Holoaden
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Holoaden
''Holoaden'', the highland frogs, is a small genus of frogs in the family Strabomantidae. The species are endemic to south-eastern Brazil. Taxonomy ''Holoaden'' is the type genus of subfamily Holoadeninae that was erected in 2008 and placed in the newly erected family Strabomantidae. Prior to this, it had been placed in the genus Brachycephalidae. Description ''Holoaden'' grow to snout–vent length at most. Head is narrower than the body. Differentiated Tympanum (anatomy), tympanic membrane and tympanic annulus are absent. Dorsum (anatomy), Dorsum is highly glandular. Venter is areolate. Species There are four recognized species in this genus: * ''Holoaden bradei'' Lutz, 1958 * ''Holoaden luederwaldti'' Miranda-Ribeiro, 1920 * ''Holoaden pholeter'' Pombal, Siqueira, Dorigo, Vrcibradic, and Rocha, 2008 * ''Holoaden suarezi'' Martins and Zaher, 2013 References

{{Taxonbar, from=Q2129401 Holoaden, Strabomantidae Endemic frogs of Brazil Amphibian genera Taxa named by Al ...
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Holoaden Bradei
''Holoaden bradei'', or the Itatiaia highland frog, is a species of frog in the family Strabomantidae. It is endemic to the Itatiaia Mountains of south-eastern Brazil. It is restricted to an area of 10 km2 at an elevation of 2,400 to 2,600, the type locality being at 22°21′S, 44°44′W at a location known as Alto Itatiaia. ''H. bradei'' is found among stones, mulch, and terrestrial bromeliads, in which it breeds through direct development. ''H. bradei'' may already be extinct in the wild because it has not been recorded in the wild for several decades.“Itatiaia Highland Frog Photos and Facts.�Arkive.org/ref> Description It is a small frog that has an olive-brown skin color with irregular dark patches around its body. The belly of the frog is a cream color, and the tips of the frog's hands are a bright yellow. It has a large, wide head with an oval body and a rounded snout. It has short limbs, and has large eyes that face forward. The Highland frog's eyes have diamon ...
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Holoaden
''Holoaden'', the highland frogs, is a small genus of frogs in the family Strabomantidae. The species are endemic to south-eastern Brazil. Taxonomy ''Holoaden'' is the type genus of subfamily Holoadeninae that was erected in 2008 and placed in the newly erected family Strabomantidae. Prior to this, it had been placed in the genus Brachycephalidae. Description ''Holoaden'' grow to snout–vent length at most. Head is narrower than the body. Differentiated Tympanum (anatomy), tympanic membrane and tympanic annulus are absent. Dorsum (anatomy), Dorsum is highly glandular. Venter is areolate. Species There are four recognized species in this genus: * ''Holoaden bradei'' Lutz, 1958 * ''Holoaden luederwaldti'' Miranda-Ribeiro, 1920 * ''Holoaden pholeter'' Pombal, Siqueira, Dorigo, Vrcibradic, and Rocha, 2008 * ''Holoaden suarezi'' Martins and Zaher, 2013 References

{{Taxonbar, from=Q2129401 Holoaden, Strabomantidae Endemic frogs of Brazil Amphibian genera Taxa named by Al ...
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Holoaden Suarezi
''Holoaden'', the highland frogs, is a small genus of frogs in the family Strabomantidae. The species are endemic to south-eastern Brazil. Taxonomy ''Holoaden'' is the type genus of subfamily Holoadeninae that was erected in 2008 and placed in the newly erected family Strabomantidae. Prior to this, it had been placed in the genus Brachycephalidae. Description ''Holoaden'' grow to snout–vent length at most. Head is narrower than the body. Differentiated tympanic membrane and tympanic annulus are absent. Dorsum is highly glandular. Venter is areolate. Species There are four recognized species in this genus: * ''Holoaden bradei ''Holoaden bradei'', or the Itatiaia highland frog, is a species of frog in the family Strabomantidae. It is endemic to the Itatiaia Mountains of south-eastern Brazil. It is restricted to an area of 10 km2 at an elevation of 2,400 to 2,600, ...'' Lutz, 1958 * '' Holoaden luederwaldti'' Miranda-Ribeiro, 1920 * '' Holoaden pholeter'' Pombal, Si ...
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Strabomantidae
The Strabomantidae, sometimes called cloud forest landfrogs, are a family of frogs native to South America. These frogs lack a free-living larval stage and hatch directly into miniature "froglets". This family includes '' Pristimantis'', the most speciose genus of any vertebrate. Based on molecular dating, the Holoadeninae in particular appears to have originated in the Eocene. The Oligocene-Miocene transition was also found to be important to their biogeographical history. However, the lack of Holoadeninae fossils and limited nucleotide Nucleotides are Organic compound, organic molecules composed of a nitrogenous base, a pentose sugar and a phosphate. They serve as monomeric units of the nucleic acid polymers – deoxyribonucleic acid (DNA) and ribonucleic acid (RNA), both o ... data ambiguates the results, by the authors' admission. Systematics The following subfamilies and genera are in the family Strabomantidae. *Holoadeninae **'' Bahius'' Dubois, Ohler, and Pyron, ...
