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Flora


Plants


Fungi


Paleomycological research

* Fossil sporocarps indistinguishable from sporocarps of members of the extant genus ''
Stemonitis ''Stemonitis'' is a distinctive genus of slime moulds found throughout the world (except Antarctica). They are characterised by the tall brown sporangia, supported on slender stalks, which grow in clusters on rotting wood. The genus was first des ...
'' are described from the Cretaceous amber from
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
by Rikkinen, Grimaldi & Schmidt (2019). * A study on the impact of major historical events such as the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden extinction event, mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million y ...
on the evolution of two major subclasses of lichen-forming fungi ( Lecanoromycetidae and Ostropomycetidae) is published by Huang ''et al.'' (2019). * Description of
crustose lichen Crustose lichens are lichens that form a crust which strongly adheres to the substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cor ...
s from European
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
amber is published by Kaasalainen ''et al.'' (2019). * Fungi belonging to the genera ''
Periconia ''Periconia'' is a genus of sac fungi in the order Pleosporales The Pleosporales is the largest order in the fungal class Dothideomycetes. By a 2008 estimate it contains 23 families, 332 genera and more than 4700 species. The majority of sp ...
'', ''
Penicillium ''Penicillium'' () is a genus of ascomycetous fungi that is part of the mycobiome of many species and is of major importance in the natural environment, in food spoilage, and in food and drug production. Some members of the genus produce ...
'' and ''
Scopulariopsis ''Scopulariopsis'' is a genus of anamorphic fungi that are saprobic and pathogenic to animals. The widespread genus contains 22 species. ''Scopulariopsis'' belongs to the group Hyphomycetes Hyphomycetes are a form classification of fungi, part ...
'', representing the first and the oldest known fossil record of these taxa, are described from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...
by Tischer ''et al.'' (2019).


Sponges


Research

*
Sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate throug ...
spicules and spicule-like structures that probably represent sponge fossils are described from four sections of the Ediacaran-Cambrian boundary interval in the
Yangtze Gorges The Three Gorges () are three adjacent gorges along the middle reaches of the Yangtze River, in the hinterland of the People's Republic of China. With a subtropical monsoon climate, they are known for their scenery. The "Three Gorges Scenic ...
(China) by Chang ''et al.'' (2019). * A study evaluating how distribution patterns of non-lithistid spiculate sponges changed during the Cambrian explosion and the
Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced w ...
is published by Botting & Muir (2019).


New taxa


Cnidarians


Research

* A study on the growth characteristics of three species of
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
corals belonging to the genus '' Agetolites'' from the Xiazhen Formation ( China), and on their implications for inferring phylogenetic relationships of this genus, is published by Sun, Elias & Lee (2019). * A study on a large colonial
rugose Rugose means "wrinkled". It may refer to: * Rugosa, an extinct order of coral, whose rugose shape earned it the name * Rugose, adjectival form of rugae Species with "rugose" in their names * ''Idiosoma nigrum'', more commonly, a black rugose trap ...
coral from the Ordovician Kope Formation (
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virgini ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) is published by Harris ''et al.'' (2019). * A study on the
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
, growth characteristics and phylogenetic relationships of the
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoi ...
tabulate coral Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to ...
'' Halysites catenularius'' is published by Liang, Elias & Lee (2019). * Fossils of tabulate corals without septa, representing the first evidence that unmetamorphosed, slightly indurated
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
sandstones crop out amidst the deposits of the Atlantic Coastal Plain Province of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
, are reported from
South Carolina )'' Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = "Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = G ...
by Landmeyer ''et al.'' (2019). This finding is strongly disputed because all other rocks of Paleozoic age in the study area are greatly metamorphosed, the rocks where the fossils were found are traditionally mapped as the Cretaceous Middendorf Formation, and it is suggested that the fossils in question are the bark of Cretaceous conifers in Cretaceous sandstone, instead of Paleozoic corals in Paleozoic sandstone. * A study aiming to determine whether ecological selection based on physiology, behavior, habitat, etc. played a role in the long-term survival of corals during the late
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
and early
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
is published by Weiss & Martindale (2019). * Fossils of ''
Acropora prolifera ''Acropora prolifera'', the fused staghorn coral, is a branching, colonial, stony coral found in shallow parts of the Caribbean Sea, the Bahamas and southern Florida. Description ''Acropora prolifera'' is very similar to staghorn coral (''Acr ...
'' dating back to the
Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
are reported by Precht ''et al.'' (2019). * A study on the distribution of reef corals during the last interglacial is published by Jones ''et al.'' (2019), who also evaluate the utility of fossil reef coral data for predictions of impact of future climate changes on reef corals. * A study on a problematic fossil specimen from the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
Ponta Grossa Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
), assigned by different authors to the species '' Serpulites sica'' or '' Euzebiola clarkei'', is published by Van Iten ''et al.'' (2019), who interpret this fossil as a
medusozoa Medusozoa is a clade in the phylum Cnidaria, and is often considered a subphylum. It includes the classes Hydrozoa, Scyphozoa, Staurozoa and Cubozoa, and possibly the parasitic Polypodiozoa. Medusozoans are distinguished by having a medusa stag ...
n capable of clonal
budding Budding or blastogenesis is a type of asexual reproduction in which a new organism develops from an outgrowth or bud due to cell division at one particular site. For example, the small bulb-like projection coming out from the yeast cell is kno ...
, and transfer it to the genus '' Sphenothallus''. *The oldest mesophotic coral ecosystems, dating back to middle Silurian, from the Lower Visby Beds on Gotland have been described by Zapalski & Berkowski. These communities, dominated by platy corals give also clues about the onset of coral-algal symbiosis. * Mihaljević (2019) describes new fossil coral collections from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
and
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
of
Sarawak Sarawak (; ) is a state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the Malaysian state of Sabah to the northeast, ...
(
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
),
Negros Island Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
and
Cebu Cebu (; ceb, Sugbo), officially the Province of Cebu ( ceb, Lalawigan sa Sugbo; tl, Lalawigan ng Cebu; hil, Kapuroan sang Sugbo), is a province of the Philippines located in the Central Visayas region, and consists of a main island and ...
(the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
). * A study on the anatomy,
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
and taxonomy of the
Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...
hydrozoa Hydrozoa (hydrozoans; ) are a taxonomic class of individually very small, predatory animals, some solitary and some colonial, most of which inhabit saline water. The colonies of the colonial species can be large, and in some cases the specializ ...
n '' Heterastridium'', based on data from fossil specimens from central
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkm ...
and south
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
, is published by Senowbari-Daryan & Link (2019).


New taxa


Arthropods


Bryozoans


Brachiopods


Molluscs


Echinoderms


Research

* A study on the
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
and phylogenetic relationships of the putative
stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushr ...
-echinoderm '' Yanjiahella biscarpa'' is published by Topper ''et al.'' (2019); the study is subsequently criticized by Zamora ''et al.'' (2020). * Soft tissue traces found in conjunction with skeletal molds are described in stylophorans by Lefebvre ''et al.'' (2019), who interpret their findings as supporting echinoderm and not
hemichordate Hemichordata is a phylum which consists of triploblastic, enterocoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and inclu ...
-like affinities of stylophorans. * A study on the morphology and phylogenetic relationships of the lepidocystoid echinoderm '' Vyscystis'' is published by Nohejlová ''et al.'' (2019). * A study on the phylogenetic relationships of
diploporita Diploporita is an extinct class of blastozoan that ranged from the Ordovician to the Devonian. These echinoderms are identified by a specialized respiratory structure, called diplopores. Diplopores are a double pore system that sit within a de ...
n
blastozoa Blastozoa is a subphylum of extinct animals belonging to Phylum Echinodermata. This subphylum is characterized by the presence of specialized respiratory structures and brachiole plates used for feeding. This subphylum ranged from the Cambrian to ...
ns is published by Sheffield & Sumrall (2019). * A study on the morphology of the feeding
ambulacral Ambulacral is a term typically used in the context of anatomical parts of the phylum Echinodermata or class Asteroidea and Edrioasteroidea. Echinoderms can have ambulacral parts that include ossicles, plates, spines, and suckers. For example, sea ...
system in the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
diploporitan '' Eumorphocystis'', as indicated by data from well-preserved specimens from the Bromide Formation ( Oklahoma,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
), is published by Sheffield & Sumrall (2019), who interpret their findings as indicating that ''Eumorphocystis'' was closely related to
crinoid Crinoids are marine animals that make up the Class (biology), class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or coma ...
s and that crinoids are nested within blastozoans; their conclusions about the relationship between ''Eumorphocystis'' and crinoids are subsequently contested by Guensburg ''et al.'' (2020). * A study on the morphology and phylogenetic relationships of '' Macurdablastus uniplicatus'' is published by Bauer, Waters & Sumrall (2019). * A study on the morphology and phylogenetic relationships of '' Hexedriocystis'' is published online by Zamora & Sumrall (2019), who consider this taxon to be a blastozoan. * A study on the paleoecology of the specimens of the
edrioasteroid Edrioasteroidea is an extinct class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrates (frequently ha ...
'' Neoisorophusella lanei'' preserved in limestone slabs from the Carboniferous (
Chesterian The Mississippian ( , also known as Lower Carboniferous or Early Carboniferous) is a subperiod in the geologic timescale or a subsystem of the geologic record. It is the earlier of two subperiods of the Carboniferous period lasting from rough ...
)
Kinkaid Formation The Pope Mega Group is a geologic unit found in the Illinois Basin of southern Illinois, southwestern Indiana, and western Kentucky.Stamm, N. 2020b''Geologic Unit: Menard, National Geologic Map Database''Willman, H.B., Atherton, Elwood, Buschbach ...
(
Illinois Illinois ( ) is a state in the Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolitan areas include, Peoria and Roc ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) is published by Shroat-Lewis, Greenwood & Sumrall (2019). * A study on the morphology of '' Cupulocrinus'' and on its implications for inferring the origin of the flexible crinoids is published by Peter (2019). * A study on the phylogenetic relationships of diplobathrid crinoids is published by Cole (2019). * A study on the biological and ecological controls on duration of diplobathrid crinoid genera is published online by Cole (2019). * A study on the macro-evolutionary patterns of body-size trends of cyrtocrinid crinoids is published by Brom (2019). * A study on patterns of paleocommunity structure and niche partitioning in crinoids from the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
( Katian) Brechin
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
(
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central C ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) is published by Cole, Wright & Ausich (2019). * A study on the anatomy of the nervous and circulatory systems of the Cretaceous crinoid '' Decameros ricordeanus'' and on the phylogenetic relationships of this species is published online by Saulsbury & Zamora (2019). * A study on the substrate preference in
stem group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor ...
sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) ...
s during the Carboniferous Period will be published by Thompson & Bottjer (2019). * A study on
Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a ...
recovery of sea urchins after the
Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, a ...
is published by Pietsch ''et al.'' (2019). * A fossil
brittle star Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to starfish. They crawl across the sea floor using their flexible arms for locom ...
belonging to the genus ''
Ophiopetra ''Ophiopetra'' is a genus of prehistoric brittle stars. It lived in lagoons in Solnhofen Plattenkalk Formation. It has only one known species, ''Ophiopetra lithographica''. Taxonomic history Both ''Ophiopetra'' and ''O. lithographica'' were firs ...
'', representing the first record of articulated brittle star from the Mesozoic of South America reported so far, is described from the
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Gloucestershire, England See also *Nizhny Nizhny (russian: Ни́жний; masculine), Nizhnyaya (; feminine), or Nizhneye (russian: Ни́� ...
Agua de la Mula Member of the Agrio Formation (
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
) by Fernández ''et al.'' (2019), who transfer the genus ''Ophiopetra'' to the family
Ophionereididae Ophionereididae are a family of brittle stars. Systematics Ophionereididae has been placed (along with Ophiocomidae) to the superfamily Ophiocomidea and infraorder Chilophiurina or suborder Chilophiurina in different classifications. Ophioner ...
within the order
Amphilepidida Amphilepidida is an order of echinoderms belonging to the class Ophiuroidea. Families: * Amphilepididae * Amphilimnidae * Amphiuridae * Hemieuryalidae * Ophiactidae * Ophiolepididae * Ophionereididae Ophionereididae are a family of brittle st ...
.


