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Dinocephalosauridae
Dinocephalosauridae is an extinct clade of marine and terrestrial archosauromorph reptiles that lived throughout the Triassic period. Like tanystropheids, they are characterized by their long necks, lengthened by either addition of cervical vertebrae or elongation of the individual bones. Dinocephalosaurids are known from Europe (Poland, Germany, Austria, Netherlands) and China. Some members (i.e. ''Dinocephalosaurus'') are solely marine animals, most likely living near the coastlines of the Tethys Ocean, while other members (i.e. '' Pectodens'' were purely terrestrial, suggesting wide ecological diversity in just the few known species in this family. Classification In 2021, a phylogenetic study was conducted by S. Spiekman, N. Fraser, and T. Schayer in an attempt to clarify the systematics of "protorosaur" groups. A total of 16 individual trees were found using different character scoring methods and unstable OTU exclusions. The results of analysis 3A, with ratio and ordere ...
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Pectodens
''Pectodens'' (meaning "comb tooth") is an extinct genus of archosauromorph reptile which lived during the Middle Triassic in China. The type and only species of the genus is ''P. zhenyuensis'', named by Chun Li and colleagues in 2017. It was a member of the Archosauromorpha, specifically part of the unnatural grouping Protorosauria. However, an unusual combination of traits similar (such as the long neck) and dissimilar (such as the absence of a hook on the fifth metatarsal bone) to other protorosaurs initially led to confusion over its evolutionary relationships. In 2021, it was placed in a newly-established group, Dinocephalosauridae, along with its closest relative ''Dinocephalosaurus''. A small, slender animal measuring long, ''Pectodens'' was named after the peculiar comb-like arrangement of long, conical teeth present in its mouth. Unlike ''Dinocephalosaurus'' and the other reptiles that it was preserved with, well-developed joints and claw-like digits indicate that ''P ...
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Dinocephalosaurus Orientalis
''Dinocephalosaurus'' (meaning "terrible-headed reptile") is a genus of long necked, aquatic protorosaur that inhabited the Triassic seas of China. The genus contains the type and only known species, ''D. orientalis'', which was named by Li in 2003. Unlike other long-necked protorosaurs (which form a group known as the tanystropheids), ''Dinocephalosaurus'' convergently evolved a long neck not through elongation of individual neck vertebrae, but through the addition of neck vertebrae that each had a moderate length. As indicated by phylogenetic analyses, it belonged in a separate lineage that also included at least its closest relative '' Pectodens'', which was named the Dinocephalosauridae in 2021. Like tanystropheids, however, ''Dinocephalosaurus'' probably used its long neck to hunt, utilizing the fang-like teeth of its jaws to ensnare prey; proposals that it employed suction feeding have not been universally accepted. It was probably a marine animal by necessity, as sugge ...
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Dinocephalosaurus
''Dinocephalosaurus'' (meaning "terrible-headed reptile") is a genus of long necked, aquatic protorosaur that inhabited the Triassic seas of China. The genus contains the type and only known species, ''D. orientalis'', which was named by Li in 2003. Unlike other long-necked protorosaurs (which form a group known as the tanystropheids), ''Dinocephalosaurus'' convergently evolved a long neck not through elongation of individual neck vertebrae, but through the addition of neck vertebrae that each had a moderate length. As indicated by phylogenetic analyses, it belonged in a separate lineage that also included at least its closest relative '' Pectodens'', which was named the Dinocephalosauridae in 2021. Like tanystropheids, however, ''Dinocephalosaurus'' probably used its long neck to hunt, utilizing the fang-like teeth of its jaws to ensnare prey; proposals that it employed suction feeding have not been universally accepted. It was probably a marine animal by necessity, as suggest ...
