in 2000 by Chris Sidor and it was immediately named as "Sidormimus" by Lyon on the Project Exploration website, with a photograph of the intended holotype. During the same year, on the National Geographic website, the same photograph of the specimen was labelled "Dogosaurus", and it was noted that the neck and ribs were exposed when the specimen was discovered. It has also been referred to as the "Gadoufaoua noasaurid" or the "unnamed Niger noasaurid", and Sidor himself confirmed via personal communication in 2005 that "Sidormimus" was the Elrhaz noasaurid. In a 2010 conference abstract, this articulated specimen was suggested to have possible fossorial (digging) adaptations, which led paleontologists including Paul Sereno to refer the specimen to as the "digging raptor".
Sinopeltosaurus
"Sinopeltosaurus" is a nomen dubium, dubious genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
thyreophoran ornithischian dinosaur described by Roman Ulansky. The type species, type and only species is "S. minimus" of the lower Jurassic Lufeng Formation of Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
China, based on an articulated set of ankle bones.[ The specimen is FMNH CUP 2338, and includes the distal tibia and fibula, distal tarsals, most metatarsals, and some phalanges. FMNH CUP 2338 was described in 2008 by Randall Irmis and Fabian Knoll, as one of the few definitive specimens of Ornithischia from the Early Jurassic based on features of the ankle and pes.] In 2016, Peter Malcolm Galton and Kenneth Carpenter identified it as a ''nomen dubium'', and listed it as Ornithischia indet., possible Thyreophora indet. Ulansky variously referred to it as "Sinopeltosaurus minimus" or "Sinopelta minima"; Galton and Carpenter, as the first revisers under International Code of Zoological Nomenclature, ICZN, made the former official.
Skaladromeus
"Skaladromeus" or the "Kaiparowits ornithopod" is an ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
from the Kaiparowits Formation named in a 2012 thesis by Clint Boyd. The intended type species is "Skaladromeus goldenii".
Sousatitan
"Sousatitan" is the name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of titanosaurian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
-aged Rio Piranhas Formation of Brazil. The intended holotype consists of a left fibula, and "Sousatitan" was coined by Ghilardi et al. (2016).
Stegotitanus
"Stegotitanus" is the informal replacement genus name given to the stegosaur ''Stegosaurus ungulatus''—resulting in the Combinatio nova, new combination "Stegotitanus" ''ungulatus''—by Gregory S. Paul in the third edition of The Princeton Field Guide to Dinosaurs in 2024. ''Stegosaurus ungulatus'' fossils are known from the Late Jurassic (Kimmeridgian) upper Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
of Wyoming, US. "Stegotitanus" was one of the largest stegosaurs, at about long and in weight.
Suciasaurus
A fossil theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
(possibly a Tyrannosauroidea, tyrannosaur) nicknamed "Suciasaurus rex" was discovered in 2012 at Sucia Island State Park in San Juan County, Washington, San Juan County of the U.S. Washington (state), State of Washington. It was the first dinosaur discovered in Washington (state), Washington state. The finding was announced when Burke Museum paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
s published a discovery paper in ''PLoS ONE''. Prompted by a petition from students at an elementary school at Parkland, Washington, Parkland, near Tacoma, the Washington State Legislature introduced a bill in 2019 to make it the official state dinosaur.[
][
][
] A renewed push came in 2021, though House Republicans, like Minority leader J. T. Wilcox, called it low priority versus the ongoing COVID-19 pandemic, and eventually the bill failed to pass, though in 2023 it passed.
Sugiyamasaurus
"Sugiyamasaurus" (meaning "Sugiyama lizard") is the informal name given to a few spatulate teeth belonging to a titanosauriform, possibly ''Fukuititan'', which lived in Japan during the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
. The name was first printed by David Lambert (author), David Lambert in 1990 in the ''Dinosaur Data Book'', and also appears in Lambert's ''Ultimate Dinosaur Book'' and in many on-line lists of dinosaurs. Since it has not been formally described, "Sugiyamasaurus" is a ''nomen nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. Remains were found near Katsuyama, Fukui, Katsuyama City and were initially referred to Camarasauridae, but might belong to ''Fukutitan'' because they were unearthed in the same quarry as the ''Fukuititan'' material.
