Stemonitis ''Stemonitis'' is a distinctive genus of slime moulds found throughout the world (except Antarctica). They are characterised by the tall brown sporangia, supported on slender stalks, which grow in clusters on rotting wood. The genus was first des ...

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...

Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Lecanoromycetidae and

Ostropomycetidae The Ostropomycetidae are a subclass of mostly lichen-forming fungi in the class Lecanoromycetes. The subclass was circumscribed in 2004 by Catherine Reeb, François M. Lutzoni, and Claude Roux. It contains ten orders and 36 families. Arctomiace ...

) is published by Huang ''et al.'' (2019). * Description of

crustose lichen Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichen ...

s from European

Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...

amber is published by Kaasalainen ''et al.'' (2019). * Fungi belonging to the genera '' Periconia'', ''

Penicillium ''Penicillium'' () is a genus of ascomycetous fungi that is part of the mycobiome of many species and is of major importance in the natural environment, in food spoilage, and in food and drug production. Some members of the genus produce pe ...

'' and ''

Scopulariopsis ''Scopulariopsis'' is a genus of teleomorph, anamorph and holomorph, anamorphic fungi that are saprobic and pathogenic to animals. The widespread genus contains 22 species. ''Scopulariopsis'' belongs to the group Hyphomycetes. These species are c ...

'', representing the first and the oldest known fossil record of these taxa, are described from the

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...

by Tischer ''et al.'' (2019).

Sponges

Research

*

Sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...

spicules and spicule-like structures that probably represent sponge fossils are described from four sections of the Ediacaran-Cambrian boundary interval in the Yangtze Gorges (China) by Chang ''et al.'' (2019). * A study evaluating how distribution patterns of non-lithistid spiculate sponges changed during the

Cambrian explosion The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil recor ...

and the

Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced wi ...

is published by Botting & Muir (2019).

New taxa

Cnidarians

Research

* A study on the growth characteristics of three species of

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...

corals belonging to the genus '' Agetolites'' from the Xiazhen Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), and on their implications for inferring phylogenetic relationships of this genus, is published by Sun, Elias & Lee (2019). * A study on a large colonial

rugose Rugose means "wrinkled". It may refer to: * Rugosa, an extinct order of coral, whose rugose shape earned it the name * Rugose, adjectival form of rugae Species with "rugose" in their names * '' Idiosoma nigrum'', more commonly, a black rugose tr ...

coral from the Ordovician Kope Formation (

Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) is published by Harris ''et al.'' (2019). * A study on the

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...

, growth characteristics and phylogenetic relationships of the

Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...

tabulate coral Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to ...

''

Halysites ''Halysites'' (meaning ''chain coral'') is an extinct genus of tabulate coral. Colonies range from less than one to tens of centimeters in diameter, and they fed upon plankton. These tabulate corals lived from the Ordovician to the Devonian (f ...

catenularius'' is published by Liang, Elias & Lee (2019). * Fossils of tabulate corals without septa, representing the first evidence that unmetamorphosed, slightly indurated

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

sandstones crop out amidst the deposits of the Atlantic Coastal Plain Province of the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

, are reported from

South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...

by Landmeyer ''et al.'' (2019). This finding is strongly disputed because all other rocks of Paleozoic age in the study area are greatly metamorphosed, the rocks where the fossils were found are traditionally mapped as the Cretaceous Middendorf Formation, and it is suggested that the fossils in question are the bark of Cretaceous conifers in Cretaceous sandstone, instead of Paleozoic corals in Paleozoic sandstone. * A study aiming to determine whether ecological selection based on physiology, behavior, habitat, etc. played a role in the long-term survival of corals during the late

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...

and early

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

is published by Weiss & Martindale (2019). * Fossils of ''

Acropora prolifera ''Acropora prolifera'', the fused staghorn coral, is a branching, colonial, stony coral found in shallow parts of the Caribbean Sea, the Bahamas and southern Florida. Description ''Acropora prolifera'' is very similar to staghorn coral (''Acr ...

'' dating back to the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

are reported by Precht ''et al.'' (2019). * A study on the distribution of reef corals during the last interglacial is published by Jones ''et al.'' (2019), who also evaluate the utility of fossil reef coral data for predictions of impact of future climate changes on reef corals. * A study on a problematic fossil specimen from the

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

Ponta Grossa Formation (

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

), assigned by different authors to the species '' Serpulites sica'' or '' Euzebiola clarkei'', is published by Van Iten ''et al.'' (2019), who interpret this fossil as a

medusozoa Medusozoa is a clade in the phylum Cnidaria, and is often considered a subphylum. It includes the classes Hydrozoa, Scyphozoa, Staurozoa and Cubozoa, and possibly the parasitic Polypodiozoa. Medusozoans are distinguished by having a medusa stage ...

n capable of clonal

budding Budding or blastogenesis is a type of asexual reproduction in which a new organism develops from an outgrowth or bud due to cell division at one particular site. For example, the small bulb-like projection coming out from the yeast cell is know ...

, and transfer it to the genus ''

Sphenothallus ''Sphenothallus'' is a problematic extinct genus lately attributed to the conulariids. It was widespread in shallow marine environments during the Paleozoic. Occurrence ''Sphenothallus'' is represented in the Cambrian period in the Kaili biot ...

''. *The oldest mesophotic coral ecosystems, dating back to middle Silurian, from the Lower Visby Beds on Gotland have been described by Zapalski & Berkowski. These communities, dominated by platy corals give also clues about the onset of coral-algal symbiosis. * Mihaljević (2019) describes new fossil coral collections from the

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

and

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

of

Sarawak Sarawak (; ) is a States and federal territories of Malaysia, state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the M ...

(

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...

),

Negros Island Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...

and

Cebu Cebu (; ceb, Sugbo), officially the Province of Cebu ( ceb, Lalawigan sa Sugbo; tl, Lalawigan ng Cebu; hil, Kapuroan sang Sugbo), is a province of the Philippines located in the Central Visayas region, and consists of a main island and 167 ...

(the

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

). * A study on the anatomy,

ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...

and taxonomy of the

Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...

hydrozoa Hydrozoa (hydrozoans; ) are a taxonomic class of individually very small, predatory animals, some solitary and some colonial, most of which inhabit saline water. The colonies of the colonial species can be large, and in some cases the specializ ...

n '' Heterastridium'', based on data from fossil specimens from central

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

and south

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...

, is published by Senowbari-Daryan & Link (2019).

New taxa

Arthropods

Bryozoans

Brachiopods

Molluscs

Echinoderms

Research

* A study on the

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...

and phylogenetic relationships of the putative

stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...

-echinoderm '' Yanjiahella biscarpa'' is published by Topper ''et al.'' (2019); the study is subsequently criticized by Zamora ''et al.'' (2020). * Soft tissue traces found in conjunction with skeletal molds are described in

stylophora The stylophorans are an extinct, possibly polyphyletic group allied to the Paleozoic Era echinoderms, comprising the prehistoric cornutes and mitrates. It is synonymous with the subphylum Calcichordata. Their unusual appearances have led to a va ...

ns by Lefebvre ''et al.'' (2019), who interpret their findings as supporting echinoderm and not

hemichordate Hemichordata is a phylum which consists of triploblastic, enterocoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and include ...

-like affinities of stylophorans. * A study on the morphology and phylogenetic relationships of the lepidocystoid echinoderm '' Vyscystis'' is published by Nohejlová ''et al.'' (2019). * A study on the phylogenetic relationships of

diploporita Diploporita is an extinct class of blastozoan that ranged from the Ordovician to the Devonian. These echinoderms are identified by a specialized respiratory structure, called diplopores. Diplopores are a double pore system that sit within a depr ...

n

blastozoa Blastozoa is a subphylum of extinct animals belonging to Phylum Echinodermata. This subphylum is characterized by the presence of specialized respiratory structures and brachiole plates used for feeding. This subphylum ranged from the Cambrian to ...

ns is published by Sheffield & Sumrall (2019). * A study on the morphology of the feeding

ambulacral Ambulacral is a term typically used in the context of anatomical parts of the phylum Echinodermata or class Sea star, Asteroidea and Edrioasteroidea. Echinoderms can have ambulacral parts that include Ossicle (echinoderm), ossicles, plates, spines ...

system in the

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...

diploporitan '' Eumorphocystis'', as indicated by data from well-preserved specimens from the

Bromide Formation The Bromide Formation is a geological formation in Oklahoma, USA. It is well known for its diverse echinoderm and trilobite fossil fauna. Location The Bromide Formation crops out in the Arbuckle and Wichita Mountains and in the Criner Hills ...

(

Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

), is published by Sheffield & Sumrall (2019), who interpret their findings as indicating that ''Eumorphocystis'' was closely related to

crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are ...

s and that crinoids are nested within blastozoans; their conclusions about the relationship between ''Eumorphocystis'' and crinoids are subsequently contested by Guensburg ''et al.'' (2020). * A study on the morphology and phylogenetic relationships of '' Macurdablastus uniplicatus'' is published by Bauer, Waters & Sumrall (2019). * A study on the morphology and phylogenetic relationships of '' Hexedriocystis'' is published online by Zamora & Sumrall (2019), who consider this taxon to be a blastozoan. * A study on the paleoecology of the specimens of the

edrioasteroid Edrioasteroidea is an extinct class (biology), class of echinoderms. The living animal would have resembled a pentamerously symmetrical disc or cushion. They were obligate encrusters and attached themselves to inorganic or biologic hard substrate ...

'' Neoisorophusella lanei'' preserved in limestone slabs from the

Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...

(

Chesterian The Mississippian ( , also known as Lower Carboniferous or Early Carboniferous) is a subperiod in the geologic timescale or a subsystem of the geologic record. It is the earlier of two subperiods of the Carboniferous period lasting from roughly ...

) Kinkaid Formation (

Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolita ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) is published by Shroat-Lewis, Greenwood & Sumrall (2019). * A study on the morphology of '' Cupulocrinus'' and on its implications for inferring the origin of the flexible crinoids is published by Peter (2019). * A study on the phylogenetic relationships of diplobathrid crinoids is published by Cole (2019). * A study on the biological and ecological controls on duration of diplobathrid crinoid genera is published online by Cole (2019). * A study on the macro-evolutionary patterns of body-size trends of cyrtocrinid crinoids is published by Brom (2019). * A study on patterns of paleocommunity structure and niche partitioning in crinoids from the

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...

(

Katian The Katian is the second stage of the Upper Ordovician. It is preceded by the Sandbian and succeeded by the Hirnantian Stage. The Katian began million years ago and lasted for about 7.8 million years until the beginning of the Hirnantian millio ...

) Brechin

Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...

(

Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) is published by Cole, Wright & Ausich (2019). * A study on the anatomy of the nervous and circulatory systems of the Cretaceous crinoid '' Decameros ricordeanus'' and on the phylogenetic relationships of this species is published online by Saulsbury & Zamora (2019). * A study on the substrate preference in

stem group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...

sea urchin Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...

s during the

Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...

Period will be published by Thompson & Bottjer (2019). * A study on

Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a un ...

recovery of sea urchins after the

Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as ...

is published by Pietsch ''et al.'' (2019). * A fossil

brittle star Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to starfish. They crawl across the sea floor using their flexible arms for locomo ...

belonging to the genus '' Ophiopetra'', representing the first record of articulated brittle star from the Mesozoic of South America reported so far, is described from the

Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...

Agua de la Mula Member of the

Agrio Formation The Agrio Formation is an Early Cretaceous geologic formation that is up to thick and is located in the southern Mendoza Province and northern-central Neuquén Province, in the Neuquén Basin of northwestern Patagonia, Argentina.Weaver, C. E. (1 ...

(

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

) by Fernández ''et al.'' (2019), who transfer the genus ''Ophiopetra'' to the family

Ophionereididae Ophionereididae are a family of brittle stars. Systematics Ophionereididae has been placed (along with Ophiocomidae) to the Taxonomic rank, superfamily Ophiocomidea and infraorder Chilophiurina or suborder Chilophiurina in different classificati ...

within the order

Amphilepidida Amphilepidida is an order of echinoderms belonging to the class Ophiuroidea Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to ...

.

New taxa

Conodonts

Research

* A study on the feeding habits of conodonts, as indicated by data from calcium stable isotopes, is published by Balter ''et al.'' (2019). * A study on the variation of conodont element crystal structure throughout their evolutionary history is published online by Medici ''et al.'' (2019). * A study on the evolution of platform-like P₁ elements in conodonts, evaluating its possible link to ecology of conodonts, is published by Ginot & Goudemand (2019). * A study on the impact of early

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

environmental changes on evolution and paleoecology of conodonts from the Canadian part of

Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although ...

is published online by Barnes (2019). * A study on the morphology, occurrences and biostratigraphical value of '' Paroistodus horridus'' is published online by Mestre & Heredia (2019). * A revision of the taxonomy and evolutionary relationships of the Late Ordovician genera '' Tasmanognathus'' and '' Yaoxianognathus'' is published by Yang ''et al.'' (2019). * A study on the composition and architecture of the apparatus of '' Erismodus quadridactylus'' is published by Dhanda ''et al.'' (2019). * A study on the

ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...

of the

Lochkovian The Lochkovian is one of three faunal stages in the Early Devonian Epoch. It lasted from 419.2 ± 3.2 million years ago to 410.8 ± 2.8 million years ago. It marked the beginning of the Devonian Period, and was followed by the Pragian Stage. It i ...

conodont species '' Ancyrodelloides carlsi'' is published by Corriga & Corradini (2019). * A study on fossils of members of the genus '' Alternognathus'' from the Upper

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

of the Kowala quarry (central

Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...

), attempting to calibrate the course of their

ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...

in days and documenting cyclic mortality events, is published by Świś (2019). * The apparatus of '' Vogelgnathus simplicatus'' is reconstructed from discrete elements from a sample of limited diversity from the

Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...

strata from

Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...

by Sanz-López, Blanco-Ferrera & Miller (2019). * Neospathodid conodont elements with partly preserved basal body (one of two main parts of conodont elements, besides the crown) are reported from the Lower Triassic of

Oman Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on the southeastern coast of the Arabian Peninsula, and spans the mouth of t ...

by Souquet & Goudemand (2019), who interpret their finding as indicating that the absence of basal bodies in post-

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

conodonts was due to a preservational bias only. * Natural assemblages of conodonts, preserving possible impressions of "eyes", are described from the Lower Triassic

pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...

black

claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone, slate, and shale. Most of the particles of which the stone is composed are less than and are too sm ...

s of the North Kitakami Belt (

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

) by Takahashi, Yamakita & Suzuki (2019). * A study on the composition of the apparatus of '' Nicoraella'', based on data from clusters from the Middle Triassic Luoping Biota (

Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...

