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Ceratopsian
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period, although ancestral forms lived earlier, in the Late Jurassic of Asia. The earliest known ceratopsian, '' Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, '' Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was '' Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no disti ...
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Triceratops
''Triceratops'' ( ; ) is a genus of Chasmosaurinae, chasmosaurine Ceratopsia, ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous Period (geology), period, about 68 to 66 million years ago on the island continent of Laramidia, now forming western North America. It was one of the last-known non-avian dinosaurs and lived until the Cretaceous-Paleogene extinction event, Cretaceous–Paleogene extinction event 66 million years ago. The name ''Triceratops'', which means 'three-horned face', is derived from the Ancient Greek, Greek words () meaning 'three', () meaning 'horn', and () meaning 'face'. Bearing a large bony neck frill, frill, three horns on the skull, and a large, four-legged body, exhibiting convergent evolution with bovines and rhinoceroses, ''Triceratops'' is one of the most recognizable of all dinosaurs and the best-known ceratopsian. It was also one of the largest, measuring around long and weighing up to . It shared the la ...
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Psittacosaurus
''Psittacosaurus'' ( ; "parrot lizard") is a genus of extinct ceratopsian dinosaur from the Early Cretaceous of what is now Asia, existing between 125 and 105 million years ago. It is notable for being the most species-rich non-avian dinosaur genus. Up to 12 species are known, from across China, Mongolia, Russia, and Thailand. The species of ''Psittacosaurus'' were obligate bipeds at adulthood, with a high skull and a robust beak. One individual was found preserved with long filaments on the tail, similar to those of '' Tianyulong''. ''Psittacosaurus'' probably had complex behaviours, based on the proportions and relative size of the brain. It may have been active for short periods of time during the day and night, and had well-developed senses of smell and vision. ''Psittacosaurus'' was one of the earliest ceratopsians, but closer to ''Triceratops'' than '' Yinlong''. Once in its own family, Psittacosauridae, with other genera like ''Hongshanosaurus'', it is now considered t ...
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Protoceratopsidae
Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most specimens found in China and Mongolia. As ceratopsians, protoceratopsids were herbivorous, with constantly replacing tooth batteries made for slicing through plants and a hooked beak for grabbing them. Protoceratopsids were small ceratopsians around in length. Their bony frill and horns were much smaller than more derived members of Ceratopsia, such as ceratopsids. Description Protoceratopsids were relatively small ceratopsians, averaging around 1-2.5 m in length from head to tail. Protoceratopsids have a frill and rostral bone characteristic of all ceratopsians. Their snout is particularly wedge-shaped with tall and narrow nostrils situated high on it. The antorbital fenestra is unusually small, and the antorbital fossa sits high on the sku ...
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Auroraceratops
''Auroraceratops'', meaning "dawn horned face", is a genus of bipedal basal neoceratopsian dinosaur, from the Early Cretaceous (Aptian age) of north central China. The etymology of the generic name refers to its status as an early ceratopsian and also to Dawn Dodson, wife of Peter Dodson, one of the palaeontologists who described it. Discovery and species Expeditions into the Gansu Province of northwestern China began with the Sino-Swedish Expedition of 1930 to 1931, where discoveries of dinosaurs including the now-dubious early ceratopsian '' Microceratops sulcidens''. These discoveries were followed by occasional observations of dinosaur bones in the Houhongquan Basin in the 1960s, and then the Gongpoquan Basin in 1986. Such observations led to the China-Canada Dinosaur Project taking a reconnaissance trip to the Gongpoquan Basin in 1988, but no further expeditions were led until the Sino-Japanese Silk Road Dinosaur Expedition of 1992 and 1993, led by Chinese paleontolog ...
