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''Ajkaceratops'' (pronounced "oi-ka-sera-tops") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
described in 2010. It lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
in the western Tethyan archipelago, in what is now
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''A. kozmai'', was originally described as a
ceratopsian Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period, although ancestral forms lived earlier, in the Late Ju ...
most closely related to forms in east
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, from where its ancestors may have migrated by island-hopping. Later research however has questioned this assignment, and treats ''Ajkaceratops'' as an ornithischian of unresolved affinity.


Discovery

The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, cataloged as MTM V2009.192.1, consists only of a few skull fragments, including snout with proposed rostral bone, fused premaxillae, and maxillae fragments (beak and jaw fragments). These fossils are kept in the Hungarian Natural History Museum, in
Budapest Budapest is the Capital city, capital and List of cities and towns of Hungary, most populous city of Hungary. It is the List of cities in the European Union by population within city limits, tenth-largest city in the European Union by popul ...
. The generic name, ''Ajkaceratops'', honors
Ajka Ajka () is a city in Hungary with about 35,000 inhabitants. It is situated in the hills of Bakony. History Around 1000 BCE, the area was inhabited by Celts. By the second century CE, the territory was conquered by the Roman Empire, Romans. The Hu ...
, a town in
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
where the fossils were first discovered, combined with the given greek nomination '' ceratops'', meaning "horned face". The specific name, "''kozmai"'', honors Károly Kozma.


Description

Although the fossils are fragmentary, the paper describing ''Ajkaceratops'' estimated a body length of . Other material includes four predentary bones, cataloged as MTM V2009.193.1, V2009.194.1, V2009.195.1, and V2009.196.1; these are also believed to have belonged to ''Ajkaceratops'', although they are proportionately smaller, and probably came from other individuals of the genus.


Classification

When first described, the fossils were stated to most closely resemble those of the Asian
protoceratopsid Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most spec ...
''
Bagaceratops ''Bagaceratops'' (meaning "small-horned face") is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 72 to 71 million years ago. ''Bagaceratops'' remains have been reported from the Barun Goyot Forma ...
''. Those similarities indicate ''Ajkaceratops'' is a
ceratopsia Ceratopsia or Ceratopia ( or ; Ancient Greek, Greek: "horned faces") is a group of herbivore, herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period (geology), Period, although ance ...
n related to the protoceratopsids, but more primitive than the '' Zuniceratops'' and the
Ceratopsidae Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', ''Centrosaurus'', and ''Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are k ...
. However, more recent research has questioned this referral. The proposed rostral bone of ''Ajkaceratops'' differs markedly from the rostral morphology of other ceratopsians, possessing a sub-circular cross section, pitted texture instead of grooves, and a lack of lateral slicing edges. Furthermore, the proposed rostral bone appears to have fully fused with the premaxilla, a feature that is only seen in the ontogenetically oldest members of derived ceratopsids. This has cast doubt as to whether ''Ajkaceratops'' possesses a rostral bone in the first place, a defining feature of the ceratopsian group, and instead the hooked beak represents a highly derived, hypertrophied premaxilla. While there is the possibility ''Ajkaceratops'' is a highly unique ceratopsian, it was referred to as an enigmatic ornithischian of uncertain affinities by the authors. In a redescription of the cranial anatomy of the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
neornithischian '' Jeholosaurus'', Bertozzo ''et al.'' tested the phylogenetic position of ''Ajkaceratops'' based on the premaxilla interpretation and recovered it as a sister taxon to '' Changmiania'', implying a position as a basal
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
.


Palaeoenvironment

The fossils of ''Ajkaceratops'' were discovered in the Csehbánya Formation, which is interpreted as a floodplain and channel deposit formed by variegated clay, silt with interbedded grey and brown sand, and sandstone beds. This strata dates to the
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
stage, around 86 to 84 million years ago. ''Ajkaceratops'' shared its environment with other dinosaurs such as '' Mochlodon'', the nodosaurid
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
s, other non-avian
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
s and enantiornithine birds, as well as
eusuchia Eusuchia is a clade of neosuchian Crocodylomorpha, crocodylomorphs that first appeared in the Early Cretaceous, which includes modern Crocodilia, crocodilians. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survi ...
n crocodiles, azhdarchid
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
s, bothremydid turtles and teiid lizards.


See also

* Timeline of ceratopsian research *
List of dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 annum, million years ago, although the exact origin and timing of the evolution of ...
* 2010 in paleontology * 2024 in paleontology * Haţeg Island


References

{{Taxonbar, from=Q516014 Ceratopsia Dinosaur genera Santonian dinosaurs Dinosaurs of Europe Fossil taxa described in 2010 Taxa named by David B. Weishampel