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Ornithocheirae
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early Cretaceous, Early to Late Cretaceous Period (geology), periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs have been discovered worldwide except Antarctica, though most genera have been recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by pteranodontians and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not ''Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also regarded to have some of the largest pterosaur wingspa ...
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Ornithocheirus
''Ornithocheirus'' (from Ancient Greek "ὄρνις", meaning bird, and "χεῖρ", meaning hand) is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco. Several species have been referred to the genus, most of which are now considered as dubious species, or members of different genera, and the genus is now often considered to include only the type species, ''Ornithocheirus simus''. Species have been referred to ''Ornithocheirus'' from the mid-Cretaceous period of North America, Europe and South America, but ''O. simus'' is known mostly from the United Kingdom, though a specimen referred to ''O.'' cf. ''simus'' is also known from the Late Cretaceous Kem Kem Group of Morocco. Because ''O. simus'' was originally named based on poorly preserved fossil material, the genus ''Ornithocheirus'' has suffered enduring problems of zoological nomenclature. Fossil remains of ''Ornithocheirus'' have been recovered mainly ...
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Anhangueridae
Anhangueridae (alternatively called Ornithocheiridae, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 90 million years ago. Anhanguerids are generally infamous for having an enormously controversial and very confusing taxonomy. Although agreements that these animals were related, and therefore similar to istiodactylids and pteranodontians, there is still no virtual consensus over the exact content and interrelationships of this group. Anhanguerids were the most successful pterosaurs during their reign, and were also the largest pterosaurs before the appearance of the azhdarchids such as ''Quetzalcoatlus''. Anhanguerids were excellent fish hunters, using various flight techniques to catch their prey, and were also capable of flying great distances without fla ...
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Targaryendraconia
Targaryendraconia is an extinct clade of lanceodontian pterosaurs that lived from the Early to Late Cretaceous period in Europe, North America, South America, and Australia. Classification Below is a cladogram following a topology by Pêgas and colleagues in 2019. In their analysis, they recovered Targaryendraconia as the sister taxon of the clade Anhangueria, both of which are within the more inclusive group Ornithocheirae. Targaryendraconia is split into two families: the Targaryendraconidae, which contains '' Aussiedraco'', '' Barbosania'', and '' Targaryendraco'', and the Cimoliopteridae, which contains '' Aetodactylus'', '' Camposipterus'', and '' Cimoliopterus''.Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, Contrastingly, a study by Alexander Averianov ...
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Tropeognathus
''Tropeognathus'' (meaning "keel jaw") is a genus of large pterosaurs from the late Early Cretaceous of South America. This genus is considered to be a member of the family Anhangueridae, however, several studies have also recovered it within another family called Ornithocheiridae. Both of these families are diverse groups of pterosaurs known for their keel-tipped snouts and large size. ''Tropeognathus'' is regarded as the largest pterosaur found in the Southern Hemisphere, only rivaled by the huge azhdarchids. The type and only species is ''Tropeognathus mesembrinus''. Fossil remains of ''Tropeognathus'' have been recovered from the Romualdo Formation, which is a Lagerstätte located in the Santana Group of the Araripe Basin in northeastern Brazil. Discovery and naming In the 1980s the German paleontology museum ''Bayerische Staatssammlung für Paläontologie und historische Geologie'' in Munich acquired a pterosaur skull from Brazilian fossil dealers that had probably been ...
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Pterosaurs
Pterosaurs are an extinct clade of flying reptiles in the Order (biology), order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved flying and gliding animals, powered flight. Their wings were formed by a membrane of skin, muscle, and other tissue (biology), tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were ...
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Istiodactylidae
Istiodactylidae is a small Family (biology), family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus ''Ornithodesmus''. Systematics and distribution Remains of taxa that can be confidently assigned to Istiodactylidae have been found in the UK and China, in rocks dating from the Early Cretaceous period (Barremian to Aptian stage). Victoria Arbour, Arbour and Currie (2011) described Canadian ''Gwawinapterus beardi'' as a member of Istiodactylidae living in the late Cretaceous (upper Campanian stage); however, Witton (2012) suggested the tooth replacement pattern in this animal does not match that of pterosaurs, suggesting that the species might be non-pterosaurian. Additional research suggested that the species was in fact a fish. The earliest known species might be ''Archaeoistiodactylus linglongtaensis'', from the Middle Jurassic of China; however, it also has been suggested that the holotype speci ...
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Boreopteridae
Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the clade containing the common ancestor of '' Boreopterus'' and '' Feilongus'' and all its descendants, which the authors reclassified as close relatives of the ornithocheirids. (''Feilongus'' had originally been considered a gallodactylid). Many possible boreopterids were subsequently described, one possible example being '' Aetodactylus'', which has been claimed to be similar to '' Boreopterus''. Originally considered close relatives of the ornithocheirids, many of these supposed boreopterids have been found to belong to other groups of the pterodactyloid lineage. In 2012, a phylogenetic analysis by Lü ''et al.'' divided the Boreopteridae into two subfamilies: Boreopterinae, comprising ''Boreopterus'' and ''Zhenyuanopterus'', and Moganopt ...
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Maaradactylus Spielbergi
''Maaradactylus'' is a genus of anhanguerid pterodactyloid pterosaur known from the Lower Cretaceous period (Aptian to Albian stages) of the Romualdo Formation of northeastern Brazil. Discovery ''Maaradactylus'' is based on the Museu Paleontologico de Santana do Cariri specimen MPSC R 2357, a skull, atlas, and axis discovered in 2010 in the Aptian—Albian-age Romualdo Formation of Sítio São Gonçalo, Santana do Cariri, Ceará, in the Araripe Basin of Brazil. ''Maaradactylus'' was described by Renan Bantim and colleagues in 2014. The type species is ''Maaradactylus kellneri''. The generic name refers to Maara, in the legends of the Cariri the daughter of a chief, by sorcery changed into a river monster with long teeth, devouring fishermen. The suffix ~''dactylus'' is common in the names of pterosaurs and is derived from Greek δάκτυλος, ''daktylos'', "finger", referring to the long (fourth) wing finger. The specific name honors Alexander Kellner, Brazil's f ...
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Period (geology)
The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (a scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardised international units of geological time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. Antarctica is the fifth-largest continent, being about 40% larger than Europe, and has an area of . Most of Antarctica is covered by the Antarctic ice sheet, with an average thickness of . Antarctica is, on average, the coldest, driest, and windiest of the continents, and it has the highest average elevation. It is mainly a polar desert, with annual Climate of Antarctica#Precipitation, precipitation of over along the coast and far less inland. About 70% of the world's freshwater reserves are frozen in Antarctica, which, if melted, would raise global sea levels by almost . Antarctica holds the record for the Lowest temperature recorded on Earth, lowest measured temperature on Earth, . The coastal regions can reach temperatures over in the ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of '' Tiktaalik'' in the arctic of Canada. Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are sometimes considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. Early edition, published online before prin ...
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