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Liaoceratops
''Liaoceratops'', meaning "Liaoning horned face", is a ceratopsian dinosaur believed to be an early relative of the horned ceratopsids. It lived in the Early Cretaceous, 126 million years ago. It was discovered in China by a team of American and Chinese scientists. ''Liaoceratops'' was much smaller than its later relatives, but offers a glimpse into the early evolution of this group of dinosaurs. Discoveries and naming ''Liaoceratops'' was discovered in the famous Liaoning Province of China, where several fossils of feathered dinosaurs have also been collected. The type species ''Liaoceratops yanzigouensis'' was in 2002 named and described by Xu Xing, Peter Makovicky, Wang Xiaolin, Mark Norell and You Hailu. The generic name is derived from Liaoning and the Greek ''keras'', "horn" and ''ops'', "face". The specific name refers to the town Yanzigou. The holotype IVPP V12738 has been found in the Yixian Formation dating from the Barremian. These beds have also yielded fossi ...
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Zuniceratops
''Zuniceratops'' ('Zuni-horned face') is a genus of ceratopsian dinosaurs that lived during the Turonian stage of the Late Cretaceous in what is now New Mexico, United States. Only a single species is known, ''Zuniceratops christopheri''. History of discovery ''Zuniceratops'' was discovered in 1996, by eight-year-old Christopher James Wolfe, son of Paleontology, paleontologist Douglas Gerald Wolfe, Douglas G. Wolfe, in the Moreno Hill Formation in west-central New Mexico where one skull and the bones from several individuals have been found. This discovery of ''Zuniceratops'' bonebed has been suggested as one of the evidence for the claim that grouping behavior could be a Apomorphy and synapomorphy, synapomorphic trait for ceratopsians. In 2001, a bone believed to be a squamosal has since been found to be an ischium of a ''Nothronychus''. The holotype specimen, MSM P2101, is either a juvenile or a subadult, while other specimens like MSM P2101 and MSM P3812 belong to adults. Th ...
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Ajkaceratops
''Ajkaceratops'' (pronounced "oi-ka-sera-tops") is a genus of ornithischian dinosaur described in 2010. It lived during the Late Cretaceous in the western Tethys Ocean, Tethyan archipelago, in what is now Europe. The type species, ''A. kozmai'', was originally described as a ceratopsian most closely related to forms in east Asia, from where its ancestors may have migrated by Oceanic dispersal, island-hopping. Later research however has questioned this assignment, and treats ''Ajkaceratops'' as an ornithischian of unresolved affinity. Discovery The holotype, cataloged as MTM V2009.192.1, consists only of a few skull fragments, including snout with proposed rostral bone, fused premaxillae, and maxillae fragments (beak and jaw fragments). These fossils are kept in the Hungarian Natural History Museum, in Budapest. The generic name, ''Ajkaceratops'', honors Ajka, a town in Hungary where the fossils were first discovered, combined with the given greek nomination ''ceratops'', meaning ...
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Mosaiceratops
''Mosaiceratops'' is a genus of ceratopsian, described by Zheng, Jin & Xu in 2015 and found in the Xiaguan Formation of Neixiang County. ''Mosaiceratops'' lived in the upper Cretaceous in what is now the Henan Province of China. Although phylogenetic analyses have found ''Mosaiceratops'' to be the most basal neoceratopsian, the authors noted that several features in the premaxilla and nasal bones are shared with ''Psittacosaurus'', indicating that neoceratopsians evolved premaxillary teeth twice and that ''Psittacosaurus'' is not as primitive as previously thought.Zheng, W., Jin, X., & Xu, X. (2015)A psittacosaurid-like basal neoceratopsian from the Upper Cretaceous of central China and its implications for basal ceratopsian evolution. ''Scientific reports'', 5. article number 14190: 1-9; doi:10.1038/srep14190 Discovery On the westbank of the river Tuanhe in Neixiang in Henan the skeleton was discovered of a small ceratopian. The fossil was prepared by Sheng Yiming and Yu Cha ...