New taxa


Conodonts


Research

* A study on the feeding habits of conodonts, as indicated by data from calcium stable isotopes, is published by Balter ''et al.'' (2019). * A study on the variation of
conodont element Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
crystal structure throughout their evolutionary history is published online by Medici ''et al.'' (2019). * A study on the evolution of platform-like P1 elements in conodonts, evaluating its possible link to ecology of conodonts, is published by Ginot & Goudemand (2019). * A study on the impact of early
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
environmental changes on evolution and paleoecology of conodonts from the Canadian part of
Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, althoug ...
is published online by Barnes (2019). * A study on the morphology, occurrences and
biostratigraphical Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Bio ...
value of '' Paroistodus horridus'' is published online by Mestre & Heredia (2019). * A revision of the taxonomy and evolutionary relationships of the Late Ordovician genera '' Tasmanognathus'' and '' Yaoxianognathus'' is published by Yang ''et al.'' (2019). * A study on the composition and architecture of the
apparatus Apparatus may refer to: *Technical term for a body of the Soviet and post-Soviet governments (see Apparatchik) *Machine *Equipment * Critical apparatus, the critical and primary source material that accompanies an edition of a text * "Apparatus" (s ...
of '' Erismodus quadridactylus'' is published by Dhanda ''et al.'' (2019). * A study on the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
of the
Lochkovian The Lochkovian is one of three faunal stages in the Early Devonian Epoch. It lasted from 419.2 ± 3.2 million years ago to 410.8 ± 2.8 million years ago. It marked the beginning of the Devonian Period, and was followed by the Pragian Stage. It ...
conodont species '' Ancyrodelloides carlsi'' is published by Corriga & Corradini (2019). * A study on fossils of members of the genus '' Alternognathus'' from the Upper
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
of the Kowala quarry (central
Poland Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 mill ...
), attempting to calibrate the course of their
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
in days and documenting cyclic mortality events, is published by Świś (2019). * The apparatus of '' Vogelgnathus simplicatus'' is reconstructed from discrete elements from a sample of limited diversity from the Carboniferous strata from
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
by Sanz-López, Blanco-Ferrera & Miller (2019). * Neospathodid conodont elements with partly preserved basal body (one of two main parts of conodont elements, besides the crown) are reported from the Lower Triassic of
Oman Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on the southeastern coast of the Arabian Peninsula, and spans the mouth of ...
by Souquet & Goudemand (2019), who interpret their finding as indicating that the absence of basal bodies in post-
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
conodonts was due to a preservational bias only. * Natural assemblages of conodonts, preserving possible impressions of "eyes", are described from the Lower Triassic
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
black
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone, slate, and shale. Most of the particles of which the stone is composed are less than and are too sm ...
s of the North Kitakami Belt ( Japan) by Takahashi, Yamakita & Suzuki (2019). * A study on the composition of the apparatus of '' Nicoraella'', based on data from clusters from the Middle Triassic Luoping Biota (
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...
, China), is published by Huang ''et al.'' (2019). * The architecture of apparatus of ''Nicoraella kockeli'' is reconstructed by Huang ''et al.'' (2019), who also evaluate proposed functional interpretations of the conodont feeding apparatus. * A study on Middle Triassic conodont assemblages from Jenzig section of the Jena Formation and Troistedt section of the
Meissner Formation Meissner, Meißner or Meisner may refer to: Geography Meissner is the name of the following geographic features: * the Meißner (range), an important mountain range in Hesse, Germany * Hoher Meißner, the highest peak of the Meißner range * Mei ...
(
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
) is published by Chen ''et al.'' (2019), who also study the morphology of the apparatuses of '' Neogondolella haslachensis'' and ''Nicoraella germanica'', and review and revise the species ''Neogondolella mombergensis''. * A study evaluating the quantitative morphological variation of P1 conodont elements within and between seven conodont morphospecies from the Pizzo Mondello section (
Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
) and their evolution within 7 million years around the
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followe ...
/
Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...
boundary is published by Guenser ''et al.'' (2019). * A study on the
taphonomy Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientist Ivan Efremo ...
of basal tissue of conodont elements is published online by Suttner & Kido (2019).


New taxa


Fishes


Amphibians


Reptiles


Synapsids


Non-mammalian synapsids


Research

* A study on the morphological diversity and morphological changes of the humeri of
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
and
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
synapsids through time is published by Lungmus & Angielczyk (2019). * A study on the diversity of patterns of skull shape (focusing on the relative lengths of the face and braincase regions of the skull) in non-mammalian synapsids is published by Krone, Kammerer & Angielczyk (2019). * Two pathologically fused tail vertebrae of a varanopid, likely affected by a metabolic bone disease closely resembling
Paget's disease of bone Paget's disease of bone (commonly known as Paget's disease or, historically, osteitis deformans) is a condition involving cellular remodeling and deformity of one or more bones. The affected bones show signs of dysregulated bone remodeling at the ...
, are described from the early
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Pale ...
Richards Spur locality ( Oklahoma,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) by Haridy ''et al.'' (2019). * Description of new skull remains of '' Echinerpeton intermedium'' and a study on the phylogenetic relationships of this species is published online by Mann & Paterson (2019). * Fossil material of a large carnivorous synapsid belonging to the family
Sphenacodontidae Sphenacodontidae (Greek: "wedge point tooth family") is an extinct family of small to large, advanced, carnivorous, Late Pennsylvanian to middle Permian pelycosaurs. The most recent one, ''Dimetrodon angelensis'', is from the late Kungurian ...
is described from the Torre del Porticciolo locality (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
) by Romano ''et al.'' (2019), representing the first carnivorous non-
therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented mor ...
synapsid Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes rep ...
from the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Pale ...
of Italy reported so far, and one of the few known from Europe. * Description of the
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
and
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vi ...
of a small
neural spine The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
from the Early
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Pale ...
Richards Spur locality ( Oklahoma,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) attributable to ''
Dimetrodon ''Dimetrodon'' ( or ,) meaning "two measures of teeth,” is an extinct genus of non- mammalian synapsid that lived during the Cisuralian (Early Permian), around 295–272 million years ago (Mya). It is a member of the family Sphenacodo ...
'' is published by Brink, MacDougall & Reisz (2019), who also report evidence from fossil teeth indicative of presence of a derived species of ''Dimetrodon'' (otherwise typical of later,
Kungurian In the geologic timescale, the Kungurian is an age or stage of the Permian. It is the latest or upper of four subdivisions of the Cisuralian Epoch or Series. The Kungurian lasted between and million years ago (Ma). It was preceded by the Art ...
localities of
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
and Oklahoma) at the Richards Spur locality. * A study on the
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vi ...
of the
skull roof The skull roof, or the roofing bones of the skull, are a set of bones covering the brain, eyes and nostrils in bony fishes and all land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparati ...
of burnetiamorph biarmosuchians is published by Kulik & Sidor (2019). *
Femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...
of a specimen of the titanosuchid species '' Jonkeria parva'' affected by
osteomyelitis Osteomyelitis (OM) is an infection of bone. Symptoms may include pain in a specific bone with overlying redness, fever, and weakness. The long bones of the arms and legs are most commonly involved in children e.g. the femur and humerus, while the ...
is described from the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Pale ...
of Karoo Basin (
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
) by Shelton, Chinsamy & Rothschild (2019). * A study on the adaptations to herbivory in the teeth of members of the family Tapinocephalidae is published by Whitney & Sidor (2019). * An almost complete skeleton of ''
Tapinocaninus ''Tapinocaninus'' is an extinct genus of therapsids in the family Tapinocephalidae, of which it is the most basal member. Only one species is known, ''Tapinocaninus pamelae'' (meaning "Pam's humble canine"). Fossils have been found dating from ...
pamelae'', providing new information on the anatomy of the appendicular skeleton of this species (including the first accurate vertebral count for a
dinocephalia Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinoceph ...
n), is described from the lowermost
Beaufort Group The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underli ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
by Rubidge, Govender & Romano (2019). * Romano & Rubidge (2019) present body mass estimates for a well preserved and complete skeleton of ''Tapinocaninus pamelae'' from the lowermost Beaufort Group of South Africa. * A study on the skull anatomy and phylogenetic relationships of ''
Styracocephalus ''Styracocephalus platyrhynchus'' ('spike head') is an extinct species of tapinocephalian therapsids that lived during the Guadalupian epoch. ''Styracocephalus''s head ornament meant that it could be recognised from a distance. The most stri ...
platyrhynchus'' is published by Fraser-King ''et al.'' (2019). * A study on the evolution of the sacral vertebrae of
dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typica ...
s is published by Griffin & Angielczyk (2019). * A study on the diversity of dicynodonts from the Upper Permian Naobaogou Formation ( China) is published by Liu (2019). * A study on skulls of South American dicynodonts, aiming to determine whether the differences in skull morphology were related to differences in feeding function, is published by Ordonez ''et al.'' (2019). * New fossil material of '' Endothiodon tolani'' is described from the Permian K5 Formation of the Metangula Graben (
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Mala ...
) by Macungo ''et al.'' (2019). * A study on the anatomy of the postcranial skeleton of ''Endothiodon bathystoma'', based on data from a new specimen from the uppermost ''Pristerognathus'' Assemblage Zone of the
