Coahuilasaurus
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''Coahuilasaurus'' (meaning "Coahuila lizard") is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of kritosaurin
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
dinosaur from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
) Cerro del Pueblo Formation of
Coahuila Coahuila, formally Coahuila de Zaragoza, officially the Free and Sovereign State of Coahuila de Zaragoza, is one of the 31 states of Mexico. The largest city and State Capital is the city of Saltillo; the second largest is Torreón and the thi ...
, Mexico. The genus contains a single species, ''C. lipani'', known from the associated tips of the upper and lower jaw and other fragmentary skull bones. It is a large kritosaurin with an estimated body length of .


Discovery and naming

The holotype specimen, IGM 6685, is a partial skull. It was originally thought to belong to the same genus as PASAC 1, a large unnamed specimen known as the "Sabinas hadrosaurid", but this referral was later rejected. IGM 6685 was later referred to as a specimen of ''
Kritosaurus ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
''. IGM 6685 was later named as a new genus and species of kritosaurin hadrosaurid in 2024. The generic name, ''Coahuilasaurus'', honors the Mexican state of Coahuila, where the fossils were found. The specific name, ''lipani'', honors the Lépai-Ndé or Lipani, a tribe of
Apache The Apache ( ) are several Southern Athabaskan language-speaking peoples of the Southwestern United States, Southwest, the Southern Plains and Northern Mexico. They are linguistically related to the Navajo. They migrated from the Athabascan ho ...
Native Americans known from the area.


Classification

Longrich et al. added ''Coahuilasaurus'' to a
phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
which found it to be a member of the Kritosaurini, while the Sabinas hadrosaur was found to be a member of Saurolophini. A cladogram adapted from that analysis is shown below:


Paleoenvironment

The Cerro del Pueblo Formation is the oldest member of the Difunta Group, which is believed to have been deposited in the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
. It is primarily made up of alternating layers of
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
s,
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
s, and gray
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of Clay mineral, clay minerals (hydrous aluminium phyllosilicates, e.g., Kaolinite, kaolin, aluminium, Al2Silicon, Si2Oxygen, O5(hydroxide, OH)4) and tiny f ...
s. These rocks preserve
fluvial A river is a natural stream of fresh water that flows on land or inside caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of its course if it ru ...
,
lacustrine A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from t ...
,
coast A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...
al, and
shallow Shallow may refer to: Places * Shallow (underwater relief), where the depth of the water is low compared to its surroundings * Shallow Bay (disambiguation), various places * Shallow Brook, New Jersey, United States * Shallow Inlet, Victoria ...
marine ecosystems. The shale layers primarily preserve marine
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s such as
ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
s and
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
s and they interbedded with sandstone and siltstone layers, suggesting that sea levels were fluctuating over the time that the formation was deposited. The terrestrial deposits, in which the contemporary ornithomimosaur ''
Paraxenisaurus ''Paraxenisaurus'' (, meaning "strange lizard") is an extinct genus of ornithomimosaurian theropod from the Late Cretaceous Cerro del Pueblo Formation of Coahuila in Mexico. The genus contains a single species, ''P. normalensis'', which is know ...
'' was found, are believed to represent a coastal
floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river. Floodplains stretch from the banks of a river channel to the base of the enclosing valley, and experience flooding during periods of high Discharge (hydrolog ...
on the margin of an
estuary An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime enviro ...
. The presence of a diverse array of crocodiles, turtles, and freshwater
bivalves Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed by a calcified exoskeleton consis ...
suggests a heavily vegetated riverine ecosystem. The abundance of dinosaur teeth and bone fragments also suggests that the region was replete with vegetation and supported a diverse assemblage of
megafauna In zoology, megafauna (from Ancient Greek, Greek μέγας ''megas'' "large" and Neo-Latin ''fauna'' "animal life") are large animals. The precise definition of the term varies widely, though a common threshold is approximately , this lower en ...
.