,

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), is published by Huang ''et al.'' (2019). * The architecture of apparatus of ''Nicoraella kockeli'' is reconstructed by Huang ''et al.'' (2019), who also evaluate proposed functional interpretations of the conodont feeding apparatus. * A study on Middle Triassic conodont assemblages from Jenzig section of the

Jena Formation The Jena Formation is a geologic formation in Germany. It preserves fossils dating back to the Triassic period. See also * List of fossiliferous stratigraphic units in Germany See also * Lists of fossiliferous stratigraphic units in Europ ...

and Troistedt section of the Meissner Formation (

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

) is published by Chen ''et al.'' (2019), who also study the morphology of the apparatuses of ''

Neogondolella ''Neogondolellla'' is an extinct genus of conodonts Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were ...

haslachensis'' and ''Nicoraella germanica'', and review and revise the species ''Neogondolella mombergensis''. * A study evaluating the quantitative morphological variation of P₁ conodont elements within and between seven conodont

morphospecies In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

from the Pizzo Mondello section (

Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...

,

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

) and their evolution within 7 million years around the

Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...

/

Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...

boundary is published by Guenser ''et al.'' (2019). * A study on the

taphonomy Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientist Ivan Efremov t ...

of basal tissue of conodont elements is published online by Suttner & Kido (2019).

New taxa

Fishes

Amphibians

Reptiles

Synapsids

Non-mammalian synapsids

Research

* A study on the morphological diversity and morphological changes of the

humeri The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a round ...

of

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

and

Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...

synapsids through time is published by Lungmus & Angielczyk (2019). * A study on the diversity of patterns of skull shape (focusing on the relative lengths of the face and braincase regions of the skull) in non-mammalian synapsids is published by Krone, Kammerer & Angielczyk (2019). * Two pathologically fused tail vertebrae of a

varanopid Varanopidae is an extinct family of amniotes that resembled monitor lizards and may have filled a similar niche, hence the name. Typically, they are considered synapsids that evolved from an ''Archaeothyris''-like synapsid in the Late Carbonifero ...

, likely affected by a metabolic bone disease closely resembling

Paget's disease of bone Paget's disease of bone (commonly known as Paget's disease or, historically, osteitis deformans) is a condition involving cellular remodeling and deformity of one or more bones. The affected bones show signs of dysregulated bone remodeling at the ...

, are described from the early

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...

Richards Spur locality (

Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) by Haridy ''et al.'' (2019). * Description of new skull remains of ''

Echinerpeton ''Echinerpeton'' is an extinct genus of synapsid, including the single species ''Echinerpeton intermedium'' from the Late Carboniferous of Nova Scotia, Canada. The name means 'spiny lizard' (Greek). Along with its contemporary ''Archaeothyris'', ...

intermedium'' and a study on the phylogenetic relationships of this species is published online by Mann & Paterson (2019). * Fossil material of a large carnivorous synapsid belonging to the family

Sphenacodontidae Sphenacodontidae (Greek: "wedge point tooth family") is an extinct family of small to large, advanced, carnivorous, Late Pennsylvanian to middle Permian pelycosaurs. The most recent one, ''Dimetrodon angelensis'', is from the late Kungurian or e ...

is described from the Torre del Porticciolo locality (

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

) by Romano ''et al.'' (2019), representing the first carnivorous non-

therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...

synapsid Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptil ...

from the

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...

of Italy reported so far, and one of the few known from Europe. * Description of the

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...

and

histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...

of a small

neural spine The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...

from the Early

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...

Richards Spur locality (

Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) attributable to ''

Dimetrodon ''Dimetrodon'' ( or ,) meaning "two measures of teeth,” is an extinct genus of non-mammalian synapsid that lived during the Cisuralian (Early Permian), around 295–272 million years ago (Mya). It is a member of the family Sphenacodontid ...

'' is published by Brink, MacDougall & Reisz (2019), who also report evidence from fossil teeth indicative of presence of a derived species of ''Dimetrodon'' (otherwise typical of later,

Kungurian In the geologic timescale, the Kungurian is an age or stage of the Permian. It is the latest or upper of four subdivisions of the Cisuralian Epoch or Series. The Kungurian lasted between and million years ago (Ma). It was preceded by the Artin ...

localities of

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

and Oklahoma) at the Richards Spur locality. * A study on the

histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...

of the

skull roof The skull roof, or the roofing bones of the skull, are a set of bones covering the brain, eyes and nostrils in bony fishes and all land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparati ...

of burnetiamorph

biarmosuchia Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelyc ...

ns is published by Kulik & Sidor (2019). *

Femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...

of a specimen of the

titanosuchid Titanosuchidae is an extinct family of dinocephalians. The titanosuchids were carnivorous to omnivorous (herbivorous?) tapinocephalians. As with other tapinocephalians, they had thick skulls probably for head-butting. They had large canine tee ...

species ''

Jonkeria ''Jonkeria'' is an extinct genus of dinocephalians. Species were very large and omnivorous (although there is some dispute to this, e.g. Colbert 1969 p. 136), from the Tapinocephalus Assemblage Zone, Lower Beaufort Group, of the South Afri ...

parva'' affected by

osteomyelitis Osteomyelitis (OM) is an infection of bone. Symptoms may include pain in a specific bone with overlying redness, fever, and weakness. The long bones of the arms and legs are most commonly involved in children e.g. the femur and humerus, while the ...

is described from the

Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...

of

Karoo Basin The Karoo Supergroup is the most widespread stratigraphy, stratigraphic unit in Africa south of the Kalahari Desert. The group (stratigraphy), supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Pennsylvan ...

(

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

) by Shelton, Chinsamy & Rothschild (2019). * A study on the adaptations to herbivory in the teeth of members of the family

Tapinocephalidae Tapinocephalidae was an advanced family of tapinocephalians. It is defined as the clade containing ''Ulemosaurus'', '' Tapinocaninus'', and the Tapinocephalinae. They are known from both Russia and South Africa. In all probability, the Tapinoceph ...

is published by Whitney & Sidor (2019). * An almost complete skeleton of '' Tapinocaninus pamelae'', providing new information on the anatomy of the appendicular skeleton of this species (including the first accurate vertebral count for a

dinocephalia Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinoceph ...

n), is described from the lowermost

Beaufort Group The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlie ...

of

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

by Rubidge, Govender & Romano (2019). * Romano & Rubidge (2019) present body mass estimates for a well preserved and complete skeleton of ''Tapinocaninus pamelae'' from the lowermost Beaufort Group of South Africa. * A study on the skull anatomy and phylogenetic relationships of ''

Styracocephalus ''Styracocephalus platyrhynchus'' ('spike head') is an extinct species of tapinocephalian therapsids that lived during the Guadalupian epoch. ''Styracocephalus''s head ornament meant that it could be recognised from a distance. The most striki ...

platyrhynchus'' is published by Fraser-King ''et al.'' (2019). * A study on the evolution of the sacral vertebrae of

dicynodont Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typicall ...

s is published by Griffin & Angielczyk (2019). * A study on the diversity of dicynodonts from the Upper Permian Naobaogou Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) is published by Liu (2019). * A study on skulls of South American dicynodonts, aiming to determine whether the differences in skull morphology were related to differences in feeding function, is published by Ordonez ''et al.'' (2019). * New fossil material of ''

Endothiodon ''Endothiodon'' (/ɛndoʊθiːoʊdɔːn/ "inner tooth" from Greek endothi (ἔνδοθῐ), "within", and odon (ὀδών), "tooth", most likely named for the characteristic of the teeth being placed internally to the maxillaBoos A. S., Schultz C ...

tolani'' is described from the Permian K5 Formation of the Metangula Graben (

Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...

) by Macungo ''et al.'' (2019). * A study on the anatomy of the postcranial skeleton of ''Endothiodon bathystoma'', based on data from a new specimen from the uppermost ''Pristerognathus'' Assemblage Zone of the

Karoo Supergroup The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a perio ...

(

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

), is published online by Maharaj, Chinsamy & Smith (2019). * Small dicynodont skull assigned to the genus ''

Digalodon ''Digalodon'' is an extinct genus of kistecephalian dicynodont. For a time it was considered synonymous with other dicynodonts as a small juvenile of a larger genus, such as ''Aulacephalodon'' or ''Dicynodontoides'', however it has since been re ...

'' is described from the

Lopingian The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic. The Lopingian is often synonymous with the informal terms l ...

upper Madumabisa Mudstone Formation (

Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most cent ...

) by Angielczyk (2019), expanding known geographic range of this genus. * Digital

endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...

of ''

Rastodon ''Rastodon'' is an extinct genus of anomodonts. It is the oldest and most basal known genus of bidentalian dicynodonts. Uniquely among dicynodonts, its tusks curve forward. The type and only species is ''R. procurvidens''. Etymology ''Rasto ...

procurvidens'' is reconstructed by de Simão-Oliveira, Kerber & Pinheiro (2019), who evaluate biological implications of the endocast morphology of this species. * Mancuso & Irmis (2019) describe an

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

of a member of the genus ''

Stahleckeria ''Stahleckeria'' is an extinct genus of Middle Triassic (Ladinian) dicynodonts.Chañares Formation The Chañares Formation is a Carnian-age geologic formation of the Ischigualasto-Villa Unión Basin, located in La Rioja Province, Argentina. It is characterized by drab-colored fine-grained volcaniclastic claystones, siltstones, and sandstones ...

(

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

), and evaluate the implications of this finding for the knowledge of the Triassic

Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...

n

biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock Stratum, strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictiona ...

and

biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...

. * A study on the body mass of ''

Lisowicia ''Lisowicia'' is an extinct genus of giant dicynodont synapsid that lived in what is now Poland during the late Norian or earliest Rhaetian age of the Late Triassic Period, about 210–205 million years ago. ''Lisowicia'' is the largest known d ...

bojani'' is published online by Romano & Manucci (2019). * A study on fossils of a putative Cretaceous dicynodont from

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

reported by

Thulborn Thulborn is an Anglo-Saxon surname dating back to the late 12th century AD, derived from the baptismal name for the "son of Thurburn". Notable people with the surname include: *Richard A. Thulborn, British paleontologist *Scott Thulborn (born 1984) ...

& Turner (2003) is published online by Knutsen & Oerlemans (2019), who consider these fossils to be of

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

age, and reinterpret it as fossils of a large mammal, probably a

diprotodontid The Diprotodontidae are an extinct family of large herbivorous marsupials, endemic to Australia and New Guinea during the Oligocene through Pleistocene periods from 28.4 million to 40,000 years ago. The family consisted of large quadrupedal te ...

. * A study aiming to determine patterns of morphological and phylogenetic diversity of

therocephalia Therocephalia is an extinct suborder of eutheriodont therapsids (mammals and their close relatives) from the Permian and Triassic. The therocephalians ("beast-heads") are named after their large skulls, which, along with the structure of their te ...

ns throughout their evolutionary history is published by Grunert, Brocklehurst & Fröbisch (2019). * A study on variation in rates of body size evolution of therocephalians is published by Brocklehurst (2019). * A study on the morphology of the manus of a new therocephalian specimen referable to the genus ''

Tetracynodon ''Tetracynodon'' is an extinct genus of therocephalian. Fossils of ''Tetracynodon'' have been found in the Karoo Basin of South Africa. Two species are known: the type species ''T. tenuis'' from the Late Permian and the species ''T. darti'' fro ...

'' from the

Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a un ...

of

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

, and on the evolution of the manus morphology of therocephalians, is published by Fontanarrosa ''et al.'' (2019). * A study on patterns of nonmammalian

cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...

species richness and the quality of their fossil record is published by Lukic-Walther ''et al.'' (2019). * A study on the morphology and bone

histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...

of the postcranial skeleton of ''

Galesaurus ''Galesaurus'' (from the Greek roots for 'weasel' and 'lizard') is an extinct genus of carnivorous cynodont therapsid that lived between the Induan and the Olenekian stages of the Early Triassic in what is now South Africa. It was incorrectly cla ...

planiceps'' is published by Butler, Abdala & Botha-Brink (2019). * Redescription of the anatomy of the skull of ''Galesaurus planiceps'' is published by Pusch, Kammerer & Fröbisch (2019). * Description of teeth of all known

diademodontid Diademodontidae is an extinct family of Triassic gomphodonts. The best-known genus is ''Diademodon'' from South Africa. ''Titanogomphodon'' from Namibia may also be a member of Diademodontidae. The Chinese genera ''Hazhenia'' and ''Ordosiodon'' ha ...

and trirachodontid cynodont taxa is published by Hendrickx, Abdala & Choiniere (2019), who also propose a standardized list of anatomical terms and abbreviations in the study of gomphodont teeth, assign '' Sinognathus'' and ''

Beishanodon ''Beishanodon'' is an extinct genus of eucynodonts from Lower Triassic of China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most p ...

'' to the family Trirachodontidae, and consider all specimens previously referred to the species ''

Cricodon ''Cricodon'' is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa.Abdala, F., J. Neveling, and J. Welman. 2006. A new trirachodontid cynodont from the lower levels of the Burg ...

kannemeyeri'' to be younger individuals of '' Trirachodon berryi''. * A study on the bone

histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...

of the

traversodontid Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Tra ...

cynodonts '' Protuberum cabralense'' and ''

Exaeretodon ''Exaeretodon'' is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. ''E. argentinus'' is from the Carnian-age (Late Triassic) Cancha de Boch ...

riograndesis'' is published by Veiga, Botha-Brink & Soares (2019). *

Hypsodont Hypsodont is a pattern of dentition with high-crowned teeth and enamel extending past the gum line, providing extra material for wear and tear. Some examples of animals with hypsodont dentition are cows and horses; all animals that feed on gritt ...

postcanine teeth of '' Menadon besairiei'' are described by Melo ''et al.'' (2019), who also study patterns of dental growth and replacement in this species. * Digital

endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...

s of '' Massetognathus ochagaviae'' and '' Probelesodon kitchingi'' are reconstructed by Hoffmann ''et al.'' (2019). * A skull of a member of the species '' Massetognathus ochagaviae'' is described from the

Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...

''Santacruzodon'' Assemblage Zone of the Santa Maria Supersequence (

Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...

,

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

) by Schmitt ''et al.'' (2019). * Description of brain endocasts of '' Siriusgnathus niemeyerorum'' and ''

Exaeretodon ''Exaeretodon'' is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. ''E. argentinus'' is from the Carnian-age (Late Triassic) Cancha de Boch ...

riograndensis'', using virtual models based on computed tomography scan data, is published by Pavanatto, Kerber & Dias-da-Silva (2019). * Description of new fossil material of '' Siriusgnathus niemeyerorum'' from the Upper Triassic

Caturrita Formation The Caturrita Formation is a geological formation, rock formation found in Rio Grande do Sul, Brazil. Its sediments were deposited in the Paraná Basin. The formation is from the Upper Triassic and forms part of the Santa Maria Supersequence in th ...

(

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

) and a study on the age of its fossils is published online by Miron ''et al.'' (2019). * A study on the evolution of infraorbital

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

ry canal in

probainognathia Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most ...

n cynodonts and on its implications for the knowledge of evolution of mobile whiskers in non-mammalian synapsids, as indicated by data from skulls of non-mammalian probainognathian cynodonts and early

mammaliaforms Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recen ...