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Leptoceratopsidae
Leptoceratopsidae is an extinct family (biology), family of neoceratopsian dinosaurs from Asia, North America and possibly Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the family (biology), families Protoceratopsidae and Ceratopsidae, but they were more primitive and generally smaller. Phylogeny Leptoceratopsidae was originally named by Franz Nopcsa von Felső-Szilvás in 1923 as a subfamily Leptoceratopsinae, and its type species is ''Leptoceratops gracilis''. Mackovicky, in 2001, defined it as a stem-based taxon and a family consisting of ''Leptoceratops gracilis'' and all species closer to ''Leptoceratops'' than to ''Triceratops horridus''.Makovicky, P.J. 2001. A ''Montanoceratops cerorhynchus'' (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta, In: Tanke, D.H. & Carpenter, K. (Eds.). ''Mesozoic Vertebrate Life''. Bloomington: Indiana University Press. Pp. 243-262. All previously published neo ...
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Asiaceratops
''Asiaceratops'' (meaning "Asian horned face") is a genus of herbivorous ceratopsian dinosaur. It lived during the Early-Late Cretaceous. The type species, ''A. salsopaludalis'' is known from Uzbekistan, while ''A. sulcidens'' is known from China and Mongolia. Discovery and naming The type species, ''Asiaceratops salsopaludalis'', was formally described by Lev Nesov, L.F. Kaznyshkina and Gennadiy Olegovich Cherepanov in 1989. The generic name combines a reference to Asia with ''~ceratops'', "horned face". The specific name means "of the salt marsh" in Latin. In the same publication '' Microceratops sulcidens'' Bohlin 1953 was renamed into a second species of ''Asiaceratops'': ''Asiaceratops sulcidens''.L.A. Nessov, L F. Kaznyshkina, and G.O. Cherepanov. (1989). esozoic ceratopsian dinosaurs and crocodiles of central Asia In: Bogdanova and Khozatskii (eds.), ''Theoretical and Applied Aspects of Modern Palaeontology'' pp 144-154 The holotype of ''Asiaceratops salsopaludalis'', C ...
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Beg Tse
''Beg tse'' (after the Himalayan war deity Beg-tse) is an extinct species of neoceratopsian dinosaurs from the early Cretaceous Ulaanoosh Formation of Mongolia. ''B. tse'' is the only species in the genus ''Beg'', known from a partial skull and very fragmentary postcrania. It represents the most basal neoceratopsian currently known. Discovery and naming The holotype, IGM 100/3652, was discovered in 2015 near the town of Tsogt-Ovoo in the Ömnögovi Province of Mongolia. Described from the Ulaanoosh Formation, the specimen is dated to 113 to 94 million years ago, at the boundary of the Lower and Upper Cretaceous. ''Beg'' is named after Beg-tse, a Himalayan deity who is the god of war in Mongolian culture. The deity is often depicted with a rugose face and/or body, similar to the appearance of the preserved skull of the dinosaur. Description Based on the size of the skull, about long, ''Beg'' was most likely a medium-sized basal ceratopsian, similar in size to ''Yin ...
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Microceratus
''Microceratus'' (meaning "small-horned") is a genus of small ceratopsian dinosaur that lived in the Cretaceous Period in Mongolia. It walked on two legs, had short front arms, a characteristic ceratopsian frill and beak-like mouth, and was around long. It was one of the first ceratopsians, or horned dinosaurs, along with '' Psittacosaurus'' also from Mongolia. Discovery The type species, ''Microceratops gobiensis'', was first described by Bohlin in 1953, and so was the second species, ''M. sulcidens'', which may belong to '' Asiaceratops'' instead. However, the generic name was already preoccupied by an ichneumon wasp (subfamily Cryptinae) with the same name. Though much of the material has since been reassigned to the genus '' Graciliceratops'', a replacement name ''Microceratus'' was created by Mateus in 2008 for the type specimen. Classification ''Microceratus'' belonged to the Ceratopsia (Ancient Greek for "horned face"), a group of herbivorous dinosaurs with parrot-li ...
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