Contemporary fauna

Dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
remains are abundant in the Cerro del Pueblo Formation, but most of the known remains are very incomplete. Many of these dinosaurs are known only from teeth, and are therefore very difficult to assign to any specific
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. Teeth from
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
s are very common and have shown that
tyrannosaurid Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to fifteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera ...
s,
dromaeosaurid Dromaeosauridae () is a family (biology), family of feathered coelurosaurian Theropoda, theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous period (geology), Period. The name Drom ...
s,
troodontids Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinos ...
, and caenagnathids lived in this environment in addition to the above-mentioned ornithomimosaurs.
Ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
ns were also common and diverse in the ecosystem, much like the contemporaneous
Dinosaur Park Dinosaur Park is a dinosaur park in Rapid City, South Dakota, United States. Dedicated on May 22, 1936, it contains seven dinosaur sculptures on a hill overlooking the city, created to capitalize on the tourists coming to the Black Hills to see ...
,
Two Medicine Two Medicine is the collective name of a region located in the southeastern section of Glacier National Park (U.S.), Glacier National Park, in the U.S. state of Montana. It has a campground alongside Two Medicine Lake. From the period starting in ...
,
Kaiparowits The Paiute Indian Tribe of Utah is a federally recognized tribe of Southern Paiute and Ute Indians in southwestern Utah. Reservation The Paiute Indian Tribe of Utah (PITU) has a reservation composed of ten separate parcels of land, located in ...
, and Kirtland formations.
Hadrosaur Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
remains are the most common dinosaur fossil material found in the Cerro del Pueblo, being known from
vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...
e, limb bones, jawbones, teeth, and shoulder bones. Some of these taxa have been named and described, such as '' Tlatolophus'', ''
Velafrons ''Velafrons'' (meaning "sailed forehead") is a genus of lambeosaurine hadrosaurid dinosaur from the Late Cretaceous of Mexico. It is known from a mostly complete skull and partial skeleton of a juvenile individual, with a bony crest on the for ...
'', and ''
Latirhinus ''Latirhinus'' (meaning "broad nose" from the Latin ''latus'' (broad) and Greek ῥίς, ''rhis'' (nose)) is an extinct genus of lambeosaurine hadrosaurid dinosaur from the Late Cretaceous of Mexico. The type species, ''Latirhinus uitstlani'', ...
'', but most remains have yet to be formally named or described, and additional species may have existed.
Ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
s are represented by several unnamed taxa, and
ceratopsid Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', '' Centrosaurus'', and '' Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are ...
s like ''
Coahuilaceratops ''Coahuilaceratops'' (meaning "horned face from Coahuila") is a genus of chasmosaurine ceratopsian dinosaur that lived during the early Maastrichtian age of the Late Cretaceous epoch, about 71.5 to 70.5 million years ago in what is now northern M ...
'' are known to have coexisted with both
chasmosaurine Chasmosaurinae is a subfamily of ceratopsid dinosaurs. They were one of the most successful groups of herbivores of their time. Chasmosaurines appeared in the early Campanian, and became extinct, along with all other non- avian dinosaurs, during ...
s and
centrosaurine Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid, a group of large quadrupedal ornithischian dinosaur. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alber ...
s. Fossilized footprints from some kind of
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
are also known from this formation. Like most
fluvial A river is a natural stream of fresh water that flows on land or inside caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of its course if it ru ...
sediments Sediment is a solid material that is transported to a new location where it is deposited. It occurs naturally and, through the processes of weathering and erosion, is broken down and subsequently sediment transport, transported by the action of ...
from the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
of
Laramidia Laramidia was an island continent that existed during the Late Cretaceous period (99.6–66 Year#SI prefix multipliers, Ma), when the Western Interior Seaway split the continent of North America in two. In the Mesozoic era, Laramidia was an island ...
, the Cerro del Pueblo Formation was home to a wide array of
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s including pleurodires,
paracryptodires __NOTOC__ Paracryptodira is an extinct group of reptiles in the clade Testudinata (which contains modern turtles and their extinct relatives), known from the Jurassic to Paleogene of North America and Europe. Initially treated as a suborder siste ...
, cheloniids,
kinosternid The Kinosternidae are a family of mostly small turtles that includes the mud turtles and musk turtles. The family contains 25 species within four genera, but taxonomic reclassification is an ongoing process, so many sources vary on the exact numb ...
s, and
trionychid Trionychidae is a family of turtles, commonly known as softshell turtles or simply softshells. The family was described by Leopold Fitzinger in 1826. Softshells include some of the world's largest freshwater turtles, though many can adapt to liv ...
s. These would have coexisted with both goniopholids and
eusuchia Eusuchia is a clade of neosuchian Crocodylomorpha, crocodylomorphs that first appeared in the Early Cretaceous, which includes modern Crocodilia, crocodilians. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survi ...
n
crocodyliformes Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseu ...
in both freshwater and saltwater environments. There is also some evidence of snakes from this environment. Microfossils have also preserved
gastropods Gastropods (), commonly known as slugs and snails, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and from the land. Ther ...
,
ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
s,
bivalves Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed by a calcified exoskeleton consis ...
, and other invertebrates.


See also

*
Timeline of hadrosaur research This timeline of hadrosaur research is a chronological listing of events in the History of paleontology, history of paleontology focused on the hadrosauroids, a group of herbivorous ornithopod dinosaurs popularly known as the duck-billed dinosa ...
*
2024 in archosaur paleontology This article records new taxa of every kind of fossil archosaur that are scheduled to be Species description, described during 2024, as well as other significant discoveries and events related to the paleontology of archosaurs published in 2024. ...


References

{{Taxonbar, from=Q130215835 Saurolophinae Dinosaur genera Campanian dinosaurs Taxa named by Nicholas R. Longrich Fossil taxa described in 2024 Dinosaurs of Mexico