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Bufonid
A true toad is any member of the family Bufonidae, in the order Anura (frogs and toads). This is the only family of anurans in which all members are known as toads, although some may be called frogs (such as harlequin frogs). The bufonids now comprise more than 35 genera, ''Bufo'' being the best known. History Bufonidae is thought to have originated in South America. Some studies date the origin of the group to after the breakup of Gondwana, about 78 - 98 million years ago in the Late Cretaceous. In contrast, other studies have dated the origin of the group to the early Paleocene. The bufonids likely radiated out of South America during the Eocene, with the entire radiation occurring during the Eocene to Oligocene, marking an extremely rapid divergence likely facilitated by the Paleogene's changing climatic conditions. Taxonomy The following phylogeny of most genera in the family is based on Portik and Papenfuss, 2015:, Chan ''et al.'', 2016, Chandramouli ''et al.'', 2016 ...
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Nimbaphrynoides
''Nimbaphrynoides'' is a monotypic genus of true toads from highlands in the Mount Nimba region of the West African countries of Guinea, Liberia, and Côte d'Ivoire. The sole species is ''Nimbaphrynoides occidentalis''. Along with '' Nectophrynoides'', '' Eleutherodactylus jasperi'', and ''Limnonectes larvaepartus'', ''Nimbaphrynoides'' is one of the only anurans that combine internal fertilization with ovoviparity.Wells (2007). ''The Ecology and Behavior of Amphibians.'' Pp. 486–487. Taxonomy and systematics ''Nimbaphrynoides occidentalis'' was first described as ''Nectophrynoides occidentalis'' by Fernand Angel in 1943. A second species from Mount Nimba, ''Nectophrynoides liberiensis'', was described in 1979. The two were then transferred to a newly erected genus ''Nimbaphrynoides'' in 1987. The two species were similar except for the larger size of ''N. liberiensis'' compared to ''N. occidentalis'' and some small colour differences. However, their calls are similar, and t ...
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Atelopus
''Atelopus'' is a large genus of Bufonidae, commonly known as harlequin frogs or toads, from Central and South America, ranging as far north as Costa Rica and as far south as Bolivia. ''Atelopus'' species are small, generally brightly colored, and diurnal. Most species are associated with mid-to-high elevation streams. This genus has been greatly affected by amphibian declines, and many species are now considered endangered, while others already are extinct. Almost 40% of the described species in the genus are considered possibly extinct; this is raised to 45% when data deficient species are added; this number may be even higher, given that the genus contains many undescribed species that could also be extinct, and many of the species considered critically endangered but extant may have gone extinct after the last surveys that detected them, or could go extinct in the future. While threatened by habitat loss, pollution, and introduced species, the primary cause of these decli ...
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Incilius
''Incilius'' is genus of toads in the true toad family, Bufonidae. They are sometimes known as the Central American toads or Middle American toads and are found in southern USA, Mexico, Central America, and northern Pacific South America (Colombia and Ecuador). They are an ecologically and biogeographically diverse group of toads, including micro-endemic species such as ''Incilius spiculatus'' that are restricted to undisturbed cloud forests, and widespread lowland species such as ''Incilius valliceps'' that predominantly occur in disturbed habitats. Taxonomy and systematics This genus was first described in 1863 by Edward Drinker Cope who designated the type species as '' Incilius coniferus''. This proved unpopular and these toads were known under the genus ''Bufo'' until the early 2000s. The current delineation of the genus follows Mendelson ''et al''. (2011) who brought ''Cranopsis''/''Cranophryne''/''Ollotis'' and ''Crepidius''/''Crepidophryne'' into synonymy with ''Inciliu ...
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Peltophryne
''Peltophryne'' is a genus of true toads in the family Bufonidae, from the Greater Antilles (Cuba, Isla de Juventud, Hispaniola, Puerto Rico). With ten endemic species, Cuba hosts the highest diversity. Hispaniola has three endemics and Puerto Rico and the Virgin Islands combined has one. Description ''Peltophryne'' range is size from the relatively small '' Peltophryne cataulaciceps'' with a snout–vent length (SVL) of to the large '' Peltophryne peltocephala'' with SVL of . The skull is as long as wide and contains some unique osteological features (thickened dermal tissue covering the snout and usually ossified into a pair of rostral bones, and squamosal-maxillary articulation). These are considered to be derived characters that set these toads apart from other bufonids. Taxonomy The genus was erected by Leopold Fitzinger in 1843, but placed in synonymy with ''Bufo'' by Albert Günther in 1859. Subsequent work has considered ''Peltophryne'' either as a valid genus, a subgen ...
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Nannophryne
''Nannophryne'' is a small genus of true toads, family Bufonidae, from South America. They are found in central Andean Peru south to southern Chile and adjacent Argentina. Taxonomy The genus was placed in synonymy with ''Bufo'' by George Albert Boulenger in 1894, but it was resurrected in 2006 by Frost when splitting the then very large ''Bufo'' into smaller, monophyletic genera. This change was controversial but has now largely been accepted. The alternative, treating ''Nannophryne'' as a subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed betw ... of ''Bufo'', still has some following. Species There are four species in this genus: * '' Nannophryne apolobambica'' (De la Riva, Ríos, and Aparicio, 2005) * '' Nannophryne cophotis'' (Boulenger, 1900) * '' Nannophryne corynetes'' (Due ...