Karoo Supergroup The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a pe ...
(
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
), is published online by Maharaj, Chinsamy & Smith (2019). * Small dicynodont skull assigned to the genus '' Digalodon'' is described from the
Lopingian The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic. The Lopingian is often synonymous with the informal term ...
upper Madumabisa Mudstone Formation (
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are ...
) by Angielczyk (2019), expanding known geographic range of this genus. * Digital
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
of ''
Rastodon ''Rastodon'' is an extinct genus of anomodonts. It is the oldest and most basal known genus of bidentalian dicynodonts. Uniquely among dicynodonts, its tusks curve forward. The type and only species is ''R. procurvidens''. Etymology ''Rastod ...
procurvidens'' is reconstructed by de Simão-Oliveira, Kerber & Pinheiro (2019), who evaluate biological implications of the endocast morphology of this species. * Mancuso & Irmis (2019) describe an
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
of a member of the genus ''
Stahleckeria ''Stahleckeria'' is an extinct genus of Middle Triassic (Ladinian) dicynodonts.Chañares Formation The Chañares Formation is a Carnian-age geologic formation of the Ischigualasto-Villa Unión Basin, located in La Rioja Province, Argentina. It is characterized by drab-colored fine-grained volcaniclastic claystones, siltstones, and sandstones ...
(
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
), and evaluate the implications of this finding for the knowledge of the Triassic
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final st ...
n
biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Bio ...
and biogeography. * A study on the body mass of '' Lisowicia bojani'' is published online by Romano & Manucci (2019). * A study on fossils of a putative Cretaceous dicynodont from Australia reported by
Thulborn Thulborn is an Anglo-Saxon surname dating back to the late 12th century AD, derived from the baptismal name for the "son of Thurburn". Notable people with the surname include: * Richard A. Thulborn, British paleontologist * Scott Thulborn (born 198 ...
& Turner (2003) is published online by Knutsen & Oerlemans (2019), who consider these fossils to be of
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
age, and reinterpret it as fossils of a large mammal, probably a
diprotodontid The Diprotodontidae are an extinct family of large herbivorous marsupials, endemic to Australia and New Guinea during the Oligocene through Pleistocene periods from 28.4 million to 40,000 years ago. The family consisted of large quadrupedal t ...
. * A study aiming to determine patterns of morphological and phylogenetic diversity of
therocephalia Therocephalia is an extinct suborder of eutheriodont therapsids (mammals and their close relatives) from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their ...
ns throughout their evolutionary history is published by Grunert, Brocklehurst & Fröbisch (2019). * A study on variation in rates of body size evolution of therocephalians is published by Brocklehurst (2019). * A study on the morphology of the
manus Manus may refer to: * Manus (anatomy), the zoological term for the distal portion of the forelimb of an animal (including the human hand) * ''Manus'' marriage, a type of marriage during Roman times Relating to locations around New Guinea * Man ...
of a new therocephalian specimen referable to the genus ''
Tetracynodon ''Tetracynodon'' is an extinct genus of therocephalian. Fossils of ''Tetracynodon'' have been found in the Karoo Basin of South Africa. Two species are known: the type species ''T. tenuis'' from the Late Permian and the species ''T. darti'' ...
'' from the
Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
, and on the evolution of the manus morphology of therocephalians, is published by Fontanarrosa ''et al.'' (2019). * A study on patterns of nonmammalian
cynodont The cynodonts () ( clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variet ...
species richness and the quality of their fossil record is published by Lukic-Walther ''et al.'' (2019). * A study on the morphology and bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vi ...
of the postcranial skeleton of '' Galesaurus planiceps'' is published by Butler, Abdala & Botha-Brink (2019). * Redescription of the anatomy of the skull of ''Galesaurus planiceps'' is published by Pusch, Kammerer & Fröbisch (2019). * Description of teeth of all known
diademodontid Diademodontidae is an extinct family of Triassic gomphodonts. The best-known genus is ''Diademodon'' from South Africa. '' Titanogomphodon'' from Namibia may also be a member of Diademodontidae. The Chinese genera ''Hazhenia'' and ''Ordosiodon'' ...
and
trirachodontid Trirachodontidae is an extinct, possibly paraphyletic family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quick ...
cynodont taxa is published by Hendrickx, Abdala & Choiniere (2019), who also propose a standardized list of anatomical terms and abbreviations in the study of
gomphodont Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent w ...
teeth, assign ''
Sinognathus ''Sinognathus'' is an extinct genus of gomphodont cynodonts from the Middle Triassic Ermaying Formation of China. Its type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type ...
'' and ''
Beishanodon ''Beishanodon'' is an extinct genus of eucynodonts from Lower Triassic of China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 ...
'' to the family Trirachodontidae, and consider all specimens previously referred to the species '' Cricodon kannemeyeri'' to be younger individuals of ''
Trirachodon ''Trirachodon'' (Greek: "three ridge tooth") is an extinct genus of cynodonts. Fossils have been found in the ''Cynognathus'' Assemblage Zone of the Beaufort Group in South Africa and the Omingonde Formation of Namibia, dating back to the Early ...
berryi''. * A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vi ...
of the traversodontid cynodonts ''
Protuberum ''Protuberum'' is an extinct genus of traversodontid cynodonts known from a single species ''Protuberum cabralense'',The specific name ''cabralense'' is "necessarily emended from the original ''P. cabralensis'' because ''Protuberum'' is a neuter ...
cabralense'' and ''
Exaeretodon ''Exaeretodon'' is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. ''E. argentinus'' is from the Carnian-age (Late Triassic) Cancha de Boch ...
riograndesis'' is published by Veiga, Botha-Brink & Soares (2019). *
Hypsodont Hypsodont is a pattern of dentition with high-crowned teeth and enamel extending past the gum line, providing extra material for wear and tear. Some examples of animals with hypsodont dentition are cows and horses; all animals that feed on grit ...
postcanine teeth of ''
Menadon ''Menadon'' is an extinct genus of traversodontid cynodonts. The type and only species is ''Menadon besairiei''. Fossils of ''Menadon'' were first found in Isalo II (the Makay Formation) of Madagascar, which preserves sediments from the Middl ...
besairiei'' are described by Melo ''et al.'' (2019), who also study patterns of dental growth and replacement in this species. * Digital
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
s of ''
Massetognathus ''Massetognathus'' ( ; Greek for "chewing muscle jaw") is an extinct genus of plant-eating traversodontid cynodonts. They lived during the Triassic Period about 235 million years ago, and are known from the Chañares Formation in Argentina and t ...
ochagaviae'' and ''
Probelesodon ''Chiniquodon'' is an extinct genus of carnivorous cynodonts, which lived during the Late Triassic ( Carnian) in South America (Argentina and Brazil) and Africa (Namibia and Madagascar). ''Chiniquodon'' was closely related to the genus ''Aleodon' ...
kitchingi'' are reconstructed by Hoffmann ''et al.'' (2019). * A skull of a member of the species ''
Massetognathus ''Massetognathus'' ( ; Greek for "chewing muscle jaw") is an extinct genus of plant-eating traversodontid cynodonts. They lived during the Triassic Period about 235 million years ago, and are known from the Chañares Formation in Argentina and t ...
ochagaviae'' is described from the
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followe ...
''Santacruzodon'' Assemblage Zone of the Santa Maria Supersequence ( Rio Grande do Sul,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Schmitt ''et al.'' (2019). * Description of brain endocasts of ''
Siriusgnathus ''Siriusgnathus'' is a traversodontid cynodont from the Carnian channel sandstones and mudstones of the Candelária Formation, belonging to the Santa Maria Supersequence of the Paraná Basin in southeastern Brazil.
niemeyerorum'' and ''
Exaeretodon ''Exaeretodon'' is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. ''E. argentinus'' is from the Carnian-age (Late Triassic) Cancha de Boch ...
riograndensis'', using virtual models based on computed tomography scan data, is published by Pavanatto, Kerber & Dias-da-Silva (2019). * Description of new fossil material of ''
Siriusgnathus ''Siriusgnathus'' is a traversodontid cynodont from the Carnian channel sandstones and mudstones of the Candelária Formation, belonging to the Santa Maria Supersequence of the Paraná Basin in southeastern Brazil.
niemeyerorum'' from the Upper Triassic
Caturrita Formation The Caturrita Formation is a rock formation found in Rio Grande do Sul, Brazil. Its sediments were deposited in the Paraná Basin. The formation is from the Upper Triassic and forms part of the Santa Maria Supersequence in the upper section of the ...
(
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) and a study on the age of its fossils is published online by Miron ''et al.'' (2019). * A study on the evolution of infraorbital
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
ry canal in probainognathian cynodonts and on its implications for the knowledge of evolution of mobile whiskers in non-mammalian synapsids, as indicated by data from skulls of non-mammalian probainognathian cynodonts and early
mammaliaforms Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent comm ...
, is published online by Benoit ''et al.'' (2019). * Digital skull
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
of a specimen of '' Riograndia guaibensis'' is reconstructed by Rodrigues ''et al.'' (2019). * Description of the anatomy of the first postcranial specimens referable to ''Riograndia guaibensis'' is published by Guignard, Martinelli & Soares (2019). * A study on the anatomy of the postcranial skeleton of '' Brasilodon quadrangularis'' is published by Guignard, Martinelli & Soares (2019). * A study on tooth wear patterns of members of the family
Tritylodontidae Tritylodontidae ("three-knob teeth", named after the shape of their cheek teeth) is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, bearing several mammalian traits like erect limbs, endothermy and details o ...
and on their possible diet is published by Kalthoff ''et al.'' (2019). * Possible cynodont teeth, which might be the most recent non-
mammaliaform Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent ...
cynodont fossils from Africa reported so far, are described from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the ...
or earliest
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
locality of Ksar Metlili (Anoual Syncline, eastern
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
) by Lasseron (2019). * A study on the origin of the mammalian
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in ...
ossicles, as indicated by the anatomy of the jaw-otic complex in 43 synapsid taxa, is published by Navarro-Díaz, Esteve-Altava & Rasskin-Gutman (2019). * A study on the evolution of the morphological complexity of the mammalian vertebral column, as indicated by data from mammals and non-mammalian synapsids, is published by Jones, Angielczyk & Pierce (2019).