Sulaimanisaurus
"Sulaimanisaurus" (meaning "Sulaiman lizard", for the Sulaiman foldbelt) is an informal taxon of titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
ian sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Balochistan (Pakistan), Balochistan, western Pakistan (also spelled "Sulaimansaurus" in some early reports).[ The proposed species is "S. gingerichi", described by M. Sadiq Malkani in 2006, and it is based on seven tail ]vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
, found in the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age Vitakri Member of the Pab Formation. Four additional tail vertebrae have been assigned to it. It was considered to be related to "#Pakisaurus, Pakisaurus" and "#Khetranisaurus, Khetranisaurus" in the family "Pakisauridae" (used as a synonym of Titanosauridae). It was considered invalid by Wilson, Barrett and Carrano (2011).
T
Teihivenator
"Teihivenator" ("strong hunter") is an improperly named taxon of Tyrannosauroidea, tyrannosauroid Coelurosauria, coelurosaur from the Navesink Formation of New Jersey. It was suggested to contain the species, "T." ''macropus'', originally classified as a species of '' Dryptosaurus'' (= "Laelaps", a name preoccupied by Laelaps (mite), a mite). It was suggested as a separate genus in 2017 by Chan-gyu Yun. The name "Teihivenator" is invalid because the publication naming it is online-only, which means that a registration with ZooBank is required to be present in the article when published. However, the ZooBank registry was only added in after initial publication, meaning that it fails the requirement to be a validly published taxon.
In 2017, a preprint paper by Chase Brownstein concluded that the remains of ''L. macropus'' are a mixture of tyrannosauroid and ornithomimidae, ornithomimid elements with no distinguishing characteristics, rendering the species a chimera and a ''nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''. In 2018, Brownstein stated that a tibia of ''L. macropus'' catalogued as specimen AMNH FARB 2550 represents a tyrannosauroid that probably was distinct from ''Dryptosaurus'', but not sufficiently to base a taxon on.
That Which Cannot Be Named
"That Which Cannot Be Named" is the name given by Darren Naish to an undescribed associated skeleton of a small coelurosaur from the Wessex Formation of the Isle of Wight. The specimen is in private ownership and is currently inaccessible to researchers. It has been suggested that the specimen is possibly a tyrannosauroid.
Tiantaisaurus
"Tiantaisaurus", alternatively spelled "Tiantaiosaurus", is the name given to a specimen of therizinosaur from the Aptian age Laijia Formation of Zhejiang, China. According to correspondence through the Dinosaur Mailing List, the former name (from a 2012 study) was the one intended to be use for an official description. After being discovered in 2005, it was first mentioned named in an unpublished manuscript written in 2007. The given species was named "T. sifengensis". The specimen consists of an , an astragalus
Astragalus may refer to:
* ''Astragalus'' (plant), a large genus of herbs and small shrubs
*Astragalus (bone)
The talus (; Latin for ankle or ankle bone; : tali), talus bone, astragalus (), or ankle bone is one of the group of foot bones known ...
, a tibia, a femur, an incomplete pubis (bone), pubis and ilium, and a large number of vertebrae
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
from across the body.
Tobasaurus
"Tobasaurus" (meaning "Toba City lizard") is the informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of sauropod dinosaur from the Euhelopodidae from the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
(Hauterivian
The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
– Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
-aged) Matsuo Group of Japan. The proposed holotype is a partial skeleton (mostly limb bones), and "Tobasaurus" grew up to when fully grown. It is the inspiration for the Vivosaur "Toba" in the video game ''Fossil Fighters''.