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Czatkowiella
''Czatkowiella'' is an extinct genus of long-necked archosauromorph known from Early Triassic (Olenekian age) rocks of Czatkowice 1, Poland. It was first named by Magdalena Borsuk−Białynicka and Susan E. Evans in 2009 and the type species is ''Czatkowiella harae''. Phylogeny Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... after Borsuk−Białynicka & Evans (2009). Cladogram after Spiekman ''et al.'' 2021: References Prehistoric archosauromorphs Prehistoric reptile genera Olenekian genera Early Triassic reptiles of Europe Fossils of Poland Fossil taxa described in 2009 {{triassic-reptile-stub ...
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Protorosaur
Protorosauria is an extinct polyphyletic group of archosauromorph reptiles from the latest Middle Permian (Capitanian stage) to the end of the Late Triassic (Rhaetian stage) of Asia, Europe and North America. It was named by the English anatomist and paleontologist Thomas Henry Huxley in 1871 as an order, originally to solely contain ''Protorosaurus''. Other names which were once considered equivalent to Protorosauria include Prolacertiformes and Prolacertilia. Protorosaurs are distinguished by their long necks formed by elongated cervical vertebrae, which have ribs that extend backward to the vertebrae behind them. Protorosaurs also have a gap between the quadrate bones and the jugal bones in the back of the skull near the jaw joint, making their skulls resemble those of lizards. While previously thought to be monophyletic, the group is now though to consist of various groups of basal archosauromorph reptiles that lie outside Crocopoda. Classification Protorosauria was cons ...
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Tanystropheid
Tanystropheidae is an extinct family of mostly marine archosauromorph reptiles that lived throughout the Triassic Period. They are characterized by their long, stiff necks formed from elongated cervical vertebrae with very long cervical ribs. Some tanystropheids such as ''Tanystropheus'' had necks that were several meters long, longer than the rest of their bodies. Tanystropheids are known from Europe, Asia (Russia, China, and Saudi Arabia), North America and probably South America (Brazil). The presence of tanystropheids in Europe and China indicate that they lived along much of the coastline of the Tethys Ocean. However, species in western North America are found in terrestrial deposits, suggesting that as a group, tanystropheids were ecologically diverse. Relationships among tanystropheid species have been difficult to resolve because most specimens were flattened during fossilization and are preserved two-dimensionally. Three-dimensional fossils are known from Europe and Nort ...
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Youngina
''Youngina'' is an extinct genus of diapsid reptile from the Late Permian Beaufort Group (''Tropidostoma''-''Dicynodon'' zones) of the Karoo Red Beds of South Africa. This, and a few related forms, make up the family Younginidae, within the Order Eosuchia (proposed by Broom in 1914). Eosuchia, having become a wastebasket taxon for many probably distantly-related primitive diapsid reptiles ranging from the Late Carboniferous to the Eocene, Romer proposed that it be replaced by Younginiformes (that included Younginidae and the Tangasauridae, ranging from the Permian to the Triassic). ''Youngina'' is known from several specimens. Many of these were attributed to as separate genera and species (such as ''Youngoides'' and ''Youngopsis''), but it was later realized that they were not distinct from ''Y. capensis''. The holotype specimen of ''Youngina'' was described briefly in 1914. The "''Youngoides romeri''" specimen was first attributed to ''Youngina'', but later given its eponymou ...
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Orovenator
''Orovenator'' is an extinct genus of diapsid from Lower Permian (Artinskian stage) deposits of Oklahoma, United States. It is known from two partial skulls from the Richards Spur locality in Oklahoma. The holotype OMNH 74606 consists of a partial skull preserving snout and mandible, and the referred specimen, OMNH 74607, a partial skull preserving the skull roof, vertebrae and palatal elements. It was first named by Robert R. Reisz, Sean P. Modesto and Diane M. Scott in 2011 and the type species is ''Orovenator mayorum''. The generic name means "mountain", ''oro'', in Greek in reference to the Richards Spur locality, which was mountainous during the Permian period and "hunter", ''venator'', in Latin. The specific name honours Bill and Julie May. ''Orovenator'' is the oldest and most basal neodiapsid to date. A 2018 redescription by David Ford and Roger Benson found that ''Orovenator'' shared many similarities with varanopids, a group of reptile-like tetrapods tra ...
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Claudiosaurus
''Claudiosaurus'' (''claudus'' is Latin for 'lameness' and ''saurus'' means 'lizard') is an extinct genus of diapsid reptiles from the Permian Sakamena Formation of the Morondava Basin, Madagascar. The pattern of the vertebrate, girle, and limbs indicates that ''Claudiosaurus'' and '' Thadeosaurus'' share a common ancestor. History and discovery ''Claudiosaurus'' is known from the Sakamena Formation of Madagascar. ''Claudiosaurus'' is found from the Late Permian. Although a paper mentions that they have been also found in Early Triassic deposits of Madagascar, citation does not mention that ''Claudiosaurus'' is from Triassic. Biology and description ''Claudiosaurus'' was one of the first members of the Neodiapsida, a group of reptiles containing diapsids more derived than the primitive Araeoscelidia. It had a relatively long body and neck, reaching on overall length of about . It is presumed to have been partially oceanic, living its life in a way similar to the modern ...
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Acerosodontosaurus
''Acerosodontosaurus'' is an extinct genus of neodiapsid reptiles that lived during the Upper Permian of Madagascar. The only species of ''Acerosodontosaurus'', ''A. piveteaui'', is known from a natural mold of a single partial skeleton including a crushed skull and part of the body and limbs. The fossil was discovered in marine deposits of the Lower Sakamena Formation. In conjunction with several skeletal characteristics, this may indicate that ''Acerosodontosaurus'' individuals were at least partially aquatic. ''Acerosodontosaurus'' has generally been considered a " younginiform", part of a paraphyletic grade of Permian diapsids which linked the most basal ("primitive") diapsids ( araeoscelidians such as ''Petrolacosaurus'') to more derived ("advanced") diapsids, including the earliest ancestors of modern reptiles such as crocodilians and lizards. However, its position within the grade is controversial. Initially considered a specimen of the contemporaneous '' Tangasaurus' ...
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Lepidosauromorpha
Lepidosauromorpha (in PhyloCode known as ''Pan-Lepidosauria'') is a group of reptiles comprising all diapsids closer to lizards than to archosaurs (which include crocodiles and birds). The only living sub-group is the Lepidosauria, which contains two subdivisions, Squamata, which contains lizards and snakes, and Rhynchocephalia, the only extant species of which is the tuatara. Lepidosauromorphs are distinguishable from archosauromorphs (reptiles closer to archosaurs) by their primitive sprawling gait (allowing for the same sinusoidal trunk and tail movement seen in fish), the sliding "joint" between the coracoids and the sternum (for a longer stride), and their pleurodont dentition. In contrast, Archosauromorphs possess a parasagittal gait, a reduction in their dermal girdle, a reduction and/or loss of the sternum, and a more thecodont dentition. Living lepidosauromorphs have retained an ectothermic (" cold blooded") metabolism, unlike the ancestral condition in archosauro ...
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Sauria
Sauria is the clade containing the most recent common ancestor of archosaurs (such as crocodilians, dinosaurs, etc.) and lepidosaurs (lizards and kin), and all its descendants. Since most molecular phylogenies recover turtles as more closely related to archosaurs than to lepidosaurs as part of Archelosauria, Sauria can be considered the crown group of diapsids, or reptiles in general. Depending on the systematics, Sauria includes all modern reptiles or most of them (including birds, a type of archosaur) as well as various extinct groups. Sauria lies within the larger total group Sauropsida, which also contains various stem-reptiles which are more closely related to reptiles than to mammals. Prior to its modern usage, "Sauria" was used as a name for the suborder occupied by lizards, which before 1800 were considered crocodilians. Systematics Recent genomic studiesCrawford, Nicholas G., et al. "More than 1000 ultraconserved elements provide evidence that turtles are the sist ...
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