Anoplotheriid
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Anoplotheriidae is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
artiodactyl Artiodactyls are placental mammals belonging to the order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other t ...
ungulates. They were endemic to Europe during the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
and
Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
epochs about 44—30 million years ago. Its name is derived from the ("unarmed") and θήριον ("beast"), translating as "unarmed beast".


Ecology

Species of Anoplotheriidae varied substantially in size''. Diplobune minor'' is suggested to have weighted about '','' while ''Anoplotherium'' is suggested to have been up to in weight. ''
Anoplotherium ''Anoplotherium'' is the type genus of the extinct Paleogene, Palaeogene artiodactyl family Anoplotheriidae, which was endemic to Western Europe. It lived from the Late Eocene to the earliest Oligocene. It was the fifth fossil mammal genus to be ...
'' is thought to have been a browser that reared up on its hind legs to feed, while ''
Diplobune ''Diplobune'' (Ancient Greek: (double) + (hill) meaning "double hill") is an extinct genus of Paleogene, Palaeogene artiodactyls belonging to the family Anoplotheriidae. It was endemic to Europe and lived from the late Eocene to the early Olig ...
'' is suggested to have been an
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
climbing animal.


Systematics and taxonomy

The family Anoplotheriidae was assigned to Belluae by Bonaparte (who named it Anoplotheriina) in 1850; to
Artiodactyla Artiodactyls are placental mammals belonging to the order (biology), order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof ...
by
Cope A cope ( ("rain coat") or ("cape")) is a liturgical long mantle or cloak, open at the front and fastened at the breast with a band or clasp. It may be of any liturgical colour. A cope may be worn by any rank of the Catholic or Anglican clerg ...
in 1889, to
Ruminantia Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principa ...
by
Gregory Gregory may refer to: People and fictional characters * Gregory (given name), including a list of people and fictional characters with the given name * Gregory (surname), a surname *Gregory (The Walking Dead), fictional character from the walkin ...
in 1910, and finally to its own
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Anoplotherioidea by
Romer Romer, Römer, Roemer, or similar may refer to: * Romer (surname), includes a list of people with the name * Romer (tool), a cartographic device also known as a reference card * Rømer scale, a disused temperature scale named after Ole Rømer * ...
in 1966. A 2019 study considered them to be closely related to
Cainotheriidae Cainotheriidae is an extinct family of artiodactyls known from the Late Eocene to Middle Miocene of Europe. They are mostly found preserved in karstic deposits. These animals were small in size, and generally did not exceed in height at the s ...
, another group of endemic European artiodactyls, with this group in turn being related to
ruminants Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principa ...
, while a 2020 study found them to be more closely related to the also European endemic
Xiphodontidae Xiphodontidae is an extinct family (biology), family of herbivorous even-toed ungulates (order (biology), order Artiodactyla), endemic to Europe during the Eocene 40.4—33.9 million years ago, existing for about 7.5 million years. ''P ...
, again as relatives of ruminants. Included genera: * Subfamily Anoplotheriinae ** ''
Anoplotherium ''Anoplotherium'' is the type genus of the extinct Paleogene, Palaeogene artiodactyl family Anoplotheriidae, which was endemic to Western Europe. It lived from the Late Eocene to the earliest Oligocene. It was the fifth fossil mammal genus to be ...
'' Cuvier, 1804 ** ''
Diplobune ''Diplobune'' (Ancient Greek: (double) + (hill) meaning "double hill") is an extinct genus of Paleogene, Palaeogene artiodactyls belonging to the family Anoplotheriidae. It was endemic to Europe and lived from the late Eocene to the early Olig ...
'' Rutimeyer, 1862 ** ''
Duerotherium ''Duerotherium'' is an extinct genus of artiodactyl that lived during the Middle Eocene and is only known from the Iberian Peninsula. The genus is a member of the family Anoplotheriidae and the subfamily Anoplotheriinae, and contains one speci ...
'' Cuesta & Badiola, 2009 ** ''
Ephelcomenus ''Ephelcomenus'' is an extinct genus of Paleogene, Palaeogene artiodactyls belonging to the Anoplotheriidae that were endemic to Western Europe. It contains one species ''E. filholi'', which was first described by Richard Lydekker in 1889 but e ...
'' Hurzeler, 1938 ** ''
Robiatherium ''Robiatherium'' is an extinct genus of Palaeogene artiodactyls containing one species ''R. cournovense''. The genus name derives from the locality of Robiac in France, where some of its fossils were described, plus the Greek /, meaning "beast" ...
'' Sudre, 1988 * Subfamily Dacrytheriinae ** ''
Dacrytherium ''Dacrytherium'' (Ancient Greek: (tear, teardrop) + (beast or wild animal) meaning "tear beast") is an extinct genus of Paleogene, Palaeogene artiodactyls belonging to the family Anoplotheriidae. It occurred from the Middle to Late Eocene of W ...
'' Filhol, 1876 ** ''
Catodontherium ''Catodontherium'' is an extinct genus of Paleogene, Palaeogene artiodactyl belonging to the family Anoplotheriidae. It was endemic to Western Europe and had a temporal range exclusive to the middle Eocene, although its earliest appearance depe ...
'' Depéret, 1908


References


Sources

* http://www.helsinki.fi/~mhaaramo/metazoa/deuterostoma/chordata/synapsida/eutheria/artiodactyla/suina/anoplotheroidea.html {{Taxonbar, from=Q567678 Tylopoda Eocene Artiodactyla Eocene first appearances Prehistoric mammals of Europe Rupelian extinctions Prehistoric mammal families