''Leptocleidus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...

, belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Leptocleididae Leptocleididae is a family of small-sized plesiosaurs that lived during the Early Cretaceous period (early Berriasian to early Albian stage). They had small bodies with small heads and short necks. '' Leptocleidus'' and '' Umoonasaurus'' had roun ...

.Smith AS, Dyke GJ. 2008. The skull of the giant predatory pliosaur ''Rhomaleosaurus cramptoni'': implications for plesiosaur phylogenetics. ''Naturwissenschaften'' e-published 2008. It was a small plesiosaur, measuring only up to .

Discovery

In short, the term ''Leptocleidus'' means "slender clavicle". It comes from a merge of the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

words ''λεπτοσ'', meaning "slender" and ''κλειδ'' (also spelled ''κλεισ'') meaning

clavicle The clavicle, collarbone, or keybone is a slender, S-shaped long bone approximately long that serves as a strut between the scapula, shoulder blade and the sternum (breastbone). There are two clavicles, one on each side of the body. The clavic ...

. ''Leptocleidus'' is known from the following sediments: *''L. capensis'' is known from the Sundays River Formation (upper

Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 137.05 ± 0.2 Ma and 132.6 ± 0.2 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretac ...

age),

Cape Province The Province of the Cape of Good Hope (), commonly referred to as the Cape Province () and colloquially as The Cape (), was a province in the Union of South Africa and subsequently the Republic of South Africa. It encompassed the old Cape Co ...

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

. *''L. clemai'' found near Kalbarri in the Carnarvon Basin (

Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...

Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...

age)

Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...

. *''L. superstes'' is known from the Upper Weald Clay (

age),

Sussex, England Sussex ( /ˈsʌsɪks/; from the Old English ''Sūþseaxe''; lit. 'South Saxons'; 'Sussex') is an area within South East England that was historically a kingdom and, later, a county. It includes the ceremonial counties of East Sussex and We ...

. A specimen from the

Vectis Formation The Vectis Formation is a geological Formation (geology), formation on the Isle of Wight and Swanage, England whose strata were formed in the Barremian, approximately 125 million years ago."Magnetostratigraphy of the Lower Cretaceous Vectis Forma ...

(lower

Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...

age),

Isle of Wight The Isle of Wight (Help:IPA/English, /waɪt/ Help:Pronunciation respelling key, ''WYTE'') is an island off the south coast of England which, together with its surrounding uninhabited islets and Skerry, skerries, is also a ceremonial county. T ...

, found in 1995 and seen as a "Leptocleidus sp.", was named as a separate genus ''

Vectocleidus ''Vectocleidus'' is an extinct genus of leptocleidid plesiosaurian known from the Early Cretaceous Vectis Formation (late Barremian stage) of Isle of Wight, in the United Kingdom. It contains a single species A species () is often defined ...

'' in 2012.

Description

With large

s and

interclavicle An interclavicle is a bone which, in most tetrapods, is located between the clavicles. Therian mammals ( marsupials and placentals) are the only tetrapods which never have an interclavicle, although some members of other groups also lack one. In ...

and small

scapula The scapula (: scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on either side ...

e, ''Leptocleidus'' resembled the Early

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...

Rhomaleosaurus ''Rhomaleosaurus'' (meaning "strong lizard") is an extinct genus of Early Jurassic (Toarcian Faunal stage, age, about 183 to 175.6 million years ago) rhomaleosaurid pliosauroid known from Northamptonshire and from Yorkshire of the United Kingdom. ...

'' and members of the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...

Polycotylidae Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. They are known as false pliosaurs. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widesprea ...

. The animal had 21

teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

on either side of its

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

and approximately 35 teeth on each side of the

mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

. The ''Leptocleidus triangle-shaped

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

had a crest running from a ridge on the end of the

nose A nose is a sensory organ and respiratory structure in vertebrates. It consists of a nasal cavity inside the head, and an external nose on the face. The external nose houses the nostrils, or nares, a pair of tubes providing airflow through the ...

to the nasal region. Differing from

pliosaurids Pliosauridae is a family (biology), family of plesiosaurian marine reptiles from the Late Triassic, Latest Triassic to the early Late Cretaceous (Rhaetian to Turonian stages). The family is more inclusive than the archetypal short-necked large he ...

, ''Leptocleidus'' had single-headed

cervical rib Cervical ribs are the ribs of the neck in many tetrapods. In most mammals, including humans, cervical ribs are not normally present as separate structures. They can, however, occur as a pathology. In humans, pathological cervical ribs are usually no ...

s and a deep depression in the

centra Centra is a convenience shop chain that operates throughout Ireland. The chain operates as a symbol group owned by Musgrave Group, the food wholesaler, meaning the individual shops are all owned by individual franchisees. The chain has three ...

of the

neck The neck is the part of the body in many vertebrates that connects the head to the torso. It supports the weight of the head and protects the nerves that transmit sensory and motor information between the brain and the rest of the body. Addition ...

vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...

e. ''Leptocleidus'', unlike many

s, lived in shallow

lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') an ...

s and likely visited

brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...

and

fresh water Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salt (chemistry), salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include ...

systems (such as the mouths of large rivers). This led Arthur Richard Ivor Cruickshank to infer that this movement to fresh water was an attempt to flee larger plesiosaurs and pliosaurs. Most species are known from The British Isles but '' L. capensis'' was discovered in

Classification

Cladogram based on Ketchum and Benson (2011):

References

External links

''Leptocleidus'' on DinoWight
*http://plesiosauria.com/leptocleidus *https://web.archive.org/web/20070922021505/http://www.palaeos.com/Vertebrates/Units/220Lepidosauromorpha/220.520.html#Leptocleidus *https://web.archive.org/web/20070922061721/http://www.plesiosaur.com/database/genusIndividual.php?i=73 {{Taxonbar, from=Q41154953 Plesiosaurs Early Cretaceous plesiosaurs of Europe Early Cretaceous plesiosaurs Fossil taxa described in 1922 Early Cretaceous reptiles of Australia Plesiosaurs of Oceania Plesiosaurs of Africa Taxa named by Charles William Andrews Sauropterygian genera

Discovery

Description

Classification

See also

References

Further reading

External links