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Acerosodontosaurus
''Acerosodontosaurus'' is an Extinction, extinct genus of Neodiapsida, neodiapsid Reptile, reptiles that lived during the Late Permian of Madagascar. The only species of ''Acerosodontosaurus'', ''A. piveteaui'', is known from a natural wiktionary:mold_fossil, mold of a single partial skeleton including a crushed skull and part of the body and limbs. The fossil was discovered in deposits of the Lower Sakamena Formation. Based on skeletal characteristics, it has been suggested that ''Acerosodontosaurus'' individuals were at least partially Aquatic animal, aquatic. ''Acerosodontosaurus'' has generally been considered a "Younginiformes, younginiform", part of a Paraphyly, paraphyletic Evolutionary grade, grade of Permian Diapsid, diapsids which linked the most basal ("primitive") diapsids (Araeoscelidia, araeoscelidians such as ''Petrolacosaurus'') to more derived ("advanced") diapsids, including the earliest ancestors of modern reptiles such as Crocodilia, crocodilians and Lizard, li ...
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Tangasaurus
''Tangasaurus'' is an extinct genus of aquatic animal, aquatic basal (phylogenetics), basal tangasaurid neodiapsid known from the Late Permian geological timescale, period (late Changhsingian stage) of Tanga Region, Tanga, northeastern Tanzania. It contains a single species, ''Tangasaurus mennelli''. Etymology ''Tangasaurus'' was first described and named by Sidney H. Haughton in 1924 in paleontology, 1924 and the type species is ''Tangasaurus mennelli''. The name of a biological genus, generic name is derived from the name of the Tanga Region in which the only known specimens were found, and Ancient Greek, Greek ''sauros'', meaning "lizard". The specific name (zoology), specific name honors F. P. Mennell, the geologist who reported and collected the specimens of ''Tangasaurus''. Discovery ''Tangasaurus'' is known only from three syntypes which represent two individuals. The small individual, which is well preserved in two specimens (Counter slab, part and counterpart) and is kno ...
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Younginiformes
Younginiformes is a group of diapsid reptiles known from the Permian-Triassic of Africa and Madagascar. It has been used as a replacement for " Eosuchia". Younginiformes (including '' Acerosodontosaurus'', '' Hovasaurus'', '' Kenyasaurus'', '' Tangasaurus'', '' Thadeosaurus'' and ''Youngina'') were historically suggested to be lepidosauromorphs, but are currently thought to be basal non- saurian neodiapsids. The group is sometimes divided into two families, Tangasauridae and Younginidae. The monophyly of the group is disputed. A 2009 study found them to be an unresolved polytomy at the base of Neodiapsida, while a 2011 study recovered the group as paraphyletic. A 2022 study recovered the Younginiformes as a monophyletic group of basal neodiapsid reptiles, also including '' Claudiosaurus'' and '' Saurosternon'' as part of the group. Some younginiforms like ''Hovasaurus'' and ''Acerosodontosaurus'' are thought to have had an amphibious lifestyle, while others like ''Kenyasaurus ...
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Pantestudines
Pantestudines or Pan-Testudines is the proposed group of all Reptile, reptiles more closely related to turtles than to any other living animal. It includes both modern turtles (crown group turtles, also known as Testudines) and all of their extinct relatives (also known as Stem-group, stem-turtles). Pantestudines with a complete shell are placed in the clade Testudinata. Classification The identity of the ancestors and closest relatives of the turtle lineage was a longstanding scientific mystery, though new discoveries and better analyses in the early 21st century began to clarify turtle relationships. They had frequently been considered relatives of the Captorhinidae, captorhinids, which also possessed an anapsid skull configuration. Later, the consensus shifted towards Testudinata's placement within Parareptilia, another "anapsid" clade. Analysis of fossil data has shown that turtles are likely diapsid reptiles, most closely related either to the archosaurs (crocodiles, bird, a ...
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Diapsid
Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The earliest traditionally identified diapsids, the araeoscelidians, appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are often placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. All modern reptiles and birds are placed within the neodiapsid subclade Sauria. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 s ...
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Younginidae
Younginidae is an extinct family of diapsid reptiles from the Late Permian. In a phylogenetic context, younginids are near the base of the clade Neodiapsida. The only species definitively a member of the Younginidae is '' Youngina capensis'' from the Late Permian of South Africa. Several genera have historically been assigned to this family, though more recent phylogenetic analyses have supported relationships with other clades. '' Heleosuchus griesbachi'' from the Late Permian of South Africa may also be a member of the family. Younginidae was traditionally assigned to Eosuchia, an order containing an assemblage of basal diapsids now thought to represent an evolutionary grade rather than a true clade. In 1945 paleontologist Alfred Romer reclassified Younginidae within a new group, Younginiformes, grouping it with the families Tangasauridae and Prolacertidae. Romer considered Younginidae to include many genera that are no longer classified as younginids: '' Paliguana'', '' P ...
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Tangasauridae
Tangasauridae is an extinct family of diapsids known from fossil specimens from Madagascar, Kenya and Tanzania that are Late Permian to Early Triassic in age. Fossils have been found of numerous specimens of common members of this family such as '' Hovasaurus'' in different stages of ontogenic development. Recent material from the Middle Sakamena Formation of the Morondava Basin of Madagascar that dates back to the early Triassic period suggests that the Tangasauridae were relatively unaffected by the Permian-Triassic extinction event. Description and phylogeny Tangasaurids are known to have been a highly derived group of diapsids. One subfamily, Kenyasaurinae, is composed of taxa that were fully terrestrial. They had long toes and highly developed sternums that made them well suited to life on land. On the other hand, the other subfamily, Tangasaurinae, is composed of taxa that were adapted to an aquatic life. They had webbed feet and a laterally compressed tails that al ...
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Araeoscelidia
Araeoscelidia or Araeoscelida is a clade of extinct tetrapods (traditionally classified as diapsid reptiles) superficially resembling lizards, extending from the Late Carboniferous to the Early Permian. The group contains the genera '' Araeoscelis'', ''Petrolacosaurus'', the possibly aquatic '' Spinoaequalis'', and less well-known genera such as ''Kadaliosaurus'' and '' Zarcasaurus''. This clade is usually considered to be the sister group to all (currently known) later diapsids. Description Araeoscelidians were small animals (less than one meter in length) looking somewhat like lizards, though they are only distantly related to true lizards. They differ from other, earlier sauropsids by their slender limbs, their elongated tail, and of course by the presence of two temporal openings, the feature defining the diapsid condition. In '' Araeoscelis'', only the upper temporal opening remains, thus resulting in a derived euryapsid condition. Genera Araeoscelidia includes well-know ...
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Sauria
Sauria is the clade of diapsids containing the most recent common ancestor of Archosauria (which includes crocodilians and birds) and Lepidosauria (which includes squamates and the tuatara), and all its descendants. Since most molecular phylogenies recover turtles as more closely related to archosaurs than to lepidosaurs as part of Archelosauria, Sauria can be considered the crown group of diapsids, or reptiles in general. Depending on the systematics, Sauria includes all modern reptiles or most of them (including birds, a type of archosaur) as well as various extinct groups. Sauria lies within the larger total group Sauropsida, which also contains various stem-reptiles which are more closely related to reptiles than to mammals. Prior to its modern usage, "Sauria" was used as a name for the suborder occupied by lizards, which before 1800 were considered crocodilians. Systematics Sauria was historically used as a partial equivalent for Squamata (which contains lizards and sna ...
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Paris
Paris () is the Capital city, capital and List of communes in France with over 20,000 inhabitants, largest city of France. With an estimated population of 2,048,472 residents in January 2025 in an area of more than , Paris is the List of cities in the European Union by population within city limits, fourth-most populous city in the European Union and the List of cities proper by population density, 30th most densely populated city in the world in 2022. Since the 17th century, Paris has been one of the world's major centres of finance, diplomacy, commerce, culture, Fashion capital, fashion, and gastronomy. Because of its leading role in the French art, arts and Science and technology in France, sciences and its early adoption of extensive street lighting, Paris became known as the City of Light in the 19th century. The City of Paris is the centre of the Île-de-France region, or Paris Region, with an official estimated population of 12,271,794 inhabitants in January 2023, or ...
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France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlantic, North Atlantic, the French West Indies, and List of islands of France, many islands in Oceania and the Indian Ocean, giving it Exclusive economic zone of France, one of the largest discontiguous exclusive economic zones in the world. Metropolitan France shares borders with Belgium and Luxembourg to the north; Germany to the northeast; Switzerland to the east; Italy and Monaco to the southeast; Andorra and Spain to the south; and a maritime border with the United Kingdom to the northwest. Its metropolitan area extends from the Rhine to the Atlantic Ocean and from the Mediterranean Sea to the English Channel and the North Sea. Its Regions of France, eighteen integral regions—five of which are overseas—span a combined area of and hav ...
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National Museum Of Natural History (France)
The French National Museum of Natural History ( ; abbr. MNHN) is the national natural history museum of France and a of higher education part of Sorbonne University. The main museum, with four galleries, is located in Paris, France, within the Jardin des Plantes on the left bank of the River Seine. It was formally founded in 1793, during the French Revolution, but was begun even earlier in 1635 as the royal garden of medicinal plants. The museum now has 14 sites throughout France. Since the 2014 reform, it has been headed by a chairman, assisted by deputy managing directors. The Museum has a staff of approximately 2,350 members, including six hundred researchers. It is a member of the national network of naturalist collections (RECOLNAT). History 17th–18th century File:Jardin du roi 1636.png, The Royal Garden of Medicinal Plants in 1636 File:Buffon statue dsc00979.jpg, Statue of Georges-Louis Leclerc, Comte de Buffon in the formal garden File:Buffon, Georges Louis - Lecler ...
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Turtle
Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including land-dwelling tortoises and freshwater terrapins. They are found on most continents, some islands and, in the case of sea turtles, much of the ocean. Like other Amniote, amniotes (reptiles, birds, and mammals) they breathe air and do not lay eggs underwater, although many species live in or around water. Turtle shells are made mostly of bone; the upper part is the domed Turtle shell#Carapace, carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scale (anatomy), scales made of keratin, the material of hair, horns, and claws. The carapace bones deve ...
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