Targaryendraconia is an extinct

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

of lanceodontian

pterosaurs Pterosaurs are an extinct clade of flying reptiles in the Order (biology), order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosau ...

that lived from the Early to

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

period in

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, and

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Classification

Below is a

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

following a topology by Pêgas and colleagues in 2019. In their analysis, they recovered Targaryendraconia as the sister taxon of the clade Anhangueria, both of which are within the more inclusive group

Ornithocheirae Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early Cretaceous, Early to Late Cretaceous Period (geology), period ...

. Targaryendraconia is split into two families: the Targaryendraconidae, which contains '' Aussiedraco'', '' Barbosania'', and '' Targaryendraco'', and the Cimoliopteridae, which contains '' Aetodactylus'', '' Camposipterus'', and '' Cimoliopterus''.Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, Contrastingly, a study by Alexander Averianov in 2025, describing the new targaryendraconian '' Saratovia'', failed to recover the existence of a cimoliopterid clade. Instead, ''Cimoliopterus cuvieri'', ''Cimoliopterus dunni'', ''Saratovia'', and a grouping of ''Camposipterus'' and ''Aetodactylus'' formed an unresolved

polytomy An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree ...

with Targaryendraconidae. He argued that the distinction between the two groups of targaryendraconian were likely caused due to the incompleteness of known remains; cimoliopterids are characterized by traits of the upper jaw, and targaryendraconids by those of the lower jaw. Most of the known genera are only known from upper or lower jaw, and the complete skull of ''Barbosania'' is preserved in a manner preventing study of the interior surface of the jaw. It was also noted that several uniting characteristics of Anhangueria and Anhangueridae are not evaluable in most or all members of Targaryendraconia.

References

Pteranodontoidea Hauterivian first appearances {{pterosaur-stub