, is published online by Benoit ''et al.'' (2019). * Digital skull

endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...

of a specimen of ''

Riograndia ''Riograndia'' is an extinct genus of tritheledontid cynodonts from the Late Triassic of South America. The type and only species is ''Riograndia guaibensis''. Remains have been found in the Caturrita Formation of the geopark of Paleorrota. I ...

guaibensis'' is reconstructed by Rodrigues ''et al.'' (2019). * Description of the anatomy of the first postcranial specimens referable to ''Riograndia guaibensis'' is published by Guignard, Martinelli & Soares (2019). * A study on the anatomy of the postcranial skeleton of ''

Brasilodon ''Brasilodon'' ("tooth from Brazil") is an extinct genus of small, mammal-like cynodonts that lived in what is now Brazil during the Norian age of the Late Triassic epoch, about 225.42 million years ago. While no complete skeletons have been foun ...

quadrangularis'' is published by Guignard, Martinelli & Soares (2019). * A study on tooth wear patterns of members of the family

Tritylodontidae Tritylodontidae ("three-knob teeth", named after the shape of their cheek teeth) is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, bearing several mammalian traits like erect limbs, endothermy and detail ...

and on their possible diet is published by Kalthoff ''et al.'' (2019). * Possible cynodont teeth, which might be the most recent non-

mammaliaform Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent com ...

cynodont fossils from Africa reported so far, are described from the

Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...

or earliest

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

locality of Ksar Metlili (Anoual Syncline, eastern

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

) by Lasseron (2019). * A study on the origin of the mammalian

middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in the ...

ossicles The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditory ...

, as indicated by the anatomy of the jaw-otic complex in 43 synapsid taxa, is published by Navarro-Díaz, Esteve-Altava & Rasskin-Gutman (2019). * A study on the evolution of the morphological complexity of the mammalian vertebral column, as indicated by data from mammals and non-mammalian synapsids, is published by Jones, Angielczyk & Pierce (2019).

New taxa

Mammals

Other animals

New taxa

Research

* A study on moulds of animals belonging to the group

Proarticulata Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. pr ...

from the southeastern

White Sea The White Sea (russian: Белое море, ''Béloye móre''; Karelian and fi, Vienanmeri, lit. Dvina Sea; yrk, Сэрако ямʼ, ''Serako yam'') is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is su ...

area (

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

), and on their implications for the knowledge of the morphology of integuments of members of Proarticulata, is published by Ivantsov, Zakrevskaya & Nagovitsyn (2019). * A study on accumulations of ''

Ernietta ''Ernietta'' is an extinct genus of Ediacaran organisms with an infaunal lifestyle. Fossil preservations and modeling indicate this organism was sessile and “sack”-shaped. It survived partly buried in substrate, with an upturned bell-shaped ...

'' from the Witputs subbasin (

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

), and on their implications for the knowledge of ecology of these organisms, is published by Gibson ''et al.'' (2019). * A diverse assemblage of tubular fossils – dominated by typical

Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...

organisms such as '' Cloudina'' and ''

Sinotubulites ''Sinotubulites'' is a genus of small, tube-shaped shelly fossils from the Ediacaran period. It is often found in association with '' Cloudina''. Its tube has a "tube-in-tube" structure composed of several thin layers. It bears prominent longit ...

'', but also preserving fossils showing similarities to early

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

shelly fossils – is described from the Ediacaran Dengying Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) by Cai ''et al.'' (2019). * Letsch ''et al.'' (2019) report late Ediacaran discoidal Ediacara-type fossils and latest Ediacaran to early

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

microfossils A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, ...

from the Tabia and the Tifnout members of the Adoudou Formation (

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

), constituting the oldest known direct evidence for presumably animal life from Northwest Africa. * A study delineating different types of the

asexual reproduction Asexual reproduction is a type of reproduction that does not involve the fusion of gametes or change in the number of chromosomes. The offspring that arise by asexual reproduction from either unicellular or multicellular organisms inherit the fu ...

for '' Cloudina'' and '' Multiconotubus'' is published by Min ''et al.'' (2019). * A study on the anatomy of ''

Charnia ''Charnia'' is a genus of frond-like lifeforms belonging to the Ediacaran biota with segmented, leaf-like ridges branching alternately to the right and left from a zig-zag medial suture (thus exhibiting glide reflection, or opposite isometry). T ...

masoni'' is published by Dunn ''et al.'' (2019). * A study evaluating whether ''

Dickinsonia ''Dickinsonia'' is an extinct genus of basal animal that lived during the late Ediacaran period in what is now Australia, China, Russia and Ukraine. The individual ''Dickinsonia'' typically resembles a bilaterally symmetrical ribbed oval. Its a ...

'' was capable of mobility is published by Evans, Gehling & Droser (2019). * A study comparing the biomechanical responses of tissues of ''Dickinsonia'' to various forces with those typical of modern organisms is published by Evans ''et al.'' (2019). * A study on the anatomy, growth and phylogenetic relationships of '' Arborea arborea'' is published by Dunn, Liu & Gehling (2019). * Xiao ''et al.'' (2019) describe a new

trace fossil A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...

from the Ediacaran Dengying Formation (China), interpreted as produced by a

bilateria The Bilateria or bilaterians are animals with bilateral symmetry as an embryo, i.e. having a left and a right side that are mirror images of each other. This also means they have a head and a tail (anterior-posterior axis) as well as a belly and ...

n animal exploring an oxygen oasis in

microbial mat A microbial mat is a multi-layered sheet of microorganisms, mainly bacteria and archaea, or bacteria alone. Microbial mats grow at interfaces between different types of material, mostly on submerged or moist surfaces, but a few survive in deserts. ...

s, and name a new

ichnotaxon An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...

'' Yichnus levis''. * A study on fossil molds and casts from the

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...

of

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

and the

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

of

New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...

, as well as on Ediacaran mold and cast fossils from

South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...

, the

White Sea The White Sea (russian: Белое море, ''Béloye móre''; Karelian and fi, Vienanmeri, lit. Dvina Sea; yrk, Сэрако ямʼ, ''Serako yam'') is a southern inlet of the Barents Sea located on the northwest coast of Russia. It is su ...

region of

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

,

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

and

Newfoundland Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...

, is published by MacGabhann ''et al.'' (2019), who evaluate how faithfully the fossils represent the original organisms, and whether the first animals to evolve on Earth could have been fossilized in a way similar to eldoniids from the Tafilalt

Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...

of Morocco. * Exceptionally preserved

phosphatized Phosphatic fossilization has occurred in unusual circumstances to preserve some extremely high-resolution microfossils in which careful preparation can even reveal preserved cellular structures. Such microscopic fossils are only visible under the sc ...

archaeocyaths and

small shelly fossils The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Camb ...

are reported from the Lower Cambrian Salaagol Formation of southwestern

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

by Pruss ''et al.'' (2019). * A study on the timing of the development of

reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock out ...

biodiversity, based on data from microbial-archaeocyathan reefs of the Salaagol Formation in Mongolia and other early

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

reefs, is published by Cordie ''et al.'' (2019). * A study on the morphological diversity of archaeocyaths is published by Cordie & Dornbos (2019). * Evidence of extensive burrowing in laminated claystone from the

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

(

Drumian The Drumian is a stage of the Miaolingian Series of the Cambrian. It succeeds the Wuliuan and precedes the Guzhangian. The base is defined as the first appearance of the trilobite ''Ptychagnostus atavus'' around million years ago. The top is def ...

) Ravens Throat River

Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...

in the Rockslide Formation (

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) is presented by Pratt & Kimmig (2019). * Description of jaw apparatus of '' Plumulites bengtsoni'' from the

Fezouata Formation The Fezouata Formation or Fezouata Shale is a geological formation in Morocco which dates to the Early Ordovician.

of

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

, evaluating its implications for the knowledge of the phylogenetic relationships of machaeridians, is published by Parry ''et al.'' (2019). * Description of internal anatomical features of '' Canadia spinosa'' identified as remnants of the nervous system is published by Parry &

Caron A caron (), háček or haček (, or ; plural ''háčeks'' or ''háčky'') also known as a hachek, wedge, check, kvačica, strešica, mäkčeň, varnelė, inverted circumflex, inverted hat, flying bird, inverted chevron, is a diacritic mark (� ...

(2019). * A study on the chemical composition,

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...

and phylogeny of fossil (

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...

,

Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...

and

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

)

annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...

tubes and tubes formerly thought to have been made by annelids, recovered from

hydrothermal vent A hydrothermal vent is a fissure on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hotspot ...

and

cold seep A cold seep (sometimes called a cold vent) is an area of the ocean floor where hydrogen sulfide, methane and other hydrocarbon-rich fluid seepage occurs, often in the form of a brine pool. ''Cold'' does not mean that the temperature of the see ...

environments, is published by Georgieva ''et al.'' (2019). * A massive deposit composed of fossil

serpulid The Serpulidae are a family of sessile, tube-building annelid worms in the class Polychaeta. The members of this family differ from other sabellid tube worms in that they have a specialized operculum that blocks the entrance of their tubes wh ...

worm tubes dating to the late

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

is reported from the Santa Monica Basin off the coast of southern

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

by Georgieva ''et al.'' (2019). * A study on the microstructure of

hyolith Hyoliths are animals with small conical surface, conical animal shell, shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, w ...

conchs and opercula from the lower

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

Xinji Formation of North China, and on its implications for inferring the phylogenetic relationships of Hyolitha, is published by Li ''et al.'' (2019). * Description of soft parts associated with the feeding apparatus of the hyolith '' Triplicatella opimus'' from the

Chengjiang biota The Maotianshan Shales are a series of Early Cambrian deposits in the Chiungchussu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales ...

of South China, and a study on the implications of this finding for the knowledge of the phylogenetic affinities of hyoliths, is published online by Liu ''et al.'' (2019). * A study on changes of conch size in tentaculitoids from the

Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...

and

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

strata is published by Wei (2019). * A study on the anatomy of '' Amiskwia sagittiformis'' is published by Vinther & Parry (2019), who interpret two reflective patches present in fossils of this species, previously interpreted as paired cerebral ganglia, as a pair of pharyngeal jaws similar to those of gnathiferans. * A study on the anatomy and phylogenetic affinities of ''Amiskwia sagittiformis'' is published by

Caron A caron (), háček or haček (, or ; plural ''háčeks'' or ''háčky'') also known as a hachek, wedge, check, kvačica, strešica, mäkčeň, varnelė, inverted circumflex, inverted hat, flying bird, inverted chevron, is a diacritic mark (� ...

& Cheung (2019). * The oldest record of

acanthocephala Acanthocephala (Greek , ', thorn + , ', head) is a phylum of parasitic worms known as acanthocephalans, thorny-headed worms, or spiny-headed worms, characterized by the presence of an eversible proboscis, armed with spines, which it uses to p ...

n parasite eggs described so far is reported from probable

crocodyliform Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...

coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...

s from the Upper Cretaceous

Adamantina Formation The Adamantina Formation is a geological formation in the Bauru Basin of western São Paulo state, in southeastern Brazil. Its strata date back to the Late Cretaceous epoch of the Cretaceous Period, during the Mesozoic Era. The formation is p ...

(

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

) by Cardia ''et al.'' (2019). * Exceptionally preserved trace and body fossils are described from the

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

File Haidar Formation (

Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...

) by Kesidis ''et al.'' (2019), who interpret these fossils as made by

priapulid Priapulida (priapulid worms, from Gr. πριάπος, ''priāpos'' 'Priapus' + Lat. ''-ul-'', diminutive), sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility ...

-like

scalidophora Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuti ...

ns. * Description of

exuviae In biology, exuviae are the remains of an exoskeleton and related structures that are left after ecdysozoans (including insects, crustaceans and arachnids) have moulted. The exuviae of an animal can be important to biologists as they can often b ...

of microscopic

scalidophora Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuti ...

n worms from the lowermost Cambrian Kuanchuanpu Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) is published by Wang ''et al.'' (2019), who interpret this finding as the oldest record of

moulting In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...

in

ecdysozoa Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerata, crustaceans, and myriapods), Nematoda, and several smaller phyla. They were first defined by Aguinaldo ''et al.'' in 1997, based mainly on phylogenetic tr ...

ns reported so far. * A reassessment of

radiodonta Radiodonta is an extinct Order (biology), order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the ...

n fossils known from the

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

Kinzers Formation The Kinzers Formation is a geologic formation in Pennsylvania. It preserves fossils dating back to the Cambrian Period. The base of the Kinzers Formation is primarily a dark-brown shale. The middle is a gray and white spotted limestone and, lo ...

(

Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) is published by Pates & Daley (2019), who argue that at least four radiodontan taxa are known from this formation, and confirm that ''

Anomalocaris ''Anomalocaris'' ("unlike other shrimp", or "abnormal shrimp") is an extinct genus of Radiodonta, radiodont, an Order (biology), order of early-diverging stem-group arthropods. The first fossils of ''Anomalocaris'' were discovered in the ''Ogygop ...

pennsylvanica'' is a distinct species from ''A. canadensis''. * A study on the

moulting In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...

behaviour of the chengjiangocaridid fuxianhuiid '' Alacaris mirabilis'' is published by Yang ''et al.'' (2019). * A study on the anatomy and phylogenetic relationships of a

stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...

-arthropod '' Guangweicaris spinatus'' is published by Wu & Liu (2019). * A fossil interpreted as a partial mold of a specimen of ''

Paropsonema ''Paropsonema'' is a discoidal animal known from the Cambrian Chengjiang biota The Maotianshan Shales are a series of Early Cambrian deposits in the Chiungchussu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the ...

cryptophya'' is described from the Middle-Upper Devonian of New York by Hagadorn & Allmon (2019), representing the most recent occurrence of the paropsonemids reported so far. * A study evaluating the utility of eye

melanosome A melanosome is an organelle found in animal cells and is the site for synthesis, storage and transport of melanin, the most common light-absorbing pigment found in the animal kingdom. Melanosomes are responsible for color and photoprotection i ...

s for determination of the phylogenetic affinities of '' Tullimonstrum'' is published by Rogers ''et al.'' (2019).

Foraminifera

Research

* A study on the morphological complexity of

planktic Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...

foraminifer

tests Test(s), testing, or TEST may refer to: * Test (assessment), an educational assessment intended to measure the respondents' knowledge or other abilities Arts and entertainment * ''Test'' (2013 film), an American film * ''Test'' (2014 film), ...

after the

Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...

is published by Lowery & Fraass (2019). * A study on the response of the larger

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular Ectoplasm (cell biology), ectoplasm for catching food and ot ...

from the

Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

to the

Paleocene–Eocene Thermal Maximum The Paleocene–Eocene thermal maximum (PETM), alternatively (ETM1), and formerly known as the "Initial Eocene" or "", was a time period with a more than 5–8 °C global average temperature rise across the event. This climate event o ...

, based on fossil evidence from south

Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...

, is published by Zhang ''et al.'' (2019).

New taxa

Other organisms

New taxa

Research

* Putative traces of life older than 3.95 Ga, reported from northern

Labrador , nickname = "The Big Land" , etymology = , subdivision_type = Country , subdivision_name = Canada , subdivision_type1 = Province , subdivision_name1 ...