New taxa


Mammals


Other animals


New taxa


Research

* A study on moulds of animals belonging to the group
Proarticulata Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. pr ...
from the southeastern
White Sea The White Sea (russian: Белое море, ''Béloye móre''; Karelian and fi, Vienanmeri, lit. Dvina Sea; yrk, Сэрако ямʼ, ''Serako yam'') is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is s ...
area (
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
), and on their implications for the knowledge of the morphology of integuments of members of Proarticulata, is published by Ivantsov, Zakrevskaya & Nagovitsyn (2019). * A study on accumulations of ''
Ernietta ''Ernietta'' is an extinct genus of Ediacaran organisms with an infaunal lifestyle. Fossil preservations and modeling indicate this organism was sessile and “sack”-shaped. It survived partly buried in substrate, with an upturned bell-shaped f ...
'' from the Witputs subbasin (
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and e ...
), and on their implications for the knowledge of ecology of these organisms, is published by Gibson ''et al.'' (2019). * A diverse assemblage of tubular fossils – dominated by typical
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and t ...
organisms such as '' Cloudina'' and '' Sinotubulites'', but also preserving fossils showing similarities to early
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
shelly fossils – is described from the Ediacaran Dengying Formation ( China) by Cai ''et al.'' (2019). * Letsch ''et al.'' (2019) report late Ediacaran discoidal Ediacara-type fossils and latest Ediacaran to early
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
microfossils A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, ...
from the Tabia and the Tifnout members of the Adoudou Formation (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
), constituting the oldest known direct evidence for presumably animal life from Northwest Africa. * A study delineating different types of the asexual reproduction for '' Cloudina'' and '' Multiconotubus'' is published by Min ''et al.'' (2019). * A study on the anatomy of ''
Charnia ''Charnia'' is a genus of frond-like lifeforms belonging to the Ediacaran biota with segmented, leaf-like ridges branching alternately to the right and left from a zig-zag medial suture (thus exhibiting glide reflection, or opposite isometry). T ...
masoni'' is published by Dunn ''et al.'' (2019). * A study evaluating whether ''
Dickinsonia ''Dickinsonia'' is an extinct genus of basal animal that lived during the late Ediacaran period in what is now Australia, China, Russia and Ukraine. The individual ''Dickinsonia'' typically resembles a bilaterally symmetrical ribbed oval. Its a ...
'' was capable of mobility is published by Evans, Gehling & Droser (2019). * A study comparing the biomechanical responses of tissues of ''Dickinsonia'' to various forces with those typical of modern organisms is published by Evans ''et al.'' (2019). * A study on the anatomy, growth and phylogenetic relationships of '' Arborea arborea'' is published by Dunn, Liu & Gehling (2019). * Xiao ''et al.'' (2019) describe a new
trace fossil A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...
from the Ediacaran Dengying Formation (China), interpreted as produced by a
bilateria The Bilateria or bilaterians are animals with bilateral symmetry as an embryo, i.e. having a left and a right side that are mirror images of each other. This also means they have a head and a tail (anterior-posterior axis) as well as a belly an ...
n animal exploring an oxygen oasis in
microbial mat A microbial mat is a multi-layered sheet of microorganisms, mainly bacteria and archaea, or bacteria alone. Microbial mats grow at interfaces between different types of material, mostly on submerged or moist surfaces, but a few survive in desert ...
s, and name a new
ichnotaxon An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...
'' Yichnus levis''. * A study on fossil molds and casts from the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
of
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
and the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
of
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...
, as well as on Ediacaran mold and cast fossils from
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
, the
White Sea The White Sea (russian: Белое море, ''Béloye móre''; Karelian and fi, Vienanmeri, lit. Dvina Sea; yrk, Сэрако ямʼ, ''Serako yam'') is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is s ...
region of
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
,
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and e ...
and
Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
, is published by MacGabhann ''et al.'' (2019), who evaluate how faithfully the fossils represent the original organisms, and whether the first animals to evolve on Earth could have been fossilized in a way similar to eldoniids from the Tafilalt
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
of Morocco. * Exceptionally preserved phosphatized archaeocyaths and
small shelly fossils The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambr ...
are reported from the Lower Cambrian Salaagol Formation of southwestern
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 millio ...
by Pruss ''et al.'' (2019). * A study on the timing of the development of
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic processes—deposition (geology), deposition of ...
biodiversity, based on data from microbial-archaeocyathan reefs of the Salaagol Formation in Mongolia and other early
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
reefs, is published by Cordie ''et al.'' (2019). * A study on the morphological diversity of archaeocyaths is published by Cordie & Dornbos (2019). * Evidence of extensive burrowing in laminated claystone from the
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
( Drumian) Ravens Throat River
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
in the
Rockslide Formation The Rockslide Formation is a geologic formation in Northwest Territories. It preserves fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples ...
(
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) is presented by Pratt & Kimmig (2019). * Description of jaw apparatus of ''
Plumulites ''Plumulites'' is an extinct genus of machaeridians, extinct annelid group. Description ''Plumulites'' is normally small animal, for example complete specimen of ''P. canadensis'' reached only , and ''P. bengtsoni'' being . However, several ...
bengtsoni'' from the
Fezouata Formation The Fezouata Formation or Fezouata Shale is a geological formation in Morocco which dates to the Early Ordovician.
of
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
, evaluating its implications for the knowledge of the phylogenetic relationships of machaeridians, is published by Parry ''et al.'' (2019). * Description of internal anatomical features of '' Canadia spinosa'' identified as remnants of the nervous system is published by Parry &
Caron A caron (), háček or haček (, or ; plural ''háčeks'' or ''háčky'') also known as a hachek, wedge, check, kvačica, strešica, mäkčeň, varnelė, inverted circumflex, inverted hat, flying bird, inverted chevron, is a diacritic mark (� ...
(2019). * A study on the chemical composition,
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
and phylogeny of fossil (
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...
,
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ...
and
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
)
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
tubes and tubes formerly thought to have been made by annelids, recovered from hydrothermal vent and
cold seep A cold seep (sometimes called a cold vent) is an area of the ocean floor where hydrogen sulfide, methane and other hydrocarbon-rich fluid seepage occurs, often in the form of a brine pool. ''Cold'' does not mean that the temperature of the see ...
environments, is published by Georgieva ''et al.'' (2019). * A massive deposit composed of fossil
serpulid The Serpulidae are a family of sessile, tube-building annelid worms in the class Polychaeta. The members of this family differ from other sabellid tube worms in that they have a specialized operculum that blocks the entrance of their tubes ...
worm tubes dating to the late
Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
is reported from the Santa Monica Basin off the coast of southern
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
by Georgieva ''et al.'' (2019). * A study on the microstructure of
hyolith Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as be ...
conchs and opercula from the lower
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
Xinji Formation Xinji () is a county-level city of Hebei Province, North China, it is under the administration of the prefecture-level city of Shijiazhuang. It is the easternmost county-level division of Shijiangzhuang. There are 8 towns () and 7 townships ...
of North China, and on its implications for inferring the phylogenetic relationships of Hyolitha, is published by Li ''et al.'' (2019). * Description of soft parts associated with the feeding apparatus of the hyolith '' Triplicatella opimus'' from the
Chengjiang biota The Maotianshan Shales are a series of Early Cambrian deposits in the Chiungchussu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales ...
of South China, and a study on the implications of this finding for the knowledge of the phylogenetic affinities of hyoliths, is published online by Liu ''et al.'' (2019). * A study on changes of conch size in tentaculitoids from the
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoi ...
and
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
strata is published by Wei (2019). * A study on the anatomy of '' Amiskwia sagittiformis'' is published by Vinther & Parry (2019), who interpret two reflective patches present in fossils of this species, previously interpreted as paired cerebral ganglia, as a pair of pharyngeal jaws similar to those of gnathiferans. * A study on the anatomy and phylogenetic affinities of ''Amiskwia sagittiformis'' is published by
Caron A caron (), háček or haček (, or ; plural ''háčeks'' or ''háčky'') also known as a hachek, wedge, check, kvačica, strešica, mäkčeň, varnelė, inverted circumflex, inverted hat, flying bird, inverted chevron, is a diacritic mark (� ...
& Cheung (2019). * The oldest record of
acanthocephala Acanthocephala (Greek , ', thorn + , ', head) is a phylum of parasitic worms known as acanthocephalans, thorny-headed worms, or spiny-headed worms, characterized by the presence of an eversible proboscis, armed with spines, which it uses to p ...
n parasite eggs described so far is reported from probable crocodyliform
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
s from the Upper Cretaceous
Adamantina Formation The Adamantina Formation is a geological formation in the Bauru Basin of western São Paulo state, in southeastern Brazil. Its strata date back to the Late Cretaceous epoch of the Cretaceous Period, during the Mesozoic Era. The formation is par ...
(
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Cardia ''et al.'' (2019). * Exceptionally preserved trace and body fossils are described from the
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
File Haidar Formation File or filing may refer to: Mechanical tools and processes * File (tool), a tool used to ''remove'' fine amounts of material from a workpiece ** Filing (metalworking), a material removal process in manufacturing ** Nail file, a tool used to g ...
( Sweden) by Kesidis ''et al.'' (2019), who interpret these fossils as made by
priapulid Priapulida (priapulid worms, from Gr. πριάπος, ''priāpos'' 'Priapus' + Lat. ''-ul-'', diminutive), sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility ...
-like
scalidophora Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous c ...
ns. * Description of
exuviae In biology, exuviae are the remains of an exoskeleton and related structures that are left after ecdysozoans (including insects, crustaceans and arachnids) have moulted. The exuviae of an animal can be important to biologists as they can often be ...
of microscopic
scalidophora Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous c ...
n worms from the lowermost Cambrian Kuanchuanpu Formation ( China) is published by Wang ''et al.'' (2019), who interpret this finding as the oldest record of
moulting In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
in
ecdysozoa Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerata, crustaceans, and myriapods), Nematoda, and several smaller phyla. They were first defined by Aguinaldo ''et al.'' in 1997, based mainly on phylogeneti ...
ns reported so far. * A reassessment of
radiodonta Radiodonta is an extinct order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two original ...
n fossils known from the
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
Kinzers Formation The Kinzers Formation is a geologic formation in Pennsylvania. It preserves fossils dating back to the Cambrian Period. The base of the Kinzers Formation is primarily a dark-brown shale. The middle is a gray and white spotted limestone and, loca ...
(
Pennsylvania Pennsylvania (; (Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, Ma ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) is published by Pates & Daley (2019), who argue that at least four radiodontan taxa are known from this formation, and confirm that ''
Anomalocaris ''Anomalocaris'' ("unlike other shrimp", or "abnormal shrimp") is an extinct genus of radiodont, an order of early-diverging stem-group arthropods. The first fossils of ''Anomalocaris'' were discovered in the ''Ogygopsis'' Shale of the Stephen Fo ...
pennsylvanica'' is a distinct species from ''A. canadensis''. * A study on the
moulting In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
behaviour of the chengjiangocaridid fuxianhuiid '' Alacaris mirabilis'' is published by Yang ''et al.'' (2019). * A study on the anatomy and phylogenetic relationships of a
stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushr ...
-arthropod '' Guangweicaris spinatus'' is published by Wu & Liu (2019). * A fossil interpreted as a partial mold of a specimen of '' Paropsonema cryptophya'' is described from the Middle-Upper Devonian of New York by Hagadorn & Allmon (2019), representing the most recent occurrence of the
paropsonemid ''Paropsonema'' is a discoidal animal known from the Cambrian Chengjiang biota and classified with the eldoniid Eldoniids are an extinct clade of enigmatic disc-shaped animals which lived in the early to middle Paleozoic (Cambrian to Devonian). ...
s reported so far. * A study evaluating the utility of eye
melanosome A melanosome is an organelle found in animal cells and is the site for synthesis, storage and transport of melanin, the most common light-absorbing pigment found in the animal kingdom. Melanosomes are responsible for color and photoprotection i ...
s for determination of the phylogenetic affinities of '' Tullimonstrum'' is published by Rogers ''et al.'' (2019).


Foraminifera


Research

* A study on the morphological complexity of
planktic Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a crucia ...
foraminifer
tests Test(s), testing, or TEST may refer to: * Test (assessment), an educational assessment intended to measure the respondents' knowledge or other abilities Arts and entertainment * ''Test'' (2013 film), an American film * ''Test'' (2014 film), ...
after the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden extinction event, mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million y ...
is published by Lowery & Fraass (2019). * A study on the response of the larger
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
from the
Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
to the
Paleocene–Eocene Thermal Maximum The Paleocene–Eocene thermal maximum (PETM), alternatively (ETM1), and formerly known as the "Initial Eocene" or "", was a time period with a more than 5–8 °C global average temperature rise across the event. This climate event o ...
, based on fossil evidence from south
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
, is published by Zhang ''et al.'' (2019).


New taxa


Other organisms


New taxa


Research

* Putative traces of life older than 3.95 Ga, reported from northern
Labrador , nickname = "The Big Land" , etymology = , subdivision_type = Country , subdivision_name = Canada , subdivision_type1 = Province , subdivision_name1 ...
(
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) by Tashiro ''et al.'' (2017) are reevaluated by Whitehouse ''et al.'' (2019). * Description of cellularly preserved microfossils from ~3.4 Ga-old deposits of the Kromberg Formation (
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
), providing information on reproduction patterns of these organisms, is published by Kaźmierczak & Kremer (2019). * El Albani ''et al.'' (2019) describe 2.1 billion-year-old fossils belonging to the
Francevillian biota The Francevillian biota (also known as Gabon macrofossils or Gabonionta) is a group of 2.1-billion-year-old Palaeoproterozoic, macroscopic organisms known from fossils found in Gabon in the Palaeoproterozoic Francevillian B Formation, a black sh ...
of
Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...
, including pyritized string-shaped structures interpreted as produced by a
multicellular A multicellular organism is an organism that consists of more than one cell, in contrast to unicellular organism. All species of animals, land plants and most fungi are multicellular, as are many algae, whereas a few organisms are partially un ...
or
syncytial A syncytium (; plural syncytia; from Ancient Greek, Greek: σύν ''syn'' "together" and κύτος ''kytos'' "box, i.e. cell") or symplasm is a multinucleate cell (biology), cell which can result from multiple cell fusions of uninuclear cells (i.e ...
organism able to migrate laterally and vertically to reach food resources. * A study on ca. 1.9 Ga hairpin-shaped trace fossils and discoid fossils from the Stirling Range Formation (
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to ...
) is published by
Retallack Retallack village is near St Columb Major, in Cornwall, England, UK, and in the civil parish of St Wenn. Most of the village was turned into a 100 acre commercial theme park called "Spirit of the West". The theme park was closed in 2009 and r ...
& Mao (2019), who interpret these fossils as evidence of early life on land. * A study on organic-walled microfossils from the Cailleach Head Formation ( Torridon Group,
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to th ...
) is published by Wacey ''et al.'' (2019), who report exceptional preservation of sub-cellular detail in selected cells. * Phosphatized three-dimensional fossil of a putative calcimicrobe ''Epiphyton'' are reported from the
Neoproterozoic The Neoproterozoic Era is the unit of geologic time from 1 billion to 538.8 million years ago. It is the last era of the Precambrian Supereon and the Proterozoic Eon; it is subdivided into the Tonian, Cryogenian, and Ediacaran periods. It is ...
Dengying Formation ( China) by Min ''et al.'' (2019). * A study on the affinities of tubular microfossils from the
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and t ...
Doushantuo Formation The Doushantuo Formation (formerly transcribed as Toushantuo or Toushantou, from ) is a geological formation in western Hubei, eastern Guizhou, southern Shaanxi, central Jiangxi, and other localities in China. It is known for the fossil Lagers ...
( China), i.e. '' Crassitubus'', '' Quadratitubus'', '' Ramitubus'' and '' Sinocyclocyclicus'', is published by Sun ''et al.'' (2019), who reject the interpretation of these taxa as early animals. * Lehn, Horodyski & Paim (2019) report the first known occurrence of Ediacaran organic-walled microfossils preserved in fine-grained
siliciclastic Siliciclastic (or ''siliclastic'') rocks are clastic noncarbonate sedimentary rocks that are composed primarily of silicate minerals, such as quartz or clay minerals. Siliciclasic rock types include mudrock, sandstone, and conglomerate Conglome ...
strata of the Camaquã Basin (southernmost
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
). * A study on the structure, developmental biology and affinities of '' Caveasphaera costata'' from the Ediacaran
Doushantuo Formation The Doushantuo Formation (formerly transcribed as Toushantuo or Toushantou, from ) is a geological formation in western Hubei, eastern Guizhou, southern Shaanxi, central Jiangxi, and other localities in China. It is known for the fossil Lagers ...
( China) is published by Yin ''et al.'' (2019). * A study on possible cells and their appendages in fossils of ''Epiphyton'' from the
Wuliuan The Wuliuan stage is the fifth stage of the Cambrian, and the first stage of the Miaolingian Series of the Cambrian. It was formally defined by the ICS in 2018. Its base is defined by the first appearance of the trilobite species '' Oryctocepha ...
of the North China Platform, and on their implications for the classification of this taxon, is published by Zhang ''et al.'' (2019). * A study on the
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
and colony organization of '' Rhyniococcus uniformis'' (a
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
organism resembling extant cyanobacteria in the genus '' Merismopedia''), based on data from new specimens, is published by Krings & Harper (2019). * A new method of assessing the morphology of fossil
radiolaria The Radiolaria, also called Radiozoa, are protozoa of diameter 0.1–0.2 mm that produce intricate mineral skeletons, typically with a central capsule dividing the cell into the inner and outer portions of endoplasm and ectoplasm. The elab ...
n specimens is presented by Kachovich, Sheng & Aitchison (2019), who apply their method to six specimens from the Cambrian
Inca Formation The Inca Empire (also known as the Incan Empire and the Inka Empire), called ''Tawantinsuyu'' by its subjects, ( Quechua for the "Realm of the Four Parts",  "four parts together" ) was the largest empire in pre-Columbian America. The adm ...
( Australia) and Ordovician
Piccadilly Formation Piccadilly () is a road in the City of Westminster, London, to the south of Mayfair, between Hyde Park Corner in the west and Piccadilly Circus in the east. It is part of the A4 road that connects central London to Hammersmith, Earl's Cou ...
(
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) and evaluate the implications of their method for the studies of radiolarian evolution.