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Amazophrynella
''Amazophrynella'' is a genus of toads in the family Bufonidae. They are found throughout the Amazon Basin. Taxonomy ''Amazophrynella'' was separated from ''Dendrophryniscus'' in 2012 based on molecular genetic evidence that indicated deep divergence between an Amazonian and an Atlantic Forest clade, the latter retaining the name ''Dendrophryniscus'' whereas the former was described as a new genus ''Amazonella'', later amended to ''Amazophrynella'' because of homonymy. The analysis suggested that ''Amazophrynella'' is the sister taxon of ''Dendrophryniscus'', but a later study has suggested a more distant relationship. Description ''Amazophrynella'' are small toads measuring in snout–vent length. The hind limbs are well developed. The parotoid glands are absent, as are vocal slits and tympana. The skin is uniformly and finely granulose. Dorsal coloration is cryptic. ''Amazophrynella'' are pond breeders, which is presumably an ancestral trait in bufonids (most ''Dendrophryni ...
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Melanophryniscus
''Melanophryniscus'' is a genus of toads in the family Bufonidae. They are found in northern half of Argentina, southern Bolivia, southern Brazil, Paraguay, and Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering .... Common name South American redbelly toads has been coined for them. Species There are 31 recognized species: The AmphibiaWeb also recognizes ''Melanophryniscus orejasmirandai'', which the Amphibian Species of the World treats as synonym of ''Melanophryniscus pachyrhynus''. References External links * Taxo''Melanophryniscus''at https://web.archive.org/web/20160606043808/http://www.itis.gov/index.html. (Accessed: May 3, 2008). * Taxo''Melanophryniscus''at https://web.archive.org/web/20080501142231/http://data.gbif.org/welcome.htm Amphibians of South ...
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Frog
A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" '' Triadobatrachus'' is known from the Early Triassic of Madagascar, but molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history. An adult frog has a stout body, protruding eyes, anteriorly-attached tongue, limbs folded underneath, and no tail (the tail of tailed frogs ...
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Oreophrynella
''Oreophrynella'', commonly known as bush toads, is a genus of true toads native to the tepuis of southern Venezuela and adjacent Guyana. The distribution of some species is restricted to a couple of tepuis or even a single tepui, as in the case of '' Oreophrynella weiassipuensis'', which occurs on Wei-Assipu-tepui. Description Species of the genus ''Oreophrynella'' are small frogs, less than in snout–vent length. They are characterized by opposable digits of the foot, dorsal skin that bears tubercules, and direct development (i.e., there is no free-living larval stage). The presence of opposable digits, unique among bufonids, in combination with an extension of the interdigital integument and the relative length/orientation of the digits, is likely to be an adaptation to facilitate life on rocky tepui summits and an exaptation to arboreality The genus also displays cranial simplification in the form of cranial fontanelles and absence of the quadratojugal, which may be driv ...
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Sclerophrys
''Sclerophrys'' is a genus of "true toads", family Bufonidae, native to Africa and the southern Arabian Peninsula. Originally, all of these species were classified in the genus ''Bufo''. The genus, originally named ''Amietophrynus'', was split due to large enough taxonomic divergence. Ohler and Dubois showed in 2016 that ''Sclerophrys capensis'' Tschudi, 1838 is the same species as '' Bufo regularis rangeri'' Hewitt, 1935, the type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ... of ''Amietophrynus''. Because the former name is older, the implication is that ''Amietophrynus'' is a junior synonym of ''Sclerophrys''. Species The following species are recognized in the genus ''Sclerophrys''. References Amphibian genera Amphibians of Africa Amphibians of Asia T ...
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Didynamipus
The four-digit toad or dwarf toad (''Didynamipus sjostedti'') is a species of toad in the family Bufonidae. It is found in Bioko (Equatorial Guinea), southwestern Cameroon, and southeastern Nigeria. It is the only species in the genus ''Didynamipus''. Etymology The specific name ''sjostedti'' honours Bror Yngve Sjöstedt, a Swedish entomologist and ornithologist who collected the type series. Description ''Didynamipus sjostedti'' are very small toads: males grow to and females to in snout–vent length. The snout is sharp in males but rounded in females; males also have a ridge on the top the snout that gives it an upturned appearance. The tympanum is absent. The hands and feet are reduced, with the two outer fingers and toes and the innermost toes present only as tubercles. No webbing is present. The dorsum is brown with lighter and darker mottling. Habitat and conservation ''Didynamipus sjostedti'' occur on forest edges and in clearings in moist forest at elevations of ...
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Rhaebo
''Rhaebo'' is a genus of true toads, family Bufonidae, from Central and South America. They are distributed from Honduras to northern South America including the Amazonian lowlands. Common name Cope toads has been suggested for them. Taxonomy The genus was removed from the synonymy of ''Bufo'' in 2006; an alternative view has been to treat it as a subgenus of ''Bufo''. At present, it is widely recognized as a genus. ''Andinophryne'', consisting of three species, was recognized as a separate genus until 2015 when it was found out that its recognition rendered ''Rhaebo'' paraphyletic. An alternative to synonymizing it with ''Rhaebo'' would have been to erect a new genus for ''Rhaebo nasicus'', but this would have caused difficulty in assigning species without molecular data to correct genus. Description ''Rhaebo'' are characterized as lacking cephalic crests, having omosternum, distinctively wide sphenethmoid, prominent and notched exoccipital condyles, and yellowish-orange skin ...
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