Tonouchisaurus
"Tonouchisaurus" (meaning "Tonouchi lizard") is the nomen nudum, informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of coelurosaurian dinosaur from the Early Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
Period (geology), Period of Mongolia. The suggested "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
", "Tonouchisaurus mongoliensis", was first informally mentioned in a Japanese news article. It was notably small: less than in length. The specimen informally dubbed "Tonouchisaurus mongoliensis" is based on limb material, and the manual and pedal remains were initially reported to incorporate a complete didactyl Manus (zoology), manus and complete pes (zoology), pes, and Rinchen Barsbold therefore initially interpreted "Tonouchisaurus" as a tyrannosauroid, but he later noted that the manus is actually tridactyl and that the pes has a sub-arcometatarsalian condition.
U
Ubirajara
"Ubirajara jubatus, Ubirajara" (meaning "Lord of the Spear") is an informal genus of Compsognathidae, compsognathid theropod known from the Early Cretaceous Crato Formation of Brazil; it was discovered in 1995 and was named in 2020 in an "In Press" article that was later withdrawn due to the specimen having been illegally smuggled from Brazil to Germany. It is considered a ''nomen manuscriptum''.
Utetitan
"Utetitan" is the informal name given to specimens of the titanosaur ''Alamosaurus'' from the Late Cretaceous (Maastrichtian) lower North Horn Formation of Utah US, by Gregory S. Paul in the third edition of The Princeton Field Guide to Dinosaurs in 2024. Other titanosaurian bones from the upper Black Peaks Formation or Texas, US, may also be referrable to this taxon. "Utetitan" was reportedly about long and in weight. The intended type species is "Utetitan zellaguymondeweyae."
V
Vectensia
In 1982 Justin Delair informally named the genus "Vectensia" based on specimen GH 981.45, an armour plate. Like the holotype of ''Polacanthus'' it was found at Barnes High, but reportedly in an older layer, of the Wessex Formation, Lower Wessex Formation. Blows in 1987 tentatively referred it to ''Polacanthus''.[Blows W.T. (1987). The armoured dinosaur ''Polacanthus foxi'', from the Lower Cretaceous of the ]Isle of Wight
The Isle of Wight (Help:IPA/English, /waɪt/ Help:Pronunciation respelling key, ''WYTE'') is an island off the south coast of England which, together with its surrounding uninhabited islets and Skerry, skerries, is also a ceremonial county. T ...
, Palaeontology. 30, 557–580
Vitakridrinda
"Vitakridrinda" is a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of abelisaurid theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Balochistan (Pakistan), Balochistan, western Pakistan. The intended type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is "V. sulaimani''"''. The discovery was made (along with other dinosaur specimens) near Vitariki by a team of palaeontologists from the Geological Survey of Pakistan, in rocks from the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age Vitakri Member of the Pab Formation.[ Informally named in an abstract by M.S. Malkani in 2004 (to which Malkani [2006] attributes the name), it is based on partial remains including two femur, thigh bones, and a tooth. A partial snout and braincase were originally referred to the holotype, and additional ]vertebra
Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e may also belong to this genus. However, the snout was later reclassified as a new genus of mesoeucrocodylian, ''Induszalim'', while the braincase was later referred to ''Gspsaurus''. Thomas R. Holtz Jr., Thomas Holtz gave a possible length of 6 meters (19.7 feet).
Vitakrisaurus
"Vitakrisaurus" is a genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of noasaurid theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaurs represented by only one known species, "Vitakrisaurus saraiki", which is the intended type species. It lived in the late Cretaceous period, approximately 70 million years ago, during the Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
, in what is today the Indian subcontinent. Its fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s were found in Pakistan's Vitakri Formation. The holotype specimen, MSM-303-2 is a right foot with a seemingly tridactyl form and robust phalanges. It may belong to Noasauridae due to similarities with the foot of ''Velocisaurus'', although inconsistencies within its brief description and a lack of comparison with other Theropoda, theropods within the article makes formal classification difficult. The generic name references the Vitakri Member of the Pab Formation and combines this with the Greek suffix "saurus", meaning "reptile". The specific name honours the Saraiki people, who primarily live in southern Pakistan. However, like most dinosaur taxa named by M. Sadiq Malkani, it is probably a ''nomen nudum''. Some authors consider "Vitakrisaurus" to be the same animal as "Vitakridrinda".
W
White Rock spinosaurid
"White Rock spinosaurid" is the nickname of a giant spinosaur from the Vectis Formation of the Isle of Wight
The Isle of Wight (Help:IPA/English, /waɪt/ Help:Pronunciation respelling key, ''WYTE'') is an island off the south coast of England which, together with its surrounding uninhabited islets and Skerry, skerries, is also a ceremonial county. T ...
described in 2022. Its remains are so fragmentary that the describers refrained from naming it, but considered the name "Vectispinus". With vertebrae comparable in dimensions to ''Spinosaurus'', it was likely among the largest theropods with a length exceeding .
X
Xinghesaurus
"Xinghesaurus" was the name given to a species of sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur, possibly a titanosauriform, in 2009, in the guidebook for the dinosaur expo "Miracle of Deserts", written by Hasegawa et al. No species name was given for the genus. Based on the skeletal mount, "Xinghesaurus" was likely around long and weighed around .
Y
Yibinosaurus
"Yibinosaurus" (meaning "Yibin lizard") is the nomen nudum, informal name given to an as yet undescribed genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of herbivorous
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
dinosaur from the Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
. It was a sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
which lived in what is now Sichuan, China. The suggested "type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
", "Yibinosaurus zhoui", is briefly mentioned in the Chongqing Natural History Museum guidebook (2001) as under description by Chinese paleontologist Ouyang Hui. It was coined as a ''nomen ex dissertationae'' by Ouyang (2003), and is based on a specimen referred to ''Gongxianosaurus'' sp. nov. by Luo and Wang (1999).
Yunxianosaurus
"Yunxianosaurus" is the provisional name for a genus of titanosaurian dinosaurs from the Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of what is now Hubei, China. The type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, "Yunxianosaurus hubeinensis", was proposed by Chinese paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Li Zhengqi in 2001. The fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s of "Yunxianosaurus" were found near the Nanyang, Henan, Nanyang Prefecture. Li stated that the name "Yunxianosaurus" was a temporary label for ease of description, but that further field work and study of the fossils would be required before the genus could be given an official name.[Li, Zhengqi. (2001). Distribution, burying and classification of dinosaur fossils in Upper Cretaceous strata at Meipu Town, Yunxian County of Hubei Province. ''Hubei Geology & Mineral Resources'', 15(4)(Total No 37): 25–31]
/ref>
Z
Zamyn Khondt oviraptorid
Zamyn Khondt oviraptorid is a nickname for oviraptorid specimen IGM or GIN 100/42. Since the type skull of ''Oviraptor'' is so poorly preserved and crushed, the skull of IGM 100/42 has become the quintessential depiction of that dinosaur, even appearing in scientific papers with the label ''Oviraptor philoceratops''.[Barsbold, R., Teresa Maryańska, Maryanska, T., and Osmolska, H. (1990). "Oviraptorosauria," in Weishampel, D.B., Dodson, P., and Osmolska, H. (eds.). ''The Dinosauria''. Berkeley: University of California Press, pp. 249–258.] However, this distinctive-looking, tall-crested species has more features of the skull in common with ''Citipati'' than it does with ''Oviraptor'' and it may represent a second species of ''Citipati osmolskae, Citipati'' or possibly an entirely new genus, pending further study.[Clark, J.M., Norell, M.A., & Barsbold, R. (2001). "Two new oviraptorids (Theropoda:Oviraptorosauria), upper Cretaceous Djadokhta Formation, Ukhaa Tolgod, Mongolia." ''Journal of Vertebrate Paleontology'' 21(2):209–213., June 2001.]
See also
*List of informally named Mesozoic reptiles
*List of dinosaur genera
References
External links
Theropod Database Blog post clarifying sauropod ''nomina nuda'' from Zhao (1985)
{{DEFAULTSORT:Informally named dinosaurs
Lists of dinosaurs
Lists of prehistoric animal genera (alphabetic)
Nomina nuda