Trace fossils


History of life in general

Research related to paleontology that concerns multiple groups of the organisms listed above. * Experiments indicating that abiotic
chemical garden Comparison of chemical gardens grown by NASA scientists on the International Space Station (left) and on the ground (right) A chemical garden while growing up Cobalt(II) chloride upA chemical garden A chemical garden is a set of complex biol ...
ing can mimic structures interpreted as the oldest known fossil microorganisms in both morphology and composition are conducted by McMahon (2019). * A study on
biomarker In biomedical contexts, a biomarker, or biological marker, is a measurable indicator of some biological state or condition. Biomarkers are often measured and evaluated using blood, urine, or soft tissues to examine normal biological processes, p ...
s recovered from cap dolomites of the Araras Group (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
), interpreted as evidence of the transition from a bacterial to
eukaryotic Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bact ...
dominated ecosystem after the Marinoan deglaciation, likely caused by massive Bacterivore, bacterivorous grazing by ciliates, is published by van Maldegem ''et al.'' (2019). * Biomarkers thought to be diagnostic for demosponges and cited as evidence of rise of animals to ecological importance prior to the Cambrian radiation are reported to be also synthesized by rhizarians by Nettersheim ''et al.'' (2019), who place the oldest unambiguous evidence for animals closer to the Cambrian Explosion. * A study on crucial conditions affecting the evolution of a proto-metabolism in early life is published by Goldford ''et al.'' (2019). * A study on the age of the
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and t ...
fossils from the Podolya Basin (southwestern Ukraine) is published by Soldatenko ''et al.'' (2019). * A study on occurrences of body and trace fossils in
Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and t ...
and lower
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
(Fortunian) rocks around the world is published by Muscente ''et al.'' (2019), who report evidence indicative of existence of a global, Cosmopolitan distribution, cosmopolitan assemblage unique to terminal Ediacaran strata, living between two episodes of biotic turnover which might be the earliest mass extinctions of complex life. * A study on the diversification of animals and their behaviour in the Ediacaran–Cambrian interval, as indicated by fossil and environmental proxy records, is published by Rachel Wood (geologist), Wood ''et al.'' (2019), who interpret the fossil record as indicating that the rise of early animals was more likely a series of successive, transitional Evolutionary radiation, radiation events which extended from the Ediacaran to the early
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
, rather than competitive or biotic replacement of the latest Ediacaran biotas by markedly distinct Cambrian ones. * A study comparing the variability of Ediacaran faunal assemblages to that of more recent fossil and modern
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
assemblages is published by Finnegan, Gehling & Droser (2019). * A study on the intensity of animal bioturbation and ecosystem engineering in trace fossil assemblages throughout the latest Ediacaran Nama Group (
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and e ...
), evaluating the implications of this data for the knowledge of the causes of the disappearance of the Ediacaran biota, is published by Cribb ''et al.'' (2019). * A study on mechanisms of skeletal biomineralization in early animals (focusing on '' Cloudina'' and
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
Hyolitha, hyoliths and halkieriids) is published by Pupa Gilbert, Gilbert ''et al.'' (2019). * A study on the relationship between atmospheric oxygen oscillations, the extent of shallow-ocean oxygenation and the animal biodiversity in the Cambrian period is published by He ''et al.'' (2019). * A study on the course of the transition from microbial-dominated reef environments to animal-based reefs in the early Cambrian, as indicated by data from strata in the western Basin and Range Province, Basin and Range of
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
and Nevada, is published by Cordie, Dornbos & Marenco (2019). * An assemblage of early Cambrian small carbonaceous fossils and acritarchs, including possible oldest known
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
remains, is described from the siltstones of the Lake Lappajärvi, Lappajärvi impact structure (Finland) by Slater & Willman (2019). * A study aiming to explain the occurrence of the variety of trace fossils associated with ''Tuzoia'' carapaces from the Cambrian Burgess Shale (British Columbia,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) is published by Mángano, Hawkes &
Caron A caron (), háček or haček (, or ; plural ''háčeks'' or ''háčky'') also known as a hachek, wedge, check, kvačica, strešica, mäkčeň, varnelė, inverted circumflex, inverted hat, flying bird, inverted chevron, is a diacritic mark (� ...
(2019). * Cambrian
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
from the Qingjiang biota (Shuijingtou Formation; Hubei, China), preserving fossils of diverse, ~518 million years old biota, is reported by Fu ''et al.'' (2019). * A study aiming to infer whether a marked drop in known diversity of marine life during the period between the Cambrian explosion and the
Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced w ...
(the Furongian Gap) is apparent, due to sampling failure or lack of rock, or real, is published by David Harper (palaeontologist), Harper ''et al.'' (2019). * A study on the marine biodiversity changes throughout the first 120 million years of the Phanerozoic is published by Rasmussen ''et al.'' (2019). * A study aiming to determine factors influencing early Palaeozoic marine biodiversity is published by Penny & Kröger (2019). * A study on rates of origination and extinction at the genus level throughout early
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...
is published by Kröger, Franeck & Rasmussen (2019), who also present estimates of longevity, taxon age and taxon life expectancy of early Paleozoic marine genera. * A review of biodiversity curves of marine organisms throughout early Paleozoic, indicating the occurrence of a large-scale, long-term radiation of life that started during late Precambrian time and was only finally interrupted in the Devonian Period, is published online by David Harper (palaeontologist), Harper, Cascales-Miñana & Servais (2019). * A study on processes causing fluctuations of biodiversity of marine invertebrates throughout the Phanerozoic is published by Rominger, Fuentes & Marquet (2019). * A study on the impact of environmental changes on the biodiversity of North American marine organisms throughout the Phanerozoic is published by Roberts & Mannion (2019). * A study testing the hypothesis that the influence of ocean chemistry and climate on the ecological success of marine calcifiers decreased throughout the Phanerozoic is published by Eichenseer ''et al.'' (2019). * A study on genus origination and extinction rates in the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
on a global scale, for the paleocontinents Baltica and
Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, althoug ...
, and for onshore and offshore areas, is published by Franeck & Liow (2019). * First Middle Ordovician (Dapingian–Darriwilian) soft-bodied fossils from northern
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final st ...
(fossils of
medusozoa Medusozoa is a clade in the phylum Cnidaria, and is often considered a subphylum. It includes the classes Hydrozoa, Scyphozoa, Staurozoa and Cubozoa, and possibly the parasitic Polypodiozoa. Medusozoans are distinguished by having a medusa stag ...
n possibly belonging to the genus ''Patanacta'', possible members of the family Wiwaxiidae and an arthropod possibly belonging to the family Pseudoarctolepidae) are described from the Valongo Formation (Portugal) by Kimmig ''et al.'' (2019). * New Konservat-
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
containing exceptionally preserved soft-bodied organisms, including the earliest record of Acoelomorpha, Turbellaria, Nemertea and Nematode, Nematoda reported so far, is described from the Ordovician ( Katian) Vaureal Formation, Vauréal Formation (
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) by Knaust & Desrochers (2019). * A review of occurrence data of latest Ordovician
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
marine organisms is published by Wang, Zhan & Percival (2019), who evaluate the implications of the studied data for the knowledge of the course of the Ordovician–Silurian extinction events, end-Ordovician mass extinction. * A revision of
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoi ...
fauna from the Pentland Hills (
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) described by Archibald Lamont in 1978 is published by Candela & Crighton (2019). * A study on the course of Graptolithina, graptolite extinctions during the middle Homerian biotic crisis and on the impact of this crisis on other marine invertebrates, as indicated by data from the Kosov Quarry section of the Prague Synform (Czech Republic), is published by Manda ''et al.'' (2019). * Well-preserved fossil cryptic biota is reported from the submarine cavities of the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
(Emsian to Givetian) mud mounds in the Hamar Laghdad area (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
) by Berkowski ''et al.'' (2019). * A study aiming to test and quantify the classification of Devonian Biogeography, biogeographic areas, based on distributional data of Devonian trilobite, brachiopod and fish taxa, is published by Dowding & Ebach (2019). * A study on patterns of local richness of terrestrial tetrapods throughout the Phanerozoic is published by Close ''et al.'' (2019). * Description of tetrapod and fish fossils from the coastal locality of Burnmouth, Scotland (Ballagan Formation), associated plant material and Sedimentology, sedimentological context of these fossils is published by Jenny Clack, Clack ''et al.'' (2019), who interpret these fossils as evidence of the potential richness of the Tournaisian fauna, running counter to the assumption of a depauperate nonmarine fauna following the end-
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
Hangenberg event. * A study on the impact of climate changes during the Carboniferous–Permian transition on the evolution of land-living vertebrates is published by Pardo ''et al.'' (2019). * A study aiming to test one of the scenarios proposed by Robert L. Carroll in 1970 to explain the origin of the Amniote, amniotic egg, based on data from Permo-Carboniferous tetrapods, is published by Didier, Chabrol & Michel Laurin, Laurin (2019). * An overview of the studies researching biodiversity changes in the Permian and their links to volcanism is published by Chen & Xu (2019). * Haridy ''et al.'' (2019) report the occurrence of overgrowth of Palate, palatal dentition of ''Cacops'' and ''Captorhinus'' by a new layer of bone to which the newest teeth are then attached (the overgrowth pattern also documented in early fishes), and evaluate the implications of this finding for the knowledge of the origin of teeth. * A study on the severity of the end-Guadalupian extinction event is published online by Rampino & Shen (2019). * A study on the ecology of Permian tetrapods from the Abrahamskraal Formation (
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
), as indicated by stable oxygen isotope compositions of phosphate from teeth and bones used as a proxy for water dependence, is published online by Rey ''et al.'' (2019). * Two Permian tetrapod assemblages, recovered from the northernmost point at which the lowest
Beaufort Group The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underli ...
has been targeted for collecting fossils, are reported from the southern Free State (province), Free State (South Africa) by Groenewald, Day & Rubidge (2019), who evaluate the implications of these fossils for the knowledge of faunal provincialism within the Middle to Late Permian Karoo Basin. * A study aiming to determine which Permian tetrapod assemblage zones are present in the vicinity of Victoria West (Northern Cape,
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