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Holoadeninae
The Strabomantidae, sometimes called cloud forest landfrogs, are a family of frogs native to South America. These frogs lack a free-living larval stage and hatch directly into miniature "froglets". This family includes ''Pristimantis'', the most speciose genus of any vertebrate. Based on molecular dating, the Holoadeninae in particular appears to have originated in the Eocene. The Oligocene-Miocene transition was also found to be important to their biogeographical history. However, the lack of Holoadeninae fossils and limited nucleotide data ambiguates the results, by the authors' admission. Systematics The following subfamilies and genera are in the family Strabomantidae. *Holoadeninae **'' Bahius'' Dubois, Ohler, and Pyron, 2021 **'' Barycholos'' Heyer, 1969 **''Bryophryne'' Hedges, Duellman, and Heinicke, 2008 **'' Euparkerella'' Griffiths, 1959 **''Holoaden'' Miranda-Ribeiro, 1920 **'' Microkayla'' De la Riva, Chaparro, Castroviejo-Fisher, and Padial, 2017 **'' Noblella'' ...
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Holoaden Luederwaldti
''Holoaden luederwaldti'' is a species of frog in the family Strabomantidae. It is endemic to Brazil. Its natural habitats are subtropical or tropical high-altitude shrubland, subtropical or tropical high-altitude grassland, and rocky areas. It is threatened by habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease .... References Holoaden Endemic frogs of Brazil Taxa named by Alípio de Miranda-Ribeiro Amphibians described in 1920 Taxonomy articles created by Polbot {{Strabomantidae-stub ...
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Alípio De Miranda-Ribeiro
Alípio de Miranda-Ribeiro (21 February 1874, Rio Preto, Minas Gerais – 8 January 1939) was a Brazilian herpetologist and ichthyologist. His son, Paulo de Miranda-Ribeiro (1901–1965), was also a zoologist. From an early age Alípio de Miranda-Ribeiro had a passion for natural history; as an adolescent he translated works of Buffon into Portuguese. He studied medicine in Rio de Janeiro, and from 1894 worked as a preparator at the National Museum of Brazil. Here he later served as secretary (from 1899), and director of the department of zoology (from 1929). During his career he explored the Amazon region many times, and under the direction of Candido Rondon (1865–1958) he took part in installing the first telegraph through the Amazon and Mato Grosso. In 1911, after visiting museums and fishery programs in Europe and the United States, he founded a fisheries inspectorate in Brazil, the first official services on fisheries in the nation. In 1911 he published the highly re ...
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Endemic Frogs Of Brazil
Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can also be referred to as an ''endemism'' or, in scientific literature, as an ''endemite''. Similarly, many species found in the Western ghats of India are examples of endemism. Endemism is an important concept in conservation biology for measuring biodiversity in a particular place and evaluating the risk of extinction for species. Endemism is also of interest in evolutionary biology, because it provides clues about how changes in the environment cause species to undergo range shifts (potentially expanding their range into a larger area or becomi ...
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Tympanum (anatomy)
The tympanum is an external hearing structure in animals such as mammals, birds, some reptiles, some amphibians and some insects. Using sound, vertebrates and many insects are capable of sensing their prey, identifying and locating their predators, warning other individuals, and locating potential mates and rivals by hearing the intentional or unintentional sounds they make. In general, any animal that reacts to sounds or communicates by means of sound, needs to have an auditory mechanism. This typically consists of a membrane capable of vibration known as the tympanum, an air-filled chamber and sensory organs to detect the auditory stimuli. Insects Amphibia Anura In frogs and toads, the tympanum is a large external oval shape membrane made up of nonglandular skin. It is located just behind the eye. It does not process sound waves; it simply transmits them to the inner parts of the amphibian's ear, which is protected from the entry of water and other foreign objects. A fr ...
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Dorsum (anatomy)
Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provides a definition of what is at the front ("anterior"), behind ("posterior") and so on. As part of defining and describing terms, the body is described through the use of anatomical planes and axes. The meaning of terms that are used can change depending on whether a vertebrate is a biped or a quadruped, due to the difference in the neuraxis, or if an invertebrate is a non-bilaterian. A non-bilaterian has no anterior or posterior surface for example but can still have a descriptor used such as proximal or distal in relation to a body part that is nearest to, or furthest from its middle. International organisations have determined vocabularies that are often used as standards for subdisciplines of anatomy. For example, '' Termi ...
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Species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. Other ways of defining species include their karyotype, DNA sequence, morphology (biology), morphology, behaviour, or ecological niche. In addition, palaeontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. About 14% of these had been described by 2011. All species (except viruses) are given a binomial nomenclature, two-part name, a "binomen". The first part of a binomen is the name of a genus to which the species belongs. The second part is called the specific name (zoology), specific name or the specific ...
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