(

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) by Tashiro ''et al.'' (2017) are reevaluated by Whitehouse ''et al.'' (2019). * Description of cellularly preserved microfossils from ~3.4 Ga-old deposits of the Kromberg Formation (

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

), providing information on reproduction patterns of these organisms, is published by Kaźmierczak & Kremer (2019). * El Albani ''et al.'' (2019) describe 2.1 billion-year-old fossils belonging to the

Francevillian biota The Francevillian biota (also known as Gabon macrofossils or Gabonionta) is a group of 2.1-billion-year-old Palaeoproterozoic, macroscopic organisms known from fossils found in Gabon in the Palaeoproterozoic Francevillian B Formation, a black sha ...

of

Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...

, including pyritized string-shaped structures interpreted as produced by a

multicellular A multicellular organism is an organism that consists of more than one cell, in contrast to unicellular organism. All species of animals, land plants and most fungi are multicellular, as are many algae, whereas a few organisms are partially uni- ...

or

syncytial A syncytium (; plural syncytia; from Greek: σύν ''syn'' "together" and κύτος ''kytos'' "box, i.e. cell") or symplasm is a multinucleate cell which can result from multiple cell fusions of uninuclear cells (i.e., cells with a single nucleus) ...

organism able to migrate laterally and vertically to reach food resources. * A study on ca. 1.9 Ga hairpin-shaped trace fossils and discoid fossils from the Stirling Range Formation (

Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...

) is published by Retallack & Mao (2019), who interpret these fossils as evidence of early life on land. * A study on organic-walled microfossils from the Cailleach Head Formation (

Torridon Group In geology, the term Torridonian is the informal name for the Torridonian Group, a series of Mesoproterozoic to Neoproterozoic arenaceous and argillaceous sedimentary rocks, which occur extensively in the Northwest Highlands of Scotland. The s ...

,

Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...

) is published by Wacey ''et al.'' (2019), who report exceptional preservation of sub-cellular detail in selected cells. *

Phosphatized Phosphatic fossilization has occurred in unusual circumstances to preserve some extremely high-resolution microfossils in which careful preparation can even reveal preserved cellular structures. Such microscopic fossils are only visible under the sc ...

three-dimensional fossil of a putative

calcimicrobe Characteristic of the Neoproterozoic and Cambrian periods, the heterogeneous group called calcimicrobes are calcareous colonial microfossils, which include many morphologically dissimilar organisms, whose effect in massive aggregations, in associat ...

''Epiphyton'' are reported from the

Neoproterozoic The Neoproterozoic Era is the unit of geologic time from 1 billion to 538.8 million years ago. It is the last era of the Precambrian Supereon and the Proterozoic Eon; it is subdivided into the Tonian, Cryogenian, and Ediacaran periods. It is ...

Dengying Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) by Min ''et al.'' (2019). * A study on the affinities of tubular microfossils from the

Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...

Doushantuo Formation The Doushantuo Formation (formerly transcribed as Toushantuo or Toushantou, from ) is a geological formation in western Hubei, eastern Guizhou, southern Shaanxi, central Jiangxi, and other localities in China. It is known for the fossil Lagerst ...

(

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), i.e. '' Crassitubus'', '' Quadratitubus'', '' Ramitubus'' and '' Sinocyclocyclicus'', is published by Sun ''et al.'' (2019), who reject the interpretation of these taxa as early animals. * Lehn, Horodyski & Paim (2019) report the first known occurrence of Ediacaran organic-walled microfossils preserved in fine-grained

siliciclastic Siliciclastic (or ''siliclastic'') rocks are clastic noncarbonate sedimentary rocks that are composed primarily of silicate minerals, such as quartz or clay minerals. Siliciclasic rock types include mudrock, sandstone Sandstone is a clastic ...

strata of the Camaquã Basin (southernmost

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

). * A study on the structure, developmental biology and affinities of ''

Caveasphaera ''Caveasphaera'' is a multicellular organism found in 609-million-year-old rocks laid down during the Ediacaran period in the Guizhou Province of South China. The organism is not easily defined as an animal or non-animal. The organism is notable ...

costata'' from the Ediacaran

Doushantuo Formation The Doushantuo Formation (formerly transcribed as Toushantuo or Toushantou, from ) is a geological formation in western Hubei, eastern Guizhou, southern Shaanxi, central Jiangxi, and other localities in China. It is known for the fossil Lagerst ...

(

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) is published by Yin ''et al.'' (2019). * A study on possible cells and their appendages in fossils of ''Epiphyton'' from the

Wuliuan The Wuliuan stage is the fifth stage of the Cambrian, and the first stage of the Miaolingian Series of the Cambrian. It was formally defined by the ICS in 2018. Its base is defined by the first appearance of the trilobite species ''Oryctocephalu ...

of the North China Platform, and on their implications for the classification of this taxon, is published by Zhang ''et al.'' (2019). * A study on the

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...

and colony organization of '' Rhyniococcus uniformis'' (a

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

organism resembling extant

cyanobacteria Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, blu ...

in the genus ''

Merismopedia ''Merismopedia'' (from the Greek ''merismos'' ivisionand the Greek ''pedion'' lain is a genus of cyanobacteria found in fresh and salt water. It is ovoid or spherical in shape and arranged in rows and flats, forming rectangular colonies held t ...

''), based on data from new specimens, is published by Krings & Harper (2019). * A new method of assessing the morphology of fossil

radiolaria The Radiolaria, also called Radiozoa, are protozoa of diameter 0.1–0.2 mm that produce intricate mineral skeletons, typically with a central capsule dividing the cell (biology), cell into the inner and outer portions of endoplasm and Ecto ...

n specimens is presented by Kachovich, Sheng & Aitchison (2019), who apply their method to six specimens from the Cambrian Inca Formation (

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

) and Ordovician Piccadilly Formation (

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) and evaluate the implications of their method for the studies of radiolarian evolution.

Trace fossils

History of life in general

Research related to paleontology that concerns multiple groups of the organisms listed above. * Experiments indicating that abiotic

chemical garden Comparison of chemical gardens grown by NASA scientists on the International Space Station (left) and on the ground (right) A chemical garden while growing up Cobalt(II) chloride upA chemical garden A chemical garden is a set of complex biol ...

ing can mimic structures interpreted as the oldest known fossil microorganisms in both morphology and composition are conducted by McMahon (2019). * A study on

biomarker In biomedical contexts, a biomarker, or biological marker, is a measurable indicator of some biological state or condition. Biomarkers are often measured and evaluated using blood, urine, or soft tissues to examine normal biological processes, ...

s recovered from cap

dolomite Dolomite may refer to: *Dolomite (mineral), a carbonate mineral *Dolomite (rock), also known as dolostone, a sedimentary carbonate rock *Dolomite, Alabama, United States, an unincorporated community *Dolomite, California, United States, an unincor ...

s of the Araras Group (

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

), interpreted as evidence of the transition from a bacterial to

eukaryotic Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacte ...

dominated ecosystem after the

Marinoan The Marinoan glaciation, sometimes also known as the Varanger glaciation, was a period of worldwide glaciation that lasted from approximately 650 to 632.3 ± 5.9 Ma (million years ago) during the Cryogenian period. This glaciation possibly cove ...

deglaciation, likely caused by massive

bacterivorous A bacterivore is an organism which obtains energy and nutrients primarily or entirely from the consumption of bacteria. The term is most commonly used to describe free-living, heterotrophic, microscopic organisms such as nematodes as well as many s ...

grazing by

ciliate The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to flagellum, eukaryotic flagella, but are in general shorter and present in much larger numbers, with a ...

s, is published by van Maldegem ''et al.'' (2019). * Biomarkers thought to be diagnostic for

demosponge Demosponges (Demospongiae) are the most diverse class in the phylum Porifera. They include 76.2% of all species of sponges with nearly 8,800 species worldwide (World Porifera Database). They are sponges with a soft body that covers a har ...

s and cited as evidence of rise of animals to ecological importance prior to the

Cambrian radiation The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil recor ...

are reported to be also synthesized by

rhizaria The Rhizaria are an ill-defined but species-rich supergroup of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus Paulinella in the phylum Cercozoa, they are all non-photosynthethic, but many foramini ...

ns by Nettersheim ''et al.'' (2019), who place the oldest unambiguous evidence for animals closer to the Cambrian Explosion. * A study on crucial conditions affecting the evolution of a proto-metabolism in early life is published by Goldford ''et al.'' (2019). * A study on the age of the

Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...

fossils from the Podolya Basin (southwestern

Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...

) is published by Soldatenko ''et al.'' (2019). * A study on occurrences of body and trace fossils in

Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and th ...

and lower

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

(

Fortunian The Fortunian age marks the beginning of the Phanerozoic Eon, the Paleozoic Era, and the Cambrian Period. It is the first of the two stages of the Terreneuvian series. Its base is defined as the first appearance of the trace fossil ''Treptichnus p ...

) rocks around the world is published by Muscente ''et al.'' (2019), who report evidence indicative of existence of a global,

cosmopolitan Cosmopolitan may refer to: Food and drink * Cosmopolitan (cocktail), also known as a "Cosmo" History * Rootless cosmopolitan, a Soviet derogatory epithet during Joseph Stalin's anti-Semitic campaign of 1949–1953 Hotels and resorts * Cosmopoli ...

assemblage unique to terminal Ediacaran strata, living between two episodes of biotic turnover which might be the earliest mass extinctions of complex life. * A study on the diversification of animals and their behaviour in the Ediacaran–Cambrian interval, as indicated by fossil and environmental proxy records, is published by

Wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...

''et al.'' (2019), who interpret the fossil record as indicating that the rise of early animals was more likely a series of successive, transitional

radiation In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or through a material medium. This includes: * ''electromagnetic radiation'', such as radio waves, microwaves, infrared, visi ...

events which extended from the Ediacaran to the early

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

, rather than competitive or biotic replacement of the latest Ediacaran biotas by markedly distinct Cambrian ones. * A study comparing the variability of Ediacaran faunal assemblages to that of more recent fossil and modern

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

assemblages is published by Finnegan, Gehling & Droser (2019). * A study on the intensity of animal

bioturbation Bioturbation is defined as the reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains. Bioturbating activities have a profound effect on the environment and are thought to be a pr ...

and ecosystem engineering in trace fossil assemblages throughout the latest Ediacaran

Nama Group The Nama Group is a megaregional Vendian to Cambrian group of stratigraphic sequences deposited in the Nama foreland basin in central and southern Namibia. The Nama Basin is a peripheral foreland basin, and the Nama Group was deposited in two e ...

(

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

), evaluating the implications of this data for the knowledge of the causes of the disappearance of the

Ediacaran biota The Ediacaran (; formerly Vendian) biota is a Taxonomy (biology), taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (). These were composed of enigmatic tubular and frond-sh ...

, is published by Cribb ''et al.'' (2019). * A study on mechanisms of skeletal

biomineralization Biomineralization, also written biomineralisation, is the process by which living organisms produce minerals, often to harden or stiffen existing tissues. Such tissues are called mineralized tissues. It is an extremely widespread phenomenon; ...

in early animals (focusing on '' Cloudina'' and

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

hyoliths Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as be ...

and

halkieriid The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is ''Halkieria'' , which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the smal ...

s) is published by

Gilbert Gilbert may refer to: People and fictional characters * Gilbert (given name), including a list of people and fictional characters *Gilbert (surname), including a list of people Places Australia * Gilbert River (Queensland) * Gilbert River (South ...

''et al.'' (2019). * A study on the relationship between atmospheric oxygen oscillations, the extent of shallow-ocean oxygenation and the animal biodiversity in the Cambrian period is published by He ''et al.'' (2019). * A study on the course of the transition from microbial-dominated reef environments to animal-based reefs in the early Cambrian, as indicated by data from strata in the western

Basin and Range Basin and range topography is characterized by alternating parallel mountain ranges and valleys. It is a result of crustal extension due to mantle upwelling, gravitational collapse, crustal thickening, or relaxation of confining stresses. The e ...

of

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

and

Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...

, is published by Cordie, Dornbos & Marenco (2019). * An assemblage of early Cambrian

small carbonaceous fossil Small carbonaceous fossils (SCFs) are sub-millimetric organic remains of organisms preserved in sedimentary strata. This category of fossils has traditionally included robust or thick-walled entities such as plant spores, acritarchs and chitinozoa ...

s and

acritarch Acritarchs are organic microfossils, known from approximately 1800 million years ago to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity refle ...

s, including possible oldest known

annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...

remains, is described from the

siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, p ...

s of the

Lappajärvi Lappajärvi is a municipality in Finland in Southern Ostrobothnia region. It is from Lappajärvi to Seinäjoki, to Kokkola and to Vaasa. The municipality has a population of () and covers an area of of which is water. The population density ...

impact structure An impact structure is a generally circular or craterlike geologic structure of deformed bedrock or sediment produced by impact on a planetary surface, whatever the stage of erosion of the structure. In contrast, an impact crater is the surface ...

(

Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...

) by Slater & Willman (2019). * A study aiming to explain the occurrence of the variety of trace fossils associated with ''

Tuzoia ''Tuzoia'' (from Mount Tuzo, a mountain in the Canadian Rockies) is an extinct genus of large bivalved arthropod known from Early to Middle Cambrian marine environments from what is now North America, Australia, China, Europe and Siberia. The larg ...

''

carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...

s from the Cambrian

Burgess Shale The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest foss ...

(

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) is published by Mángano, Hawkes &

Caron A caron (), háček or haček (, or ; plural ''háčeks'' or ''háčky'') also known as a hachek, wedge, check, kvačica, strešica, mäkčeň, varnelė, inverted circumflex, inverted hat, flying bird, inverted chevron, is a diacritic mark (� ...

(2019). * Cambrian

Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...

from the Qingjiang biota ( Shuijingtou Formation;

Hubei Hubei (; ; alternately Hupeh) is a landlocked province of the People's Republic of China, and is part of the Central China region. The name of the province means "north of the lake", referring to its position north of Dongting Lake. The prov ...

,

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), preserving fossils of diverse, ~518 million years old biota, is reported by Fu ''et al.'' (2019). * A study aiming to infer whether a marked drop in known diversity of marine life during the period between the

Cambrian explosion The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil recor ...

and the

Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced wi ...

(the

Furongian The Furongian is the fourth and final epoch and series of the Cambrian. It lasted from to million years ago. It succeeds the Miaolingian series of the Cambrian and precedes the Lower Ordovician Tremadocian Stage. It is subdivided into three sta ...

Gap) is apparent, due to sampling failure or lack of rock, or real, is published by

Harper Harper may refer to: Names * Harper (name), a surname and given name Places ;in Canada * Harper Islands, Nunavut *Harper, Prince Edward Island ;In the United States *Harper, former name of Costa Mesa, California in Orange County * Harper, Il ...

''et al.'' (2019). * A study on the marine biodiversity changes throughout the first 120 million years of the

Phanerozoic The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when anima ...

is published by Rasmussen ''et al.'' (2019). * A study aiming to determine factors influencing early Palaeozoic marine biodiversity is published by Penny & Kröger (2019). * A study on rates of origination and extinction at the genus level throughout early

Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ' ...

is published by Kröger, Franeck & Rasmussen (2019), who also present estimates of longevity, taxon age and taxon life expectancy of early Paleozoic marine genera. * A review of biodiversity curves of marine organisms throughout early Paleozoic, indicating the occurrence of a large-scale, long-term radiation of life that started during late Precambrian time and was only finally interrupted in the Devonian Period, is published online by

Harper Harper may refer to: Names * Harper (name), a surname and given name Places ;in Canada * Harper Islands, Nunavut *Harper, Prince Edward Island ;In the United States *Harper, former name of Costa Mesa, California in Orange County * Harper, Il ...

, Cascales-Miñana & Servais (2019). * A study on processes causing fluctuations of biodiversity of marine invertebrates throughout the Phanerozoic is published by Rominger, Fuentes & Marquet (2019). * A study on the impact of environmental changes on the biodiversity of North American marine organisms throughout the Phanerozoic is published by Roberts & Mannion (2019). * A study testing the hypothesis that the influence of

ocean chemistry Marine chemistry, also known as ocean chemistry or chemical oceanography, is influenced by plate tectonics and seafloor spreading, turbidity currents, sediments, pH levels, atmospheric constituents, metamorphic activity, and ecology. The fiel ...

and climate on the ecological success of marine calcifiers decreased throughout the Phanerozoic is published by Eichenseer ''et al.'' (2019). * A study on genus origination and extinction rates in the

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...

on a global scale, for the

paleocontinent A paleocontinent or palaeocontinent is a distinct area of continental crust that existed as a major landmass in the geological past. There have been many different landmasses throughout Earth's time. They range in sizes, some are just a collection ...

s

Baltica Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, is mo ...

and

Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although ...

, and for onshore and offshore areas, is published by Franeck & Liow (2019). * First

Middle Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The ...

(

Dapingian The Dapingian is the third stage of the Ordovician period and the first stage of the Middle Ordovician series. It is preceded by the Floian and succeeded by the Darriwilian. The top of the Floian is defined as the first appearance of the conodont ...

–

Darriwilian The Darriwilian is the upper stage of the Middle Ordovician. It is preceded by the Dapingian and succeeded by the Upper Ordovician Sandbian Stage. The lower boundary of the Darriwilian is defined as the first appearance of the graptolite species '' ...

) soft-bodied fossils from northern

Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...

(fossils of

medusozoa Medusozoa is a clade in the phylum Cnidaria, and is often considered a subphylum. It includes the classes Hydrozoa, Scyphozoa, Staurozoa and Cubozoa, and possibly the parasitic Polypodiozoa. Medusozoans are distinguished by having a medusa stage ...

n possibly belonging to the genus '' Patanacta'', possible members of the family Wiwaxiidae and an

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

possibly belonging to the family Pseudoarctolepidae) are described from the Valongo Formation (

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

) by Kimmig ''et al.'' (2019). * New Konservat-

Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...

containing exceptionally preserved soft-bodied organisms, including the earliest record of

Acoelomorpha Acoelomorpha is a subphylum of very simple and small soft-bodied animals with planula-like features which live in marine or brackish waters. They usually live between grains of sediment, swimming as plankton, or crawling on other organisms, suc ...

,

Turbellaria The Turbellaria are one of the traditional sub-divisions of the phylum Platyhelminthes (flatworms), and include all the sub-groups that are not exclusively parasitic. There are about 4,500 species, which range from to large freshwater forms more ...

,

Nemertea Nemertea is a phylum of animals also known as ribbon worms or proboscis worms, consisting of 1300 known species. Most ribbon worms are very slim, usually only a few millimeters wide, although a few have relatively short but wide bodies. Many h ...

and

Nematoda The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broa ...

reported so far, is described from the Ordovician (

Katian The Katian is the second stage of the Upper Ordovician. It is preceded by the Sandbian and succeeded by the Hirnantian Stage. The Katian began million years ago and lasted for about 7.8 million years until the beginning of the Hirnantian millio ...

) Vauréal Formation (

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) by Knaust & Desrochers (2019). * A review of occurrence data of latest Ordovician

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

marine organisms is published by Wang, Zhan & Percival (2019), who evaluate the implications of the studied data for the knowledge of the course of the end-Ordovician mass extinction. * A revision of

Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...

fauna from the

Pentland Hills The Pentland Hills are a range of hills southwest of Edinburgh, Scotland. The range is around in length, and runs southwest from Edinburgh towards Biggar and the upper Clydesdale. Etymology The name is first recorded for the farm of Pentlan ...

(

Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...

) described by

Archibald Lamont Archibald Lamont (21 October 1907 – 16 March 1985) was a Scottish geologist, palaeontologist, Scottish Nationalist writer, poet and politician. He named the trilobite genus '' Wallacia'' after William Wallace. Life Born on 21 October 1907 a ...

in 1978 is published by Candela & Crighton (2019). * A study on the course of

graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and ...

extinctions during the middle

Homerian In the geologic timescale, the Homerian is an age of the Wenlock Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 430.5 ± 0.7 Ma and 427.4 ± 0.5 Ma (million years ago), approximately. The H ...

biotic crisis and on the impact of this crisis on other marine invertebrates, as indicated by data from the Kosov Quarry section of the Prague Synform (

Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...

), is published by Manda ''et al.'' (2019). * Well-preserved fossil cryptic biota is reported from the submarine cavities of the

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

(

Emsian The Emsian is one of three faunal stages in the Early Devonian Epoch. It lasted from 407.6 ± 2.6 million years ago to 393.3 ± 1.2 million years ago. It was preceded by the Pragian Stage and followed by the Eifelian Stage. It is named after the ...

to

Givetian The Givetian is one of two faunal stages in the Middle Devonian Period. It lasted from million years ago to million years ago. It was preceded by the Eifelian Stage and followed by the Frasnian Stage. It is named after the town of Givet in Fra ...

) mud mounds in the Hamar Laghdad area (

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

) by Berkowski ''et al.'' (2019). * A study aiming to test and quantify the classification of Devonian

biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...

areas, based on distributional data of Devonian trilobite, brachiopod and fish taxa, is published by Dowding & Ebach (2019). * A study on patterns of local richness of terrestrial tetrapods throughout the Phanerozoic is published by Close ''et al.'' (2019). * Description of tetrapod and fish fossils from the coastal locality of Burnmouth, Scotland (Ballagan Formation), associated plant material and

sedimentological Sedimentology encompasses the study of modern sediments such as sand, silt, and clay, and the processes that result in their formation (erosion and weathering), transport, deposition and diagenesis. Sedimentologists apply their understanding of mo ...

context of these fossils is published by Clack ''et al.'' (2019), who interpret these fossils as evidence of the potential richness of the Tournaisian fauna, running counter to the assumption of a depauperate nonmarine fauna following the end-

Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...

Hangenberg event The Hangenberg event, also known as the Hangenberg crisis or end-Devonian extinction, is a mass extinction that occurred at the end of the Famennian stage, the last stage in the Devonian Period (roughly 358.9 ± 0.4 million years ago). It is usuall ...

. * A study on the impact of climate changes during the Carboniferous–Permian transition on the evolution of land-living vertebrates is published by Pardo ''et al.'' (2019). * A study aiming to test one of the scenarios proposed by

Robert L. Carroll Robert "Bob" Lynn Carroll (May 5, 1938 – April 8, 2020) was an American–Canadian vertebrate paleontologist who specialised in Paleozoic and Mesozoic amphibians and reptiles. Biography Carroll was an only child and grew up on a farm ne ...

in 1970 to explain the origin of the amniotic egg, based on data from Permo-Carboniferous tetrapods, is published by Didier, Chabrol &

Laurin Laurin is both a surname and a given name. Notable people with the name include: Surname * Anna-Lena Laurin (born 1962), Swedish composer * Camille Laurin (1922–1999), psychiatrist and politician in Quebec, Canada * Dan Laurin (born 1960), Swe ...

(2019). * An overview of the studies researching biodiversity changes in the Permian and their links to volcanism is published by Chen & Xu (2019). * Haridy ''et al.'' (2019) report the occurrence of overgrowth of

palatal The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly separ ...

dentition of ''

Cacops ''Cacops'' ("ugly look" for its strange appearance), is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. ''Cacops'' is one of the few olsoniforms (dissorophids and the larger trematopids) w ...

'' and ''

Captorhinus ''Captorhinus'' (from el, καπτō , 'to gulp down' and el, ῥῑνός , 'nose') is an extinct genus of captorhinid reptiles that lived during the Permian period. Its remains are known from Oklahoma, Texas, Europe, India, the Pedra de Fogo ...

'' by a new layer of bone to which the newest teeth are then attached (the overgrowth pattern also documented in early fishes), and evaluate the implications of this finding for the knowledge of the origin of teeth. * A study on the severity of the end-Guadalupian extinction event is published online by Rampino & Shen (2019). * A study on the ecology of Permian tetrapods from the Abrahamskraal Formation (

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

), as indicated by stable oxygen isotope compositions of phosphate from teeth and bones used as a proxy for water dependence, is published online by Rey ''et al.'' (2019). * Two Permian tetrapod assemblages, recovered from the northernmost point at which the lowest

Beaufort Group The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlie ...

has been targeted for collecting fossils, are reported from the southern Free State (South Africa) by Groenewald, Day & Rubidge (2019), who evaluate the implications of these fossils for the knowledge of faunal provincialism within the Middle to Late Permian Karoo Basin. * A study aiming to determine which Permian tetrapod assemblage zones are present in the vicinity of Victoria West (

Northern Cape The Northern Cape is the largest and most sparsely populated province of South Africa. It was created in 1994 when the Cape Province was split up. Its capital is Kimberley. It includes the Kalahari Gemsbok National Park, part of the Kgalagadi T ...

,

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

), and to reassess the

biostratigraphic Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Bio ...

provenance of specimens collected historically in this area (including the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

of ''

Lycaenops ''Lycaenops'' ("wolf-face") is a genus of carnivorous therapsids. It lived during the Middle Permian to the early Late Permian, about 260 mya, in what is now South Africa. Description ''Lycaenops'' measured about and weighed up to . Like the ...

ornatus''), is published by Day & Rubidge (2019). * A study on the course of the turnover from the ''

Daptocephalus ''Daptocephalus'' is an extinct genus of non- mammalian synapsid anomodont dicynodont, it which was found in Late Permian strata, in a biozone known precisely for the presence of fossils of this dicynodont, the ''Daptocephalus'' Zone, in the Ka ...

'' to ''Lystrosaurus'' Assemblage Zones of the Karoo Basin is published by Gastaldo ''et al.'' (2019). * A study on the timing of the extinction of latest Permian vertebrates in the Karoo Basin of South Africa is published online by Rampino ''et al.'' (2019). * A study on the identification and position of the terrestrial end-Permian mass extinction in southern African sediments, based on data from a new site in the South African Karoo Basin, is published online by Botha ''et al.'' (2019). * A study on the functional diversity of middle Permian and Early Triassic marine paleocommunities in the area of present-day western

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

, and on its implications for the knowledge of functional re-organization of these communities in the aftermath of the Permian–Triassic extinction event, is published by Dineen, Roopnarine & Fraiser (2019). * A study aiming to explain high biodiversity preserved in the Triassic Cassian Formation (

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

) is published online by Roden ''et al.'' (2019). * A study on shark, sizable carnivorous archosaur, big herbivorous tetrapod and probable turtle

bromalite Bromalites are the fossilized remains of material sourced from the digestive system of organisms. As such, they can be broadly considered to be trace fossils. The most well-known types of bromalites are fossilized faeces or coprolites A co ...

s (

coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...

s and possibly some cololites) from a turtle-dominated fossil assemblage from the

Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. Th ...

Poręba site (

Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...

) is published by Bajdek ''et al.'' (2019), who evaluate the implications of their findings for inferring the diet of the Triassic turtle ''

Proterochersis ''Proterochersis'' is an extinct genus of turtle from the Late Triassic period ( Norian stage) of Europe. It is known from a large number of fossils uncovered in Germany and Poland.Fraas E (1913)"''Proterochersis'', eine pleurodire Schildkröte aus ...

porebensis''. * A study on seawater oxygenation during the

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...

and its impact on the recovery of marine

benthos Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone.Triassic–Jurassic extinction event The Triassic–Jurassic (Tr-J) extinction event, often called the end-Triassic extinction, marks the boundary between the Triassic and Jurassic periods, , and is one of the top five major extinction events of the Phanerozoic eon, profoundly affect ...

, as indicated by data from

Blue Lias The Blue Lias is a geological formation in southern, eastern and western England and parts of South Wales, part of the Lias Group. The Blue Lias consists of a sequence of limestone and shale layers, laid down in latest Triassic and early Jurassi ...

Formation (

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

), is published by Atkinson & Wignall (2019). * A study on the patterns and processes of recovery of marine fauna after the

Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...

oceanic

anoxic event Oceanic anoxic events or anoxic events ( anoxia conditions) describe periods wherein large expanses of Earth's oceans were depleted of dissolved oxygen (O2), creating toxic, euxinic (anoxic and sulfidic) waters. Although anoxic events have not ...

, as indicated by data from the

Cleveland Basin The Cleveland Basin is a sedimentary basin in Yorkshire, England. Formed initially by rifting during the Mississippian period of the Early Carboniferous. It is West–East trending and lies between the intrabasinal highs of the Askrigg Block and t ...

(

Yorkshire Yorkshire ( ; abbreviated Yorks), formally known as the County of York, is a Historic counties of England, historic county in northern England and by far the largest in the United Kingdom. Because of its large area in comparison with other Eng ...

, United Kingdom), is published by Caswell & Dawn (2019). * A study on changes of land vegetation resulting from the Toarcian oceanic anoxic event is published by Slater ''et al.'' (2019). * Skeletal elements of Oxfordian ichthyosaurs and plesiosaurs are reported from the Kingofjeld mountain (north-east

Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...

) by Delsett & Alsen (2019). * New marine reptile-bearing localities documenting the

Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...

–

Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...

transition at the High

Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...

(

Mendoza Province Mendoza, officially Province of Mendoza, is a province of Argentina, in the western central part of the country in the Cuyo region. It borders San Juan to the north, La Pampa and Neuquén to the south, San Luis to the east, and the republic o ...

,

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

) are reported by Fernández ''et al.'' (2019). * A study on microvertebrate fossils from the Upper Jurassic or Lower Cretaceous of Ksar Metlili (Anoual Syncline,

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

), evaluating their palaeobiogeographical implications, and on the age of this fauna, is published online by Lasseron ''et al.'' (2019). * Description of mid-Cretaceous invertebrate fauna from

Batavia Knoll Batavia may refer to: Historical places * Batavia (region), a land inhabited by the Batavian people during the Roman Empire, today part of the Netherlands * Batavia, Dutch East Indies, present-day Jakarta, the former capital of the Dutch East In ...

(eastern Indian Ocean), and a study on its similarities to other Cretaceous faunas from around the Indian Ocean, is published by Wild & Stilwell (2019). * A study on the age of the vertebrate fauna from the Cretaceous

Cerro Barcino Formation The Cerro Barcino Formation (also known as the Gorro Frigio Formation) is a geological formation in South America whose strata span the Early Cretaceous to the earliest Late Cretaceous. The top age for the formation has been estimated to be Cenoman ...

(

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

) is published online by Krause ''et al.'' (2019). * Possible amphibian, gastropod and insect egg masses are described from the Cretaceous amber from

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...

by Xing ''et al.'' (2019). * A study on

coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is de ...

s from the Upper Cretaceous deposits in the Münster Basin (northwestern

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

), evaluating their implications for the knowledge of Cretaceous

trophic Trophic, from Ancient Greek τροφικός (''trophikos'') "pertaining to food or nourishment", may refer to: * Trophic cascade * Trophic coherence * Trophic egg * Trophic function * Trophic hormone * Trophic level index * Trophic level * Trop ...

structures and predator–prey interactions, is published by Qvarnström ''et al.'' (2019). * New vertebrate assemblage from the upper

Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...

Schönleiten Formation of Gams bei Hieflau (

Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...

) is described by Ősi ''et al.'' (2019). * Turonian marine vertebrate fossils from the Huehuetla quarry (

Puebla Puebla ( en, colony, settlement), officially Free and Sovereign State of Puebla ( es, Estado Libre y Soberano de Puebla), is one of the 32 states which comprise the Federal Entities of Mexico. It is divided into 217 municipalities and its cap ...

,

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

) are described by Alvarado-Ortega ''et al.'' (2019). * A study on the

biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...

of Cretaceous terrestrial tetrapods is published by Kubo (2019). * A study on the structure and contents of a large piece of amber attached to a jaw of a specimen of ''

Prosaurolophus ''Prosaurolophus'' (; meaning "before ''Saurolophus''", in comparison to the later dinosaur with a similar head crest) is a genus of hadrosaurid (or duck-billed) dinosaur from the Late Cretaceous of North America. It is known from the remains of ...

maximus'' from the Cretaceous

Dinosaur Park Formation The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76. ...

(

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

), evaluating the implications of this finding for the knowledge of the habitat and taphonomy of the dinosaur, is published by McKellar ''et al.'' (2019). * An accumulation of fossil eggshells of bird,

crocodylomorph Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cro ...

and

gekkota Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from . Geckos ar ...

n eggs is reported from the Late Cretaceous Oarda de Jos locality in the vicinity of the city of

Sebeș Sebeș (; German: ''Mühlbach''; Hungarian: ''Szászsebes''; Transylvanian Saxon dialect: ''Melnbach'') is a city in Alba County, central Romania, southern Transylvania. Geography The city lies in the Mureș River valley and straddles the rive ...

(

Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...

) by Fernández ''et al.'' (2019). * A review of the fossil record of Late Cretaceous and Paleogene vertebrates from the

Seymour Island Seymour Island or Marambio Island, is an island in the chain of 16 major islands around the tip of the Graham Land on the Antarctic Peninsula. Graham Land is the closest part of Antarctica to South America. It lies within the section of the isla ...

(

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...

) is published by Reguero (2019). * A study on the evolutionary history of the family

Pospiviroidae The Pospiviroidae are a family of viroids, including the first viroid to be discovered, PSTVd. Their secondary structure is key to their biological activity. The classification of this family is based on differences in the conserved central re ...

, aiming to assess possible impact of the

Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...

on the divergence rates in this family, is published by Bajdek (2019). * A study on calcareous

nanoplankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a crucia ...

and planktic foraminiferal assemblages in a Cretaceous-Paleogene section from the peak ring of the

Chicxulub crater The Chicxulub crater () is an impact crater buried underneath the Yucatán Peninsula in Mexico. Its center is offshore near the community of Chicxulub, after which it is named. It was formed slightly over 66 million years ago when a large ast ...

, and on their implications for the knowledge of recovery of plankton after the Cretaceous–Paleogene extinction event, is published by Jones, Lowery & Bralower (2019). * A study on the course of recovery of the nanoplankton communities after the Cretaceous–Paleogene extinction event is published by Alvarez ''et al.'' (2019), who report evidence indicative of 1.8 million years of exceptional volatility of post-extinction communities and indicating that the emergence of a more stable equilibrium-state community coincided with indicators of

carbon cycle The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and Earth's atmosphere, atmosphere of the Earth. Carbon is the main component of biological compounds as well as ...

restoration and a fully functioning

biological pump The biological pump (or ocean carbon biological pump or marine biological carbon pump) is the ocean's biologically driven sequestration of carbon from the atmosphere and land runoff to the ocean interior and seafloor sediments.Sigman DM & GH ...

. * A study on the timing and nature of recovery of

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

marine ecosystems of

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...

after the Cretaceous–Paleogene mass extinction, as indicated by data from fossils of benthic molluscs, is published by Whittle ''et al.'' (2019). * A study on the drivers and tempo of biotic recovery after Cretaceous–Paleogene mass extinction, as indicated by data from the Corral Bluffs section of the

Denver Basin The Denver Basin, variously referred to as the Julesburg Basin, Denver-Julesburg Basin (after Julesburg, Colorado), or the D-J Basin, is a geologic structural basin centered in eastern Colorado in the United States, but extending into southeast Wyo ...

(

Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

), is published by Lyson ''et al.'' (2019). * Description of the vertebrate assemblage from the

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

Shine Us locality in the Khaliun Basin (

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

) is published by Daxner-Höck ''et al.'' (2019). * Description of reptile and amphibian fossils from the early Miocene localities of the Kilçak section (

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...

) is published by Syromyatnikova ''et al.'' (2019). * Description of fossil fish, amphibian and reptilian fauna from the middle Miocene locality Gračanica (

Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...

) is published online by Vasilyan (2019). * A study on the vertebrate fossils from the early

Clarendonian The Clarendonian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 13,600,000 to 10,300,000 years BP, a period of . It is u ...

localities within the

Goliad Formation The Goliad Formation (Tg)Page et al., 2005, p.5 is a Formation (geology), geologic formation in Texas. It preserves fossils dating back to the Serravallian to earliest Pliocene stages (Clarendonian, Hemphillian and earliest Blancan in the North Ame ...

in

Bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...

and

Live Oak Live oak or evergreen oak is any of a number of oaks in several different sections of the genus ''Quercus'' that share the characteristic of evergreen foliage. These oaks are not more closely related to each other than they are to other oaks. ...

Counties in

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

(comprising the Lapara Creek Fauna), and on the

stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostrati ...

context of these localities, is published by May (2019). * New late Miocene vertebrate assemblage, including turtle, rodent and xenarthran fossils (among which is the oldest record of an

armadillo Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along wi ...

belonging to the genus '' Dasypus'' reported so far), is described from the Los Alisos locality (Guanaco Formation,

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

) by Ercoli ''et al.'' (2019). * Description of a diverse late Miocene marine fauna from the Bloomfield Quarry ( Wilson Grove Formation;

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

), including the most diverse assemblage of fossil

walruses The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fami ...

yet reported worldwide from a single locality, is published by Powell ''et al.'' (2019). * Fish, turtle and mammals fossils are described from a locality near Whitehorse (

Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

), probably of Miocene age, by Eberle ''et al.'' (2019). * A study on microscopic traces of hominin and animal activities in the

Denisova Cave Denisova Cave (russian: Денисова пещера, lit= the cave of Denis, translit= Denísova peshchéra; alt, Аю-Таш, lit= Bear Rock, translit= Ayu Tash) is a cave in the Bashelaksky Range of the Altai mountains, Siberia, Russia. The ...

(

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

), providing the information on the use of this cave over the last 300,000 years, is published by Morley ''et al.'' (2019). * A study on the age of the Pleistocene vertebrate assemblage from the Khok Sung locality (

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...

) is published by Duval ''et al.'' (2019). * Revision of reptile and amphibian fossils from the late Pleistocene collection of the "Caverne Marie-Jeanne" (Hastière-Lavaux,

Namur Province Namur (; nl, Namen ; wa, Nameur) is a province of Wallonia, one of the three regions of Belgium. It borders (clockwise from the West) on the Walloon provinces of Hainaut, Walloon Brabant, Liège and Luxembourg in Belgium, and the French depar ...

,

Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...

) is published by Blain ''et al.'' (2019). * New late Pleistocene site Tsaramody (Sambaina basin,

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

), preserving diverse

subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...

remains of vertebrates, is reported by Samonds ''et al.'' (2019). * A study on the paleoecology and diet of late Pleistocene terrestrial vertebrates known from an asphalt deposit (Project 23, Deposit 1) at

Rancho La Brea Rancho La Brea was a Mexican land grant in present-day Los Angeles County, California, given in 1828 to Antonio Jose Rocha and Nemisio Dominguez by José Antonio Carrillo, the alcalde of Los Angeles. Rancho La Brea consisted of one square leagu ...

(

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) is published online by Fuller ''et al.'' (2019). * A study on changes of vegetation in southern

Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...

over the past 40,000 calibrated years BP, as indicated by data from Saleh Cave (

South Kalimantan South Kalimantan ( id, Kalimantan Selatan) is a Provinces of Indonesia, province of Indonesia. It is the smallest province in Kalimantan, the Indonesian territory of Borneo. The provincial capital was Banjarmasin until 15 February 2022 when it wa ...

,

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...

), is published by Wurster ''et al.'' (2019). * Late Quaternary fossils of vertebrates are described from caves in the Manning Karst Region of eastern

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

(

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

) by Price ''et al.'' (2019). * A study aiming to determine the relationships between extinctions of

megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...

, climatic changes and patterns of human appearance in south-eastern Australia over the last 120,000 years is published by Saltré ''et al.'' (2019). * A study on the causes of

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...

extinction of

megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...

of

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

is published by

Godfrey Godfrey may refer to: People * Godfrey (name), a given name and surname * Godfrey (comedian), American comedian, actor Places In the United States * Godfrey, Georgia, an unincorporated community * Godfrey, Illinois, a village * Godfrey, Kansas, an ...

''et al.'' (2019). * A review discussing possible links between the fossil record of marine biodiversity, nutrient availability and

primary productivity In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through c ...

is published online by Martin & Servais (2019). * A study on factors which determined the relative intensity of marine extinctions during greenhouse–icehouse transitions in the Late

Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start ...

and the

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...

is published online by Saupe ''et al.'' (2019). * A study on the possible relationship between

speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...

and extinction rates of different groups of organisms and the ages of these groups, as indicated by data from extant and fossil species, is published by Henao Diaz ''et al.'' (2019). * A study on the evolution of bite force of

amniote Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are disti ...

s, as indicated by data from extant and fossil taxa, is published by Sakamoto,

Ruta ''Ruta'' (commonly known as rue) is a genus of strongly scented evergreen subshrubs, 20–60 cm tall, in the family Rutaceae, native to the Mediterranean region, Macaronesia and southwest Asia. About ten species are accepted in the genus. T ...

& Venditti (2019). * A study on the phylogenetic distribution, morphological variation and functions of apicobasal ridges (elevated ridges of

tooth enamel Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The ...

) in aquatic reptiles and mammals, as indicated by data from extant and fossil taxa, is published by McCurry ''et al.'' (2019). * A study on the impact of uncertainty of stratigraphic age of fossils on studies estimating species divergence times which incorporate fossil taxa, based on data from the fossil record of North American mammals and from the dataset of extant and fossil cetaceans, is published by Barido-Sottani ''et al.'' (2019). * A study evaluating the impact of information about stratigraphic ranges of fossil taxa on the analyses of timing of evolutionary divergence is published online by Püschel ''et al.'' (2019). * A study on anatomical distribution, abundance, geometry,

melanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...

chemistry and elemental inventory of

melanosome A melanosome is an organelle found in animal cells and is the site for synthesis, storage and transport of melanin, the most common light-absorbing pigment found in the animal kingdom. Melanosomes are responsible for color and photoprotection i ...

s in tissues of extant vertebrates, evaluating their implications for reconstructions of internal soft-tissue anatomy in fossil vertebrates, is published by Rossi ''et al.'' (2019). * A study on the

chronostratigraphy Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time. The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geologica ...

and

biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock Stratum, strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictiona ...

of Cenozoic vertebrate (mostly mammal) fossils from the

South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...

Coastal Plain is published by Albright ''et al.'' (2019).

Other research

Other research related to paleontology, including research related to

geology Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other Ear ...

,

palaeogeography Palaeogeography (or paleogeography) is the study of historical geography, generally physical landscapes. Palaeogeography can also include the study of human or cultural environments. When the focus is specifically on landforms, the term pale ...

,

paleoceanography Paleoceanography is the study of the history of the oceans in the geology, geologic past with regard to circulation, chemistry, biology, geology and patterns of sedimentation and biological productivity. Paleoceanographic studies using environmen ...

and

paleoclimatology Paleoclimatology (British spelling, palaeoclimatology) is the study of climates for which direct measurements were not taken. As instrumental records only span a tiny part of Earth's history, the reconstruction of ancient climate is important to ...

. * A study on the biological oxygen production during the

Mesoarchean The Mesoarchean (, also spelled Mesoarchaean) is a geologic era in the Archean Eon, spanning , which contains the first evidence of modern-style plate subduction and expansion of microbial life. The era is defined chronometrically and is not r ...

, as indicated by data from Mesoarchean shales of the Mozaan Group (Pongola Supergroup,

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

) preserving record of a shallow ocean "oxygen oasis", is published by Ossa Ossa ''et al.'' (2019). * A study on the extent of the oxygenation of ocean waters over

continental shelves A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

before the

Great Oxidation Event The Great Oxidation Event (GOE), also called the Great Oxygenation Event, the Oxygen Catastrophe, the Oxygen Revolution, the Oxygen Crisis, or the Oxygen Holocaust, was a time interval during the Paleoproterozoic era when the Earth's atmosphere ...

, as indicated by data from 2.5-billion-year-old Mount McRae Shale (

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

), is published by Ostrander ''et al.'' (2019). * A study on the extent of the oxygenation of shallow oceans 2.45 billion years ago is published by Rasmussen ''et al.'' (2019), who interpret their findings as indicating that oxygen levels both the surface oceans and atmosphere were exceedingly low before the Great Oxidation Event, which the authors interpret as directly caused by evolution of oxygenic

photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...

. * A study aiming to determine whether the overall size of the

biosphere The biosphere (from Greek βίος ''bíos'' "life" and σφαῖρα ''sphaira'' "sphere"), also known as the ecosphere (from Greek οἶκος ''oîkos'' "environment" and σφαῖρα), is the worldwide sum of all ecosystems. It can also be ...

decreased at the end of the Great Oxidation Event, based on data on isotope

geochemistry Geochemistry is the science that uses the tools and principles of chemistry to explain the mechanisms behind major geological systems such as the Earth's crust and its oceans. The realm of geochemistry extends beyond the Earth, encompassing the e ...

of

sulfate The sulfate or sulphate ion is a polyatomic anion with the empirical formula . Salts, acid derivatives, and peroxides of sulfate are widely used in industry. Sulfates occur widely in everyday life. Sulfates are salts of sulfuric acid and many ar ...

minerals from the Belcher Group (subarctic

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

), is published by Hodgskiss ''et al.'' (2019). * Evidence of a burst of mantle activity at the end of the

Archean The Archean Eon ( , also spelled Archaean or Archæan) is the second of four geologic eons of Earth's history, representing the time from . The Archean was preceded by the Hadean Eon and followed by the Proterozoic. The Earth Earth ...

(around 2.5 billion years ago) is presented by Marty ''et al.'' (2019), who interpret their findings as lending credence to models advocating a magmatic origin for environmental changes such as the Great Oxidation Event. * A study aiming to determine the effects of competition of early anoxygenic

phototroph Phototrophs () are organisms that carry out photon capture to produce complex organic compounds (e.g. carbohydrates) and acquire energy. They use the energy from light to carry out various cellular metabolic processes. It is a common misconcep ...

s and primitive oxygenic phototrophs on the Earth system, especially on the large-scale oxygenation of Earth's atmosphere ~2.3 billion years ago, is published by Ozaki ''et al.'' (2019). * A study on the

geochemistry Geochemistry is the science that uses the tools and principles of chemistry to explain the mechanisms behind major geological systems such as the Earth's crust and its oceans. The realm of geochemistry extends beyond the Earth, encompassing the e ...

of mat-related structures and their host sediments from the Francevillian Formation (

Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the north ...

) is published by Aubineau ''et al.'' (2019), who evaluate the implications of their findings for the knowledge whether ancient microbes induced illitisation (conversion of

smectite A smectite (from ancient Greek ''σμηκτός'' smektos 'lubricated'; ''σμηκτρίς'' smektris 'walker's earth', 'fuller's earth'; rubbing earth; earth that has the property of cleaning) is a mineral mixtures of various swelling sheet sil ...

to

illite Illite is a group of closely related non-expanding clay minerals. Illite is a secondary mineral precipitate, and an example of a phyllosilicate, or layered alumino-silicate. Its structure is a 2:1 sandwich of silica tetrahedron (T) – alumina ...

–smectite mixed-layer minerals), and for the knowledge of Earth's climate and ocean chemistry in the

Paleoproterozoic The Paleoproterozoic Era (;, also spelled Palaeoproterozoic), spanning the time period from (2.5–1.6 Ga), is the first of the three sub-divisions (eras) of the Proterozoic Eon. The Paleoproterozoic is also the longest era of the Earth's ...

. * A study on the organic geochemical (biomarker) signatures of the 1.38-billion-years-old black siltstones of the Velkerri Formation (

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

), and on their implications for inferring the microbial diversity and palaeoenvironment of the

Proterozoic The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided ...

Roper Seaway, is published by Jarrett ''et al.'' (2019). * A study on the origins of putative

stromatolite Stromatolites () or stromatoliths () are layered sedimentary formations (microbialite) that are created mainly by photosynthetic microorganisms such as cyanobacteria, sulfate-reducing bacteria, and Pseudomonadota (formerly proteobacteria). The ...

s and associated carbonate minerals from lacustrine sedimentary rocks of the 1.1-billion-years-old

Stoer Group In geology, the term Torridonian is the informal name for the Torridonian Group, a series of Mesoproterozoic to Neoproterozoic arenaceous and argillaceous sedimentary rocks, which occur extensively in the Northwest Highlands of Scotland. The st ...

is published by Brasier ''et al.'' (2019). * A study suggesting a link between early evolution and diversification of animals and high availability of

copper Copper is a chemical element with the symbol Cu (from la, cuprum) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkis ...

in the late Neoproterozoic is published by Parnell & Boyce (2019). * A study aiming to determine the cause of the uniquely high amplitudes of Neoproterozoic δ¹³C excursions is published by Shields ''et al.'' (2019). * A study evaluating the possible relationship between the

Cryogenian The Cryogenian (from grc, κρύος, krýos, meaning "cold" and , romanized: , meaning "birth") is a geologic period that lasted from . It forms the second geologic period of the Neoproterozoic Era, preceded by the Tonian Period and followed by ...

magmatic activity and the evolution of early life, based on data from the Cryogenian Yaolinghe Group (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), is published by Long, Zhang & Luo (2019). * Evidence for oxygenated waters near ice sheet grounding lines during the Cryogenian is presented by Lechte ''et al.'' (2019). * A study on ocean oxygen levels during the Ediacaran Shuram negative C-isotope Excursion and the middle Ediacaran, and on their implications for the evolution of the

Ediacaran biota The Ediacaran (; formerly Vendian) biota is a Taxonomy (biology), taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (). These were composed of enigmatic tubular and frond-sh ...

, is published by Zhang ''et al.'' (2019). * A study on the causes of widespread preservation of soft-bodied organisms in sandstones of the Ediacara Member in South Australia is published by Liu ''et al.'' (2019). * A study on the seafloor oxygen

fugacity In chemical thermodynamics, the fugacity of a real gas is an effective partial pressure which replaces the mechanical partial pressure in an accurate computation of the chemical equilibrium constant. It is equal to the pressure of an ideal gas whic ...

in the time of the emergence of the earliest known

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

animals, as inferred from data from the latest Ediacaran Dengying Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), is published by Ding ''et al.'' (2019). * A study on the process of fossilization of Ediacaran organisms, and on its impact on the preservation of the external shape of these organisms, is published by Bobrovskiy ''et al.'' (2019). * A study on the global extent of the oxygenation of seafloor, surface oceans and atmosphere during early Cambrian is published by Dahl ''et al.'' (2019), who report evidence of two major oceanic anoxic events in the early Cambrian. * A study on nitrogen isotope and organic carbon isotope data from the lower Cambrian Niutitang Formation (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) is published online by Xu ''et al.'' (2019), who link

nitrogen cycle The nitrogen cycle is the biogeochemical cycle by which nitrogen is converted into multiple chemical forms as it circulates among atmospheric, terrestrial, and marine ecosystems. The conversion of nitrogen can be carried out through both biologi ...

perturbations to animal diversification during the early Cambrian. * A study on the paleoecological characteristics of Cambrian marine ecosystems of central

Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...

(

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

) is published by Romero ''et al.'' (2019). * A study on seawater temperatures during the Cambrian, as indicated by data from oxygen isotope analyses of Cambrian brachiopod shells, is published by Wotte ''et al.'' (2019). * A study on bottom-water

redox Redox (reduction–oxidation, , ) is a type of chemical reaction in which the oxidation states of substrate (chemistry), substrate change. Oxidation is the loss of Electron, electrons or an increase in the oxidation state, while reduction ...

conditions in the late Cambrian

Alum Shale The Alum Shale Formation (also known as alum schist and alum slate) is a formation of black shale of Middle Cambrian to Tremadocian (Lower Ordovician) in age found predominantly in southern Scandinavia. It is shale or clay slate containing py ...

Sea, as indicated by sedimentary

molybdenum Molybdenum is a chemical element with the symbol Mo and atomic number 42 which is located in period 5 and group 6. The name is from Neo-Latin ''molybdaenum'', which is based on Ancient Greek ', meaning lead, since its ores were confused with lea ...

contents of the Alum Shale, is published by Dahl ''et al.'' (2019), who interpret their findings as indicating that

anoxic The term anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts: * Anoxic waters, sea water, fresh water or groundwater that are depleted of diss ...

sulfidic bottom waters were an intermittent rather than persistent feature of Cambrian oceans, and that early animals invaded the seafloor during oxygenated periods. * A study on the paleogeographic position of all major Phanerozoic

arc-continent A continental arc is a type of volcanic arc occurring as an "arc-shape" topographic high region along a continental margin. The continental arc is formed at an active continental margin where two tectonic plates meet, and where one plate has conti ...

collisions In physics, a collision is any event in which two or more bodies exert forces on each other in a relatively short time. Although the most common use of the word ''collision'' refers to incidents in which two or more objects collide with great f ...

, comparing it with the latitudinal distribution of ice-sheets throughout the Phanerozoic, is published by Macdonald ''et al.'' (2019). * A study aiming to determine whether the

Ordovician meteor event The Ordovician meteor event was a dramatic increase in the rate at which L chondrite meteorites fell to Earth during the Middle Ordovician period, about 467.5±0.28 million years ago. This is indicated by abundant fossil L chondrite meteorites in ...

directly affected Earth's climate and biota is published by Schmitz ''et al.'' (2019). * A review of the evidence of evolutionary radiation of animals throughout the

Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced wi ...

, and of environmental changes coincident with these biotic changes, is published by Stigall ''et al.'' (2019). * A study on conodont oxygen isotope compositions in Ordovician samples from

Argentine Argentines (mistakenly translated Argentineans in the past; in Spanish (masculine) or (feminine)) are people identified with the country of Argentina. This connection may be residential, legal, historical or cultural. For most Argentines, s ...

Precordillera Precordillera is a Spanish geographical term for hills and mountains lying before a greater range, foothills. The term is derived from ''cordillera'' (mountain range)—literally "pre-mountain range"—and applied usually to the Andes. Some places ...

and

Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although ...

, and on their implications for the knowledge of palaeothermometry and drift of the Precordillera in the early Paleozoic, is published online by Albanesi ''et al.'' (2019). * A study on carbon isotope data from

stratigraphic section A stratigraphic section is a sequence of layers of rocks in the order they were deposited. It is based on the principle of original horizontality, which states that layers of sediment are originally deposited horizontally under the action of gr ...

s at Germany Valley (

West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...

) and Union Furnace (

Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...

) in the Central Appalachian Basin, evaluating its implications for the knowledge of change in atmospheric oxygen levels during the late Ordovician and its possible relationship with early diversification of land plants, is published by Adiatma ''et al.'' (2019). * Signatures of Devonian (Famennian) forests and soils preserved in black shales in the southernmost

Appalachian Basin The geology of the Appalachians dates back to more than 480 million years ago. A look at rocks exposed in today's Appalachian Mountains reveals elongate belts of folded and thrust faulted marine sedimentary rocks, volcanic rocks and slivers of ...

(

Chattanooga Shale The Chattanooga Shale is a geologic formation in Alabama, Arkansas, Kentucky, Missouri and Tennessee. It preserves conodont fossils dating to the Devonian Period. It occurs mostly as a subsurface geologic formation composed of layers of shale. It ...

;

Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) are presented by Lu ''et al.'' (2019). * A study examining the intensity of explosive volcanism from 400 to 200 million years ago, and evaluating its impact on the late Paleozoic Ice Age, is published by Soreghan, Soreghan & Heavens (2019). * Description of Cisuralian charcoal from the Barro Branco coal seam (Siderópolis Member of the

Rio Bonito Formation The Rio Bonito Formation is a Formation (stratigraphy), geological formation of the Paraná Basin, Paraná and Pelotas Basins of Permian age. It is represented by a succession of cyclic sedimentary packages of sandstones, siltstones and shales whi ...

,

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

), and a study on its implications for reconstruction of palaeo-wildfire occurrences in peat-forming vegetation through the Late Palaeozoic in

Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...

, is published by Benicio ''et al.'' (2019). * A study on the extent and causes of the end-

Capitanian In the geologic timescale, the Capitanian is an age or stage of the Permian. It is also the uppermost or latest of three subdivisions of the Guadalupian Epoch or Series. The Capitanian lasted between and million years ago. It was preceded by th ...

extinction event, based on data from the Middle to Late Permian section of the Sverdrup Basin (

Ellesmere Island Ellesmere Island ( iu, script=Latn, Umingmak Nuna, lit=land of muskoxen; french: île d'Ellesmere) is Canada's northernmost and List of Canadian islands by area, third largest island, and the List of islands by area, tenth largest in the world. ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

), is published online by Bond, Wignall & Grasby (2019). * A study on the ocean chemistry during the

Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as ...

, as indicated by data from a new

stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostrati ...

section in

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

, and on its implications for the knowledge of the causes of this extinction, is published by Burger, Estrada & Gustin (2019). * A study aiming to determine the

stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostrati ...

position of the end-Permian biotic crisis in the

Sydney Basin The Sydney Basin is an interim Australian bioregion and is both a structural entity and a depositional area, now preserved on the east coast of New South Wales, Australia and with some of its eastern side now subsided beneath the Tasman Sea. ...

(

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

) is published by Fielding ''et al.'' (2019), who also attempt to determine the climate changes in this region concurrent with the end-Permian extinction. * A study on shifts in volcanic activity across the Permian-Triassic boundary, as indicated by measurements of

mercury Mercury commonly refers to: * Mercury (planet), the nearest planet to the Sun * Mercury (element), a metallic chemical element with the symbol Hg * Mercury (mythology), a Roman god Mercury or The Mercury may also refer to: Companies * Merc ...

in marine sections across the Northern Hemisphere, is published by Shen ''et al.'' (2019). * A study on mercury enrichments in Permian-Triassic boundary sections from Lubei (South China craton) and Dalongkou (Junggar terrane), and on their implications for the knowledge of volcanic activity during the Permian-Triassic transition, is published by Shen ''et al.'' (2019). * Evidence of the environmental transition from meandering to braided rivers and of the development of desert-like conditions in the earliest Triassic is reported from Permian-Triassic boundary sections in

Shanxi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-lev ...

(China) by Zhu ''et al.'' (2019). * A study on the nitrogen isotope variations in oceanic waters in the aftermath of the end-Permian mass extinction is published by Sun ''et al.'' (2019), whose conceptual model indicates

ammonium The ammonium cation is a positively-charged polyatomic ion with the chemical formula or . It is formed by the protonation of ammonia (). Ammonium is also a general name for positively charged or protonated substituted amines and quaternary a ...

intoxication of the oceans during this time period. * A study on microbially induced sedimentary structures from the Lower Triassic Blind Fiord Formation (Arctic

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

), evaluating their implications for the knowledge of the course of biotic recovery in the aftermath of the Permian–Triassic extinction event, is published online by Wignall ''et al.'' (2019). * A study on the oxygen isotope compositions of discrete conodont elements from the Lower Triassic Mianwali Formation (

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...

), and on their implications for inferring the timing of temperature changes and the interrelationship between climate and biodiversity patterns during the Smithian-Spathian biotic crisis, is published by Goudemand ''et al.'' (2019). * A study on nutrient availability through the Early to Middle Triassic along the northern margin of Pangea is published online by Grasby ''et al.'' (2019). * A study on the character and extent of the Triassic Boreal Ocean delta plain across the area of the present-day

Barents Sea The Barents Sea ( , also ; no, Barentshavet, ; russian: Баренцево море, Barentsevo More) is a marginal sea of the Arctic Ocean, located off the northern coasts of Norway and Russia and divided between Norwegian and Russian territo ...

, interpreted as the largest delta plain reported so far, is published by Klausen, Nyberg & Helland-Hansen (2019). * A study aiming to determine links between volcanic activity in the

Central Atlantic magmatic province The Central Atlantic magmatic province (CAMP) is the Earth's largest continental large igneous province, covering an area of roughly 11 million km2. It is composed mainly of basalt that formed before Pangaea broke up in the Mesozoic Era, near the ...

, elevated concentrations of

mercury Mercury commonly refers to: * Mercury (planet), the nearest planet to the Sun * Mercury (element), a metallic chemical element with the symbol Hg * Mercury (mythology), a Roman god Mercury or The Mercury may also refer to: Companies * Merc ...

in marine and terrestrial sediments and abnormalities of fossil fern spores across the Triassic-Jurassic boundary in southern Scandinavia and northern Germany is published by Lindström ''et al.'' (2019). * A study aiming to reconstruct the palaeoenvironmental changes of the late

Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 190.8 ± 1.5 Ma and 182.7 ± 1.5 Ma (million years ago). The Plien ...

outside of Western

Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

and to test their temporal relation to

large igneous province A large igneous province (LIP) is an extremely large accumulation of igneous rocks, including intrusive (sills, dikes) and extrusive (lava flows, tephra deposits), arising when magma travels through the crust towards the surface. The formation ...

volcanism is published by De Lena ''et al.'' (2019). * Krencker, Lindström & Bodin (2019) present

sedimentological Sedimentology encompasses the study of modern sediments such as sand, silt, and clay, and the processes that result in their formation (erosion and weathering), transport, deposition and diagenesis. Sedimentologists apply their understanding of mo ...

, paleontological and

geochemical Geochemistry is the science that uses the tools and principles of chemistry to explain the mechanisms behind major geological systems such as the Earth's crust and its oceans. The realm of geochemistry extends beyond the Earth, encompassing the e ...

evidence from the Central

High Atlas High Atlas, also called the Grand Atlas ( ar, الأطلس الكبير, Al-Aṭlas al-Kabīr; french: Haut Atlas; shi, ⴰⴷⵔⴰⵔ ⵏ ⴷⵔⵏ ''Adrar n Dern''), is a mountain range in central Morocco, North Africa, the highest part of t ...

Basin (

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

) and

Jameson Land Jameson Land is a peninsula in eastern Greenland. Geography Jameson Land is bounded to the southwest by Scoresby Sound (the world's largest fjord), to the northwest by the Stauning Alps, to the north by Scoresby Land, to the northeast by the Flemi ...

(

Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...

) indicative of the occurrence of a major sea-level drop prior to the onset of the Toarcian oceanic anoxic event. * A study on the duration of the Toarcian oceanic anoxic event, as indicated by data from the Talghemt section in the

High Atlas High Atlas, also called the Grand Atlas ( ar, الأطلس الكبير, Al-Aṭlas al-Kabīr; french: Haut Atlas; shi, ⴰⴷⵔⴰⵔ ⵏ ⴷⵔⵏ ''Adrar n Dern''), is a mountain range in central Morocco, North Africa, the highest part of t ...

(

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...

), is published by Boulila ''et al.'' (2019). * A study on the

Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...

palaeoenvironment of La Voulte (

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...

), as indicated by data from exceptionally preserved eyes of the

polychelida Polychelida is an infraorder of decapod crustaceans. Fossil representatives are known dating from as far back as the Upper Triassic. A total of 38 extant species, all in the family Polychelidae, and 55 fossil species have been described. Histor ...

n lobster '' Voulteryon parvulus'' and from

epibiont An epibiont (from the Ancient Greek meaning "living on top of") is an organism that lives on the surface of another living organism, called the basibiont ("living underneath"). The interaction between the two organisms is called epibiosis. An epi ...

ic brachiopods associated with ''V. parvulus'', is published by Audo ''et al.'' (2019). * A study comparing the Jurassic floras of the Ayuquila Basin and the Otlaltepec Basin (

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

) and evaluating their implications for the knowledge of the Jurassic environments of these basins is published by Velasco-de León ''et al.'' (2019). * A study on

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...

paleomagnetism Paleomagnetism (or palaeomagnetismsee ), is the study of magnetic fields recorded in rocks, sediment, or archeological materials. Geophysicists who specialize in paleomagnetism are called ''paleomagnetists.'' Certain magnetic minerals in rock ...

, based on an updated set of Jurassic paleopoles from

Adria Adria is a town and ''comune'' in the province of Rovigo in the Veneto region of northern Italy, situated between the mouths of the rivers Adige and Po River, Po. The remains of the Etruria, Etruscan city of Atria or Hatria are to be found below ...

(

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

), is published by Muttoni & Kent (2019). * A study on the

chronostratigraphy Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time. The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geologica ...

of the Upper Jurassic

Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...

is published by Maidment & Muxworthy (2019). * Evidence of repeated significant oceanic and biotic turnovers in the area of the present-day

Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...

at the Jurassic-Cretaceous transition is presented by Zell ''et al.'' (2019). * A study on the age of the dinosaur-bearing Upper Jurassic–Lower Cretaceous sediments of western

Maestrazgo Basin The Maestrazgo () or Maestrat () is a natural and historical mountainous region, located at the eastern end of the Sistema Ibérico mountain range, in Spain. It encompasses the north of the Autonomous Community of Valencia, in Castellón provinc ...

and South-Iberian Basin (eastern

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

), aiming to also reconstruct the palaeoenvironments of this area on the basis of data from these sediments, is published by Campos-Soto ''et al.'' (2019). * A review of data on the Jurassic and Cretaceous climates of Siberia is published by Rogov ''et al.'' (2019). * A study on global climatic changes during the Early Cretaceous, focusing on the duration and magnitude of Early Cretaceous cold episodes, is published by Vickers ''et al.'' (2019). * Evidence from the Lower Cretaceous strata around the southern margin of the

Eromanga Basin The Eromanga Basin is a large Mesozoic sedimentary basin in central and northern Australia. It covers parts of Queensland, the Northern Territory, South Australia, and New South Wales, and is a major component of the Great Artesian Basin. The E ...

(

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

) indicative of cold (limited glacial and/or seasonal freezing) conditions persisting in Southern Australia through the

Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...

and the

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

is presented by Alley, Hore & Frakes (2019). * A study on

phototropism Phototropism is the growth of an organism in response to a light stimulus. Phototropism is most often observed in plants, but can also occur in other organisms such as fungi. The cells on the plant that are farthest from the light contain a hor ...

in extant trees from Beijing and

Jilin Jilin (; alternately romanized as Kirin or Chilin) is one of the three provinces of Northeast China. Its capital and largest city is Changchun. Jilin borders North Korea (Rasŏn, North Hamgyong, Ryanggang and Chagang) and Russia (Prim ...

Provinces and fossil tree trunks from the

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...

Tiaojishan and Tuchengzi formations in

Liaoning Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmost ...

and Beijing regions (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

), and on its implications for inferring the history of the rotation of the North China Block, is published by Jiang ''et al.'' (2019). * A study on the age of the Cretaceous

Cloverly Formation The Cloverly Formation is a geological formation of Early and Late Cretaceous age (Valanginian to Cenomanian stage) that is present in parts of Montana, Wyoming, Colorado and Utah in the western United States. It was named for a post office on t ...

is published by D'Emic ''et al.'' (2019). * Evidence from the

chronostratigraphy Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time. The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geologica ...

, fossil content, bracketing

facies In geology, a facies ( , ; same pronunciation and spelling in the plural) is a body of rock with specified characteristics, which can be any observable attribute of rocks (such as their overall appearance, composition, or condition of formatio ...

and ages of the Cretaceous

Wayan Formation The Wayan Formation is a geological formation in Idaho whose strata date back to the latest Early Cretaceous and the earliest Late Cretaceous. Dinosaur, other reptile, mammal, and micro and macro-floral remains are among the fossils that have been ...

of

Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...

and Vaughn Member of the Blackleaf Formation of

Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...

, indicating that they represent the same depositional system prior to disruption by subsequent tectonic and volcanic events, is presented by Krumenacker (2019). * A study on

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

plants from the Redmond no.1 mine near Schefferville ( Redmond Formation;

Labrador Peninsula The Labrador Peninsula, or Quebec-Labrador Peninsula, is a large peninsula in eastern Canada. It is bounded by the Hudson Bay to the west, the Hudson Strait to the north, the Labrador Sea to the east, and the Gulf of Saint Lawrence to the southe ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) and on their implications for the knowledge of paleoclimate of this site is published by Demers-Potvin & Larsson (2019). * The first high-resolution record of

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

–

Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...

paleotemperatures from the Southern Hemisphere, as indicated by data from the

Ocean Drilling Program The Ocean Drilling Program (ODP) was a multinational effort to explore and study the composition and structure of the Earth's oceanic basins. ODP, which began in 1985, was the successor to the Deep Sea Drilling Project initiated in 1968 by th ...

Site 1138 on the

Kerguelen Plateau The Kerguelen Plateau (, ), also known as the Kerguelen–Heard Plateau, is an oceanic plateau and a large igneous province (LIP) located on the Antarctic Plate, in the southern Indian Ocean. It is about to the southwest of Australia and is ...

, is presented by Robinson ''et al.'' (2019). * A study on the impact of marine biogeochemical processes on the

Cretaceous Thermal Maximum The Cretaceous Thermal Maximum (CTM), also known as Cretaceous Thermal Optimum, was a period of climatic warming that reached its peak approximately 90 million years ago (90 Ma) during the Turonian age of the Late Cretaceous epoch. The CTM is notab ...

is published by Wallmann ''et al.'' (2019). * A study on the age of the

Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

Wadi Milk Formation The Wadi Milk Formation is a geological formation in Sudan whose strata date back to the Late Cretaceous. Originally, the formation was thought to be Albian to Cenomanian, later research has provided dating to the Campanian to Maastrichtian. Dinosa ...

(

Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...

) is published by Owusu Agyemang ''et al.'' (2019). * A study on

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

to

Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...

polar environmental conditions at eight locations in northeast

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

and northern

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

is published online by Spicer ''et al.'' (2019). * A study on variability of carbon, oxygen and nitrogen isotopes in multiple tissues from a wide array of extant vertebrate taxa from the

Atchafalaya River The Atchafalaya River ( french: La Rivière Atchafalaya, es, Río Atchafalaya) is a distributary of the Mississippi River and Red River in south central Louisiana in the United States. It flows south, just west of the Mississippi River, and ...

Basin in

Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...

(inferred to be an environmental analogue to the Late Cretaceous coastal floodplains of North America), and on its implications for formulating and testing predictions about ancient ecological communities based on stable isotope data from fossil specimens, is published by Cullen ''et al.'' (2019). * A study on the general distribution and

stratigraphy Stratigraphy is a branch of geology concerned with the study of rock (geology), rock layers (Stratum, strata) and layering (stratification). It is primarily used in the study of sedimentary rock, sedimentary and layered volcanic rocks. Stratigrap ...

of the lower shale member of the

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

Aguja Formation The Aguja Formation is a geological formation in North America, exposed in Texas, United States and Chihuahua and Coahuila in Mexico, whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered f ...

(

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

), and a revision of all significant larger vertebrate fossil specimens from these strata, is published by Lehman ''et al.'' (2019). * High-precision dating for the

Battle Formation The Battle Formation is a Formation (geology), geologic formation of Late Cretaceous (Maastrichtian) age in the Western Canadian Sedimentary Basin, Western Canada Sedimentary Basin. It is present throughout much of the central Alberta plains, wh ...

(

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) is presented by Eberth & Kamo (2019). * High-precision dating and the first calibrated

chronostratigraphy Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time. The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geologica ...

for the

Horseshoe Canyon Formation The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller. The Horseshoe Canyon Formation is part of the ...

(

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

,

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

) is presented by Eberth & Kamo (2019). * A study on the

Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...

climate of Arctic

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

, based on data from the

Prince Creek Formation The Prince Creek Formation is a geological formation in Alaska with strata dating to the Early Maastrichtian stage of the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et a ...

, is published by Salazar-Jaramillo ''et al.'' (2019). * Studies on the timing of the

Deccan Traps The Deccan Traps is a large igneous province of west-central India (17–24°N, 73–74°E). It is one of the largest volcanic features on Earth, taking the form of a large shield volcano. It consists of numerous layers of solidified flood ...

volcanism close to the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

-

Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...

boundary are published by Schoene ''et al.'' (2019), who interpret their findings as indicative of four high-volume eruptive periods close to the Cretaceous-Paleogene boundary, the first of which occurred tens of thousands of years prior to both the Chicxulub bolide impact and

Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...

and by Sprain ''et al.'' (2019), who interpret their findings as indicating that a steady eruption of the flood basalts mostly occurred in the earliest Paleogene. * A study on the environmental variability before and across the Cretaceous-Paleogene mass extinction, as inferred from data on the calcium isotope ratios of

aragonitic Aragonite is a carbonate mineral, one of the three most common naturally occurring crystal forms of calcium carbonate, (the other forms being the minerals calcite and vaterite). It is formed by biological and physical processes, including pre ...

mollusc shells from the

Lopez de Bertodano Formation The Lopez de Bertodano Formation is a geological formation in the James Ross archipelago of the Antarctic Peninsula. The strata date from the end of the Late Cretaceous (upper-lower Maastrichtian stage) to the Danian stage of the lower Paleocene ...

(

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...

), is published online by Linzmeier ''et al.'' (2019). * A turbulently deposited sediment package directly overlain by the Cretaceous–Paleogene boundary tonstein is reported from the Tanis site (

Hell Creek Formation The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of ...

,

North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the Native Americans in the United States, indigenous Dakota people, Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north a ...

,

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

) by DePalma ''et al.'' (2019), who interpret their findings as indicating that deposition occurred shortly after a major bolide impact, and might have been caused by the

Chicxulub The Chicxulub crater () is an impact crater buried underneath the Yucatán Peninsula in Mexico. Its center is offshore near the community of Chicxulub, after which it is named. It was formed slightly over 66 million years ago when a large a ...

impact. * A study on the immediate aftermath of the Chicxulub impact at the Cretaceous–Paleogene boundary, based on data from the

Chicxulub crater The Chicxulub crater () is an impact crater buried underneath the Yucatán Peninsula in Mexico. Its center is offshore near the community of Chicxulub, after which it is named. It was formed slightly over 66 million years ago when a large ast ...

, is published by Gulick ''et al.'' (2019). * Evidence of rapid ocean acidification in the aftermath of the Chicxulub impact and of the protracted Earth system recovery after the Cretaceous–Paleogene extinction event is presented by Henehan ''et al.'' (2019). * The longest, highest resolution, stratigraphically continuous, single-species

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular Ectoplasm (cell biology), ectoplasm for catching food and ot ...

l carbon and oxygen isotope records for the Late

Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...

to Early

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

from a single site in the South

Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

, providing information on the evolution of climate and carbon-cycling during this time period, are presented by Barnet ''et al.'' (2019). * O'Leary ''et al.'' (2019) publish a monograph on the sedimentology and sequence stratigraphy of the part of

Mali Mali (; ), officially the Republic of Mali,, , ff, 𞤈𞤫𞤲𞥆𞤣𞤢𞥄𞤲𞤣𞤭 𞤃𞤢𞥄𞤤𞤭, Renndaandi Maali, italics=no, ar, جمهورية مالي, Jumhūriyyāt Mālī is a landlocked country in West Africa. Mali ...

which was covered by an ancient

epeiric sea An inland sea (also known as an epeiric sea or an epicontinental sea) is a continental body of water which is very large and is either completely surrounded by dry land or connected to an ocean by a river, strait, or "arm of the sea". An inland se ...

known as