), and to reassess the Biostratigraphy, biostratigraphic provenance of specimens collected historically in this area (including the holotype of ''Lycaenops ornatus''), is published by Day & Rubidge (2019). * A study on the course of the turnover from the ''Daptocephalus Assemblage Zone, Daptocephalus'' to Lystrosaurus Assemblage Zone, ''Lystrosaurus'' Assemblage Zones of the Karoo Basin is published by Gastaldo ''et al.'' (2019). * A study on the timing of the extinction of latest Permian vertebrates in the Karoo Basin of South Africa is published online by Michael R. Rampino, Rampino ''et al.'' (2019). * A study on the identification and position of the terrestrial end-Permian mass extinction in southern African sediments, based on data from a new site in the South African Karoo Basin, is published online by Botha ''et al.'' (2019). * A study on the functional diversity of middle Permian and Early Triassic marine paleocommunities in the area of present-day western
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
, and on its implications for the knowledge of functional re-organization of these communities in the aftermath of the Permian–Triassic extinction event, is published by Dineen, Roopnarine & Fraiser (2019). * A study aiming to explain high biodiversity preserved in the Triassic Cassian Formation (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
) is published online by Roden ''et al.'' (2019). * A study on shark, sizable carnivorous archosaur, big herbivorous tetrapod and probable turtle bromalites (
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
s and possibly some cololites) from a turtle-dominated fossil assemblage from the Upper Triassic Poręba site (
Poland Poland, officially the Republic of Poland, , is a country in Central Europe. Poland is divided into Voivodeships of Poland, sixteen voivodeships and is the fifth most populous member state of the European Union (EU), with over 38 mill ...
) is published by Bajdek ''et al.'' (2019), who evaluate the implications of their findings for inferring the diet of the Triassic turtle ''Proterochersis porebensis''. * A study on seawater oxygenation during the Early Jurassic and its impact on the recovery of marine benthos after the Triassic–Jurassic extinction event, as indicated by data from Blue Lias Formation (United Kingdom), is published by Atkinson & Wignall (2019). * A study on the patterns and processes of recovery of marine fauna after the Toarcian oceanic anoxic event, as indicated by data from the Cleveland Basin (Yorkshire, United Kingdom), is published by Caswell & Dawn (2019). * A study on changes of land vegetation resulting from the Toarcian oceanic anoxic event is published by Slater ''et al.'' (2019). * Skeletal elements of Oxfordian (stage), Oxfordian ichthyosaurs and plesiosaurs are reported from the Kingofjeld mountain (north-east Greenland) by Delsett & Alsen (2019). * New marine reptile-bearing localities documenting the Tithonian–Berriasian transition at the High Andes (Mendoza Province,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
) are reported by Fernández ''et al.'' (2019). * A study on microvertebrate fossils from the Upper Jurassic or Lower Cretaceous of Ksar Metlili (Anoual Syncline,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
), evaluating their palaeobiogeographical implications, and on the age of this fauna, is published online by Lasseron ''et al.'' (2019). * Description of mid-Cretaceous invertebrate fauna from Batavia Knoll (eastern Indian Ocean), and a study on its similarities to other Cretaceous faunas from around the Indian Ocean, is published by Wild & Stilwell (2019). * A study on the age of the vertebrate fauna from the Cretaceous Cerro Barcino Formation (
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
) is published online by Krause ''et al.'' (2019). * Possible amphibian, gastropod and insect egg masses are described from the Cretaceous amber from
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
by Xing ''et al.'' (2019). * A study on
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...
s from the Upper Cretaceous deposits in the Münster Basin (northwestern
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
), evaluating their implications for the knowledge of Cretaceous Trophic level, trophic structures and predator–prey interactions, is published by Qvarnström ''et al.'' (2019). * New vertebrate assemblage from the upper Turonian Schönleiten Formation of Gams bei Hieflau (Austria) is described by Ősi ''et al.'' (2019). * Turonian marine vertebrate fossils from the Huehuetla quarry (Puebla, Mexico) are described by Alvarado-Ortega ''et al.'' (2019). * A study on the biogeography of Cretaceous terrestrial tetrapods is published by Kubo (2019). * A study on the structure and contents of a large piece of amber attached to a jaw of a specimen of ''Prosaurolophus maximus'' from the Cretaceous Dinosaur Park Formation (Alberta,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
), evaluating the implications of this finding for the knowledge of the habitat and taphonomy of the dinosaur, is published by McKellar ''et al.'' (2019). * An accumulation of fossil eggshells of bird, Crocodylomorpha, crocodylomorph and gekkotan eggs is reported from the Late Cretaceous Oarda de Jos locality in the vicinity of the city of Sebeș (Romania) by Fernández ''et al.'' (2019). * A review of the fossil record of Late Cretaceous and Paleogene vertebrates from the Seymour Island (Antarctica) is published by Reguero (2019). * A study on the evolutionary history of the family Pospiviroidae, aiming to assess possible impact of the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden extinction event, mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million y ...
on the divergence rates in this family, is published by Bajdek (2019). * A study on calcareous nanoplankton and planktic foraminiferal assemblages in a Cretaceous-Paleogene section from the peak ring of the Chicxulub crater, and on their implications for the knowledge of recovery of plankton after the Cretaceous–Paleogene extinction event, is published by Jones, Lowery & Bralower (2019). * A study on the course of recovery of the nanoplankton communities after the Cretaceous–Paleogene extinction event is published by Alvarez ''et al.'' (2019), who report evidence indicative of 1.8 million years of exceptional volatility of post-extinction communities and indicating that the emergence of a more stable equilibrium-state community coincided with indicators of carbon cycle restoration and a fully functioning biological pump. * A study on the timing and nature of recovery of
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
marine ecosystems of Antarctica after the Cretaceous–Paleogene mass extinction, as indicated by data from fossils of benthic molluscs, is published by Whittle ''et al.'' (2019). * A study on the drivers and tempo of biotic recovery after Cretaceous–Paleogene mass extinction, as indicated by data from the Corral Bluffs section of the Denver Basin (Colorado,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
), is published by Lyson ''et al.'' (2019). * Description of the vertebrate assemblage from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
Shine Us locality in the Khaliun Basin (
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 millio ...
) is published by Daxner-Höck ''et al.'' (2019). * Description of reptile and amphibian fossils from the early Miocene localities of the Kilçak section (
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
) is published by Syromyatnikova ''et al.'' (2019). * Description of fossil fish, amphibian and reptilian fauna from the middle Miocene locality Gračanica (Bosnia and Herzegovina) is published online by Vasilyan (2019). * A study on the vertebrate fossils from the early Clarendonian localities within the Goliad Formation in Bee County, Texas, Bee and Live Oak County, Texas, Live Oak Counties in
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
(comprising the Lapara Creek Fauna), and on the Stratigraphy, stratigraphic context of these localities, is published by May (2019). * New late Miocene vertebrate assemblage, including turtle, rodent and xenarthran fossils (among which is the oldest record of an armadillo belonging to the genus ''Dasypus'' reported so far), is described from the Los Alisos locality (Guanaco Formation,
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
) by Ercoli ''et al.'' (2019). * Description of a diverse late Miocene marine fauna from the Bloomfield Quarry (Wilson Grove Formation;
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
), including the most diverse assemblage of fossil Odobenidae, walruses yet reported worldwide from a single locality, is published by Powell ''et al.'' (2019). * Fish, turtle and mammals fossils are described from a locality near Whitehorse (Yukon,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
), probably of Miocene age, by Eberle ''et al.'' (2019). * A study on microscopic traces of hominin and animal activities in the Denisova Cave (
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
), providing the information on the use of this cave over the last 300,000 years, is published by Morley ''et al.'' (2019). * A study on the age of the Pleistocene vertebrate assemblage from the Khok Sung locality (Thailand) is published by Duval ''et al.'' (2019). * Revision of reptile and amphibian fossils from the late Pleistocene collection of the "Caverne Marie-Jeanne" (Hastière-Lavaux, Namur Province, Belgium) is published by Blain ''et al.'' (2019). * New late Pleistocene site Tsaramody (Sambaina basin, Madagascar), preserving diverse subfossil remains of vertebrates, is reported by Samonds ''et al.'' (2019). * A study on the paleoecology and diet of late Pleistocene terrestrial vertebrates known from an asphalt deposit (Project 23, Deposit 1) at Rancho La Brea (
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) is published online by Fuller ''et al.'' (2019). * A study on changes of vegetation in southern Borneo over the past 40,000  calibrated years Before Present, BP, as indicated by data from Saleh Cave (South Kalimantan, Indonesia), is published by Wurster ''et al.'' (2019). * Late Quaternary fossils of vertebrates are described from caves in the Manning Karst Region of eastern New South Wales ( Australia) by Price ''et al.'' (2019). * A study aiming to determine the relationships between extinctions of megafauna, climatic changes and patterns of human appearance in south-eastern Australia over the last 120,000 years is published by Saltré ''et al.'' (2019). * A study on the causes of Holocene extinction of megafauna of Madagascar is published by Laurie Godfrey, Godfrey ''et al.'' (2019). * A review discussing possible links between the fossil record of marine biodiversity, nutrient availability and primary productivity is published online by Martin & Servais (2019). * A study on factors which determined the relative intensity of marine extinctions during greenhouse–icehouse transitions in the Late
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
and the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...
is published online by Saupe ''et al.'' (2019). * A study on the possible relationship between speciation and extinction rates of different groups of organisms and the ages of these groups, as indicated by data from extant and fossil species, is published by Henao Diaz ''et al.'' (2019). * A study on the evolution of bite force of amniotes, as indicated by data from extant and fossil taxa, is published by Sakamoto, Marcello Ruta, Ruta & Venditti (2019). * A study on the phylogenetic distribution, morphological variation and functions of apicobasal ridges (elevated ridges of tooth enamel) in aquatic reptiles and mammals, as indicated by data from extant and fossil taxa, is published by McCurry ''et al.'' (2019). * A study on the impact of uncertainty of stratigraphic age of fossils on studies estimating species divergence times which incorporate fossil taxa, based on data from the fossil record of North American mammals and from the dataset of extant and fossil cetaceans, is published by Barido-Sottani ''et al.'' (2019). * A study evaluating the impact of information about stratigraphic ranges of fossil taxa on the analyses of timing of evolutionary divergence is published online by Püschel ''et al.'' (2019). * A study on anatomical distribution, abundance, geometry, melanin chemistry and elemental inventory of
melanosome A melanosome is an organelle found in animal cells and is the site for synthesis, storage and transport of melanin, the most common light-absorbing pigment found in the animal kingdom. Melanosomes are responsible for color and photoprotection i ...
s in tissues of extant vertebrates, evaluating their implications for reconstructions of internal soft-tissue anatomy in fossil vertebrates, is published by Rossi ''et al.'' (2019). * A study on the chronostratigraphy and
biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Bio ...
of Cenozoic vertebrate (mostly mammal) fossils from the
South Carolina )'' Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = "Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = G ...
Coastal Plain is published by Albright ''et al.'' (2019).


Other research

Other research related to paleontology, including research related to geology, palaeogeography, paleoceanography and paleoclimatology. * A study on the biological oxygen production during the Mesoarchean, as indicated by data from Mesoarchean shales of the Mozaan Group (Pongola Supergroup,
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
) preserving record of a shallow ocean "oxygen oasis", is published by Ossa Ossa ''et al.'' (2019). * A study on the extent of the oxygenation of ocean waters over Continental shelf, continental shelves before the Great Oxidation Event, as indicated by data from 2.5-billion-year-old Mount McRae Shale ( Australia), is published by Ostrander ''et al.'' (2019). * A study on the extent of the oxygenation of shallow oceans 2.45 billion years ago is published by Rasmussen ''et al.'' (2019), who interpret their findings as indicating that oxygen levels both the surface oceans and atmosphere were exceedingly low before the Great Oxidation Event, which the authors interpret as directly caused by evolution of oxygenic photosynthesis. * A study aiming to determine whether the overall size of the biosphere decreased at the end of the Great Oxidation Event, based on data on isotope geochemistry of sulfate minerals from the Belcher Group (subarctic
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
), is published by Hodgskiss ''et al.'' (2019). * Evidence of a burst of Earth's mantle, mantle activity at the end of the Archean (around 2.5 billion years ago) is presented by Marty ''et al.'' (2019), who interpret their findings as lending credence to models advocating a magmatic origin for environmental changes such as the Great Oxidation Event. * A study aiming to determine the effects of competition of early Anoxygenic photosynthesis, anoxygenic phototrophs and primitive Photosynthesis, oxygenic phototrophs on the Earth system, especially on the large-scale oxygenation of Earth's atmosphere ~2.3 billion years ago, is published by Ozaki ''et al.'' (2019). * A study on the geochemistry of mat-related structures and their host sediments from the Francevillian biota, Francevillian Formation (
Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...
) is published by Aubineau ''et al.'' (2019), who evaluate the implications of their findings for the knowledge whether ancient microbes induced illitisation (conversion of smectite to illite–smectite mixed-layer minerals), and for the knowledge of Earth's climate and ocean chemistry in the Paleoproterozoic. * A study on the organic geochemical (biomarker) signatures of the 1.38-billion-years-old black siltstones of the Velkerri Formation ( Australia), and on their implications for inferring the microbial diversity and palaeoenvironment of the Proterozoic Roper Seaway, is published by Jarrett ''et al.'' (2019). * A study on the origins of putative stromatolites and associated carbonate minerals from lacustrine sedimentary rocks of the 1.1-billion-years-old Stoer Group is published by Brasier ''et al.'' (2019). * A study suggesting a link between early evolution and diversification of animals and high availability of copper in the late Neoproterozoic is published by Parnell & Boyce (2019). * A study aiming to determine the cause of the uniquely high amplitudes of Neoproterozoic Δ13C, δ13C excursions is published by Shields ''et al.'' (2019). * A study evaluating the possible relationship between the Cryogenian magmatic activity and the evolution of early life, based on data from the Cryogenian Yaolinghe Group ( China), is published by Long, Zhang & Luo (2019). * Evidence for oxygenated waters near ice sheet grounding lines during the Cryogenian is presented by Lechte ''et al.'' (2019). * A study on ocean oxygen levels during the Ediacaran Ediacaran#Upper and lower boundaries, Shuram negative C-isotope Excursion and the middle Ediacaran, and on their implications for the evolution of the Ediacaran biota, is published by Zhang ''et al.'' (2019). * A study on the causes of widespread preservation of soft-bodied organisms in sandstones of the Ediacara Member in South Australia is published by Liu ''et al.'' (2019). * A study on the seafloor oxygen fugacity in the time of the emergence of the earliest known
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
animals, as inferred from data from the latest Ediacaran Dengying Formation ( China), is published by Ding ''et al.'' (2019). * A study on the process of fossilization of Ediacaran organisms, and on its impact on the preservation of the external shape of these organisms, is published by Bobrovskiy ''et al.'' (2019). * A study on the global extent of the oxygenation of seafloor, surface oceans and atmosphere during early Cambrian is published by Dahl ''et al.'' (2019), who report evidence of two major oceanic anoxic events in the early Cambrian. * A study on nitrogen isotope and organic carbon isotope data from the lower Cambrian Niutitang Formation ( China) is published online by Xu ''et al.'' (2019), who link nitrogen cycle perturbations to animal diversification during the early Cambrian. * A study on the paleoecological characteristics of Cambrian marine ecosystems of central Sonora (Mexico) is published by Romero ''et al.'' (2019). * A study on seawater temperatures during the Cambrian, as indicated by data from oxygen isotope analyses of Cambrian brachiopod shells, is published by Wotte ''et al.'' (2019). * A study on bottom-water redox conditions in the late Cambrian Alum Shale Formation, Alum Shale Sea, as indicated by sedimentary molybdenum contents of the Alum Shale, is published by Dahl ''et al.'' (2019), who interpret their findings as indicating that Anoxic waters, anoxic sulfidic bottom waters were an intermittent rather than persistent feature of Cambrian oceans, and that early animals invaded the seafloor during oxygenated periods. * A study on the Palaeogeography, paleogeographic position of all major Phanerozoic arc-continent Continental collision, collisions, comparing it with the latitudinal distribution of ice-sheets throughout the Phanerozoic, is published by Macdonald ''et al.'' (2019). * A study aiming to determine whether the Ordovician meteor event directly affected Earth's climate and biota is published by Schmitz ''et al.'' (2019). * A review of the evidence of evolutionary radiation of animals throughout the
Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced w ...
, and of environmental changes coincident with these biotic changes, is published by Stigall ''et al.'' (2019). * A study on conodont oxygen isotope compositions in Ordovician samples from Argentina, Argentine Precordillera and
Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, althoug ...
, and on their implications for the knowledge of Paleothermometer, palaeothermometry and drift of the Precordillera in the early Paleozoic, is published online by Albanesi ''et al.'' (2019). * A study on carbon isotope data from stratigraphic sections at Germany Valley (West Virginia) and Union Furnace (
Pennsylvania Pennsylvania (; (Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, Ma ...
) in the Central Appalachian Basin, evaluating its implications for the knowledge of change in atmospheric oxygen levels during the late Ordovician and its possible relationship with early diversification of land plants, is published by Adiatma ''et al.'' (2019). * Signatures of Devonian (Famennian) forests and soils preserved in black shales in the southernmost Appalachian Basin (Chattanooga Shale; Alabama,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) are presented by Lu ''et al.'' (2019). * A study examining the intensity of explosive volcanism from 400 to 200 million years ago, and evaluating its impact on the Late Paleozoic icehouse, late Paleozoic Ice Age, is published by Soreghan, Soreghan & Heavens (2019). * Description of Cisuralian charcoal from the Barro Branco coal seam (Siderópolis Member of the Rio Bonito Formation,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
), and a study on its implications for reconstruction of palaeo-wildfire occurrences in peat-forming vegetation through the Late Palaeozoic in
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final st ...
, is published by Benicio ''et al.'' (2019). * A study on the extent and causes of the end-Capitanian extinction event, based on data from the Middle to Late Permian section of the Sverdrup Basin (Ellesmere Island,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
), is published online by Bond, Wignall & Grasby (2019). * A study on the ocean chemistry during the
Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, a ...
, as indicated by data from a new Stratigraphy, stratigraphic section in Utah, and on its implications for the knowledge of the causes of this extinction, is published by Burger, Estrada & Gustin (2019). * A study aiming to determine the Stratigraphy, stratigraphic position of the end-Permian biotic crisis in the Sydney Basin ( Australia) is published by Fielding ''et al.'' (2019), who also attempt to determine the climate changes in this region concurrent with the end-Permian extinction. * A study on shifts in volcanic activity across the Permian-Triassic boundary, as indicated by measurements of Mercury (element), mercury in marine sections across the Northern Hemisphere, is published by Shen ''et al.'' (2019). * A study on mercury enrichments in Permian-Triassic boundary sections from Lubei (South China craton) and Dalongkou (Junggar terrane), and on their implications for the knowledge of volcanic activity during the Permian-Triassic transition, is published by Shen ''et al.'' (2019). * Evidence of the environmental transition from meandering to braided rivers and of the development of desert-like conditions in the earliest Triassic is reported from Permian-Triassic boundary sections in Shanxi (China) by Zhu ''et al.'' (2019). * A study on the nitrogen isotope variations in oceanic waters in the aftermath of the end-Permian mass extinction is published by Sun ''et al.'' (2019), whose conceptual model indicates ammonium intoxication of the oceans during this time period. * A study on microbially induced sedimentary structures from the Lower Triassic Blind Fiord Formation (Arctic
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
), evaluating their implications for the knowledge of the course of biotic recovery in the aftermath of the Permian–Triassic extinction event, is published online by Wignall ''et al.'' (2019). * A study on the oxygen isotope compositions of discrete conodont elements from the Lower Triassic Mianwali Formation (Pakistan), and on their implications for inferring the timing of temperature changes and the interrelationship between climate and biodiversity patterns during the Early Triassic#Smithian-Spathian extinction, Smithian-Spathian biotic crisis, is published by Goudemand ''et al.'' (2019). * A study on nutrient availability through the Early to Middle Triassic along the northern margin of Pangea is published online by Grasby ''et al.'' (2019). * A study on the character and extent of the Triassic Boreal Ocean delta plain across the area of the present-day Barents Sea, interpreted as the largest delta plain reported so far, is published by Klausen, Nyberg & Helland-Hansen (2019). * A study aiming to determine links between volcanic activity in the Central Atlantic magmatic province, elevated concentrations of Mercury (element), mercury in marine and terrestrial sediments and abnormalities of fossil fern spores across the Triassic-Jurassic boundary in southern Scandinavia and northern Germany is published by Lindström ''et al.'' (2019). * A study aiming to reconstruct the palaeoenvironmental changes of the late Pliensbachian outside of Western
Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
and to test their temporal relation to large igneous province volcanism is published by De Lena ''et al.'' (2019). * Krencker, Lindström & Bodin (2019) present Sedimentology, sedimentological, paleontological and Geochemistry, geochemical evidence from the Central High Atlas Basin (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
) and Jameson Land (Greenland) indicative of the occurrence of a major sea-level drop prior to the onset of the Toarcian oceanic anoxic event. * A study on the duration of the Toarcian oceanic anoxic event, as indicated by data from the Talghemt section in the High Atlas (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
), is published by Boulila ''et al.'' (2019). * A study on the Middle Jurassic palaeoenvironment of La Voulte-sur-Rhône (lagerstätte), La Voulte (France), as indicated by data from exceptionally preserved eyes of the polychelidan lobster ''Voulteryon parvulus'' and from epibiontic brachiopods associated with ''V. parvulus'', is published by Audo ''et al.'' (2019). * A study comparing the Jurassic floras of the Ayuquila Basin and the Otlaltepec Basin (Mexico) and evaluating their implications for the knowledge of the Jurassic environments of these basins is published by Velasco-de León ''et al.'' (2019). * A study on Jurassic paleomagnetism, based on an updated set of Jurassic paleopoles from Adria (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
), is published by Muttoni & Kent (2019). * A study on the chronostratigraphy of the Upper Jurassic Morrison Formation is published by Maidment & Muxworthy (2019). * Evidence of repeated significant oceanic and biotic turnovers in the area of the present-day Gulf of Mexico at the Jurassic-Cretaceous transition is presented by Zell ''et al.'' (2019). * A study on the age of the dinosaur-bearing Upper Jurassic–Lower Cretaceous sediments of western Maestrazgo Basin and South-Iberian Basin (eastern Spain), aiming to also reconstruct the palaeoenvironments of this area on the basis of data from these sediments, is published by Campos-Soto ''et al.'' (2019). * A review of data on the Jurassic and Cretaceous climates of Siberia is published by Rogov ''et al.'' (2019). * A study on global climatic changes during the Early Cretaceous, focusing on the duration and magnitude of Early Cretaceous cold episodes, is published by Vickers ''et al.'' (2019). * Evidence from the Lower Cretaceous strata around the southern margin of the Eromanga Basin ( Australia) indicative of cold (limited glacial and/or seasonal freezing) conditions persisting in Southern Australia through the Hauterivian and the Aptian is presented by Alley, Hore & Frakes (2019). * A study on phototropism in extant trees from Beijing and Jilin Provinces and fossil tree trunks from the Jurassic Tiaojishan Formation, Tiaojishan and Tuchengzi Formation, Tuchengzi formations in Liaoning and Beijing regions ( China), and on its implications for inferring the history of the rotation of the North China Block, is published by Jiang ''et al.'' (2019). * A study on the age of the Cretaceous Cloverly Formation is published by D'Emic ''et al.'' (2019). * Evidence from the chronostratigraphy, fossil content, bracketing facies and ages of the Cretaceous Wayan Formation of Idaho and Vaughn Member of the Blackleaf Formation of Montana, indicating that they represent the same depositional system prior to disruption by subsequent tectonic and volcanic events, is presented by Krumenacker (2019). * A study on Cenomanian plants from the Redmond no.1 mine near Schefferville (Redmond Formation; Labrador Peninsula,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) and on their implications for the knowledge of paleoclimate of this site is published by Demers-Potvin & Larsson (2019). * The first high-resolution record of Cenomanian–Turonian paleotemperatures from the Southern Hemisphere, as indicated by data from the Ocean Drilling Program Site 1138 on the Kerguelen Plateau, is presented by Robinson ''et al.'' (2019). * A study on the impact of marine biogeochemical processes on the Cretaceous Thermal Maximum is published by Wallmann ''et al.'' (2019). * A study on the age of the Upper Cretaceous Wadi Milk Formation (Sudan) is published by Owusu Agyemang ''et al.'' (2019). * A study on Cenomanian to Coniacian polar environmental conditions at eight locations in northeast
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
and northern Alaska is published online by Spicer ''et al.'' (2019). * A study on variability of carbon, oxygen and nitrogen isotopes in multiple tissues from a wide array of extant vertebrate taxa from the Atchafalaya River Atchafalaya Basin, Basin in Louisiana (inferred to be an environmental analogue to the Late Cretaceous coastal floodplains of North America), and on its implications for formulating and testing predictions about ancient ecological communities based on stable isotope data from fossil specimens, is published by Cullen ''et al.'' (2019). * A study on the general distribution and stratigraphy of the lower shale member of the Campanian Aguja Formation (
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
), and a revision of all significant larger vertebrate fossil specimens from these strata, is published by Lehman ''et al.'' (2019). * High-precision dating for the Battle Formation (Alberta,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) is presented by Eberth & Kamo (2019). * High-precision dating and the first calibrated chronostratigraphy for the Horseshoe Canyon Formation (Alberta,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
) is presented by Eberth & Kamo (2019). * A study on the Maastrichtian climate of Arctic Alaska, based on data from the Prince Creek Formation, is published by Salazar-Jaramillo ''et al.'' (2019). * Studies on the timing of the Deccan Traps volcanism close to the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
-
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
boundary are published by Schoene ''et al.'' (2019), who interpret their findings as indicative of four high-volume eruptive periods close to the Cretaceous-Paleogene boundary, the first of which occurred tens of thousands of years prior to both the Chicxulub bolide impact and
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden extinction event, mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million y ...
and by Sprain ''et al.'' (2019), who interpret their findings as indicating that a steady eruption of the flood basalts mostly occurred in the earliest Paleogene. * A study on the environmental variability before and across the Cretaceous-Paleogene mass extinction, as inferred from data on the calcium isotope ratios of Aragonite, aragonitic mollusc shells from the Lopez de Bertodano Formation (Antarctica), is published online by Linzmeier ''et al.'' (2019). * A turbulently deposited sediment package directly overlain by the Cretaceous–Paleogene boundary tonstein is reported from the Tanis site (Hell Creek Formation, North Dakota,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
) by DePalma ''et al.'' (2019), who interpret their findings as indicating that deposition occurred shortly after a major bolide impact, and might have been caused by the Chicxulub crater, Chicxulub impact. * A study on the immediate aftermath of the Chicxulub impact at the Cretaceous–Paleogene boundary, based on data from the Chicxulub crater, is published by Gulick ''et al.'' (2019). * Evidence of rapid ocean acidification in the aftermath of the Chicxulub impact and of the protracted Earth system recovery after the Cretaceous–Paleogene extinction event is presented by Henehan ''et al.'' (2019). * The longest, highest resolution, stratigraphically continuous, single-species
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
l carbon and oxygen isotope records for the Late Maastrichtian to Early
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
from a single site in the South Atlantic Ocean, providing information on the evolution of climate and carbon-cycling during this time period, are presented by Barnet ''et al.'' (2019). * O'Leary ''et al.'' (2019) publish a monograph on the sedimentology and sequence stratigraphy of the part of Mali which was covered by an ancient Inland sea (geology), epeiric sea known as the Trans-Saharan Seaway during the Late Cretaceous and early
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago (Mya) to the beginning of ...
, provide the first formal description of and nomenclature for the Upper Cretaceous and lower Paleogene geological formations of this region, and revise fossil flora and fauna of this region. * Zeebe & Lourens (2019) provide a new absolute astrochronology up to 58 Year#SI prefix multipliers, Ma and a new Paleocene–Eocene boundary age. * A study on stomata of fossil specimens of members of the family Lauraceae from the
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
of Australia and New Zealand, evaluating their implications for reconstructions of Eocene PCO2, ''p''CO2 levels, is published by Steinthorsdottir ''et al.'' (2019). * Climate simulations capturing major climatic features of the Early Eocene and the
Paleocene–Eocene Thermal Maximum The Paleocene–Eocene thermal maximum (PETM), alternatively (ETM1), and formerly known as the "Initial Eocene" or "", was a time period with a more than 5–8 °C global average temperature rise across the event. This climate event o ...
in a state-of-the-art Earth system model are presented by Zhu, Poulsen & Tierney (2019). * A study evaluating the utility of membrane lipids of members of Thaumarchaeota (now Nitrososphaerota) as proxies for the carbon isotope excursion and surface ocean warming, and assessing their implications for the knowledge of the source and size of carbon emissions during the Paleocene–Eocene Thermal Maximum, is published by Elling ''et al.'' (2019). * A study on abundant black charcoal shards from Paleogene sites of Wilson Lake B (New Jersey) and Randall's Farm (Maryland) is published by Fung ''et al.'' (2019), who interpret these shards as most likely to be evidence of widespread wildfires at the Paleocene-Eocene boundary caused by extraterrestrial impact. * A study on the impact of carbon-based greenhouse gas fluxes associated with the North Atlantic Igneous Province on the onset of the Paleocene–Eocene Thermal Maximum is published by Jones ''et al.'' (2019). * Evidence from the Deep Ivorian Basin offshore West Africa (equatorial Atlantic Ocean), indicating that peak warming during the Middle Eocene Climatic Optimum was associated with upper-ocean Stratification (water), stratification, decreased Marine snow#Study, export production, and possibly harmful algal blooms, is presented by Cramwinckel ''et al.'' (2019). * New stable isotopes record of the Middle Eocene Climatic Optimum event is reported from eastern
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
by Giorgioni ''et al.'' (2019). * A study on variations of ocean circulation and marine bioproductivity related to the beginnings of the formation of the Antarctic Circumpolar Current, based on data from Eocene and Oligocene sedimentary drift deposits east of New Zealand, is published by Sarkar ''et al.'' (2019). * A study on changes in surface water temperature in the eastern North Sea Basin during the late Priabonian to earliest Rupelian is published by Śliwińska ''et al.'' (2019). * A study linking the onset or strengthening of an Atlantic meridional overturning circulation to the closure of the Arctic–Atlantic gateway at the Eocene–Oligocene transition is published by Hutchinson ''et al.'' (2019). * A study on the timing of the uplift of the Tibetan Plateau, as indicated by the discovery of the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
Arecaceae, palm fossils in the Lunpola Basin in Tibet, is published by Su ''et al.'' (2019). * A review of vertebrate fossils from the Tibetan Plateau, evaluating their implications for inferring the course of the uplift of the Tibetan Plateau, is published by Deng ''et al.'' (2019). * A study on the impact of changing Eocene paleogeography and climate on the utility of stable isotope paleoaltimetry methods in the studies aiming to reconstruct the elevation history of the Tibetan Plateau is published by Botsyun ''et al.'' (2019). * A study on the causes of the long-term climate cooling during the Neogene is published by Rugenstein, Ibarra & von Blanckenburg (2019). * A study on the climatic and environmental conditions in the Loperot site (Kenya) in the early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
is published by Liutkus-Pierce ''et al.'' (2019). * A study on the timing and course of the separation of the Indian Ocean and the Mediterranean Sea in the Miocene is published by Bialik ''et al.'' (2019). * A study comparing changes of the export of intermediate-depth Pacific waters to the western North Atlantic prior to the closure of the Central American Seaway with records of strength of the Atlantic meridional overturning circulation, evaluating the implications of this data for the knowledge of the timing of closure of the Central American Seaway, is published by Kirillova ''et al.'' (2019). * A study on climatic and environmental changes in central Andes during the late Miocene is published by Carrapa, Clementz & Feng (2019). * A study on the exact age of the marine fauna from the Miocene Chilcatay Formation, Chilcatay and Pisco Formation, Pisco formations (Peru), and on its implications for reconstructions of local paleoenvironment, is published online by Bosio ''et al.'' (2019). * A study on the origin of the African C4 carbon fixation, C4 savannah grasslands is published by Polissar ''et al.'' (2019). * A study on the anatomical traits of teeth and inferred diet of Bovidae, bovids, Suidae, suids and Rhinoceros, rhinocerotids from Kanapoi, and on their implications for reconstructing the environments of this site, is published online by Dumouchel & Bobe (2019). * New spatial data on the Plio-Pleistocene Bolt's Farm pits from the Cradle of Humankind site (
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
) is presented by Edwards ''et al.'' (2019), who also attempt to provide key Biochronology, biochronological ages for the Bolt's Farm deposits. * A study on the global mean sea level during the Pliocene mid-Piacenzian Warm Period is published by Dumitru ''et al.'' (2019). * A study on the amplitude of sea-level variations during the Pliocene is published by Grant ''et al.'' (2019). * Simulations of coevolution of climate, ice sheets and carbon cycle over the past 3 million years are presented by Willeit ''et al.'' (2019). * A study on the age of the Sahara, as indicated by data from Pliocene and Pleistocene paleosols from the Canary Islands, is published by Muhs ''et al.'' (2019). * A study on the latest Villafranchian climate and environment of the area of southern
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
, as indicated by amphibian and reptile fossil record from the Pirro Nord karstic complex, is published by Blain ''et al.'' (2019). * A study on atmospheric gas levels before and after the shift from glacial cycles of 100 thousand years to 40-thousand-year cycles around one million years ago, as inferred from data from ice core samples from the Allan Hills Blue Ice Area (East Antarctica), is published by Yan ''et al.'' (2019). * A study on PCO2, pCO2 levels from 2.6 to 0.8 Year#SI prefix multipliers, Ma is published by Da ''et al.'' (2019), who find no evidence indicating that the Mid-Pleistocene Transition was caused by the decline of pCO2. * A study on changes in winter rainfall in the Mediterranean over the past 1.36 million years is published by Wagner ''et al.'' (2019). * Results of stable carbon and oxygen isotope analyses of tooth enamel samples from
Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
mammals from the Yugong Cave and Baxian Cave ( China) are presented by Sun ''et al.'' (2019), who evaluate the implications of their findings for the knowledge of Pleistocene climatic and environmental changes in South China. * A study on Pleistocene mammal fossils from the Yai Ruak Cave (Krabi Province, Thailand), including the southernmost known record of ''Spotted hyena, Crocuta crocuta ultima'', is published by Suraprasit ''et al.'' (2019), who evaluate the implications of these fossils for reconstructions of the environment in the area of the Malay Peninsula in the Pleistocene. * A study on Acheulean and Middle Stone Age sites from the Eastern Desert (Sudan), preserving stone artifacts, is published by Masojć ''et al.'' (2019), who interpret these sites as evidence of green corridor or corridors across Sahara which made early hominin dispersal possible. * Evidence from oxygen isotope data from Soreq Cave speleothems (Israel), indicative of the occurrence of summer monsoon rainfall in the Middle East during recurrent intervals of the Eemian, last interglacial period (overlapping with archeological indicators of human migration), is presented by Orland ''et al.'' (2019). * A study on the spatial and temporal distribution of ancient peatlands in the past 130,000 years is published by Treat ''et al.'' (2019). * A study on the size of fossil rabbits from 14 late Pleistocene and Holocene archaeological sites in Portugal, and on its implications for the knowledge of temperatures and environment in the area of Portugal during the last glaciation, is published by Davis (2019). * A study on Pleistocene small mammal remains from Stratigraphic Unit V from El Salt site (Alcoy, Spain), evaluating their implications for the knowledge of climatic conditions in the eastern Iberian Peninsula at the time of the disappearance of local Neanderthal populations during Marine isotope stage, Marine Isotope Stage 3, is published by Fagoaga ''et al.'' (2019). * A study on the sedimentary sequence from the Pilauco Bajo, Pilauco site in Chile, evaluating whether evidence from this site is consistent with the Younger Dryas impact hypothesis, is published by Mario Pino Quivira, Pino ''et al.'' (2019). * A study on variations of size of fossil Murinae, murine rodents from Liang Bua (Flores, Indonesia) through time, and on their implications for reconstructions of paleoclimate and paleoenvironment of Flores, is published by Veatch ''et al.'' (2019). * A study on human land use worldwide from 10,000 years before the present to 1850 CE, indicating that Earth was to a large extent transformed by human activity by 3000 years ago, is published by Stephens ''et al.'' (2019). * Evidence for synchronous cyclical changes in monsoon climate, human activity and prehistoric cultural development in the area of northeast China throughout the Holocene is presented by Xu ''et al.'' (2019). * A study on Andean plate tectonics since the late Mesozoic is published by Chen, Wu & Suppe (2019). * A study on the course of the Continental collision, collision of India and Asia, as indicated by Paleomagnetism, palaeomagnetic data from the Burma Terrane, is published by Westerweel ''et al.'' (2019). * A scenario for the genesis of tropical cyclones throughout the Cenozoic is presented by Yan ''et al.'' (2019). * A study on the extent of ice sheets in the Northern Hemisphere throughout the Quaternary is published by Batchelor ''et al.'' (2019). * A new method of concentration of proteins from fossil specimens with high humic content and of removal of humic substances is presented by Schroeter ''et al.'' (2019).


References

* {{reflist 2019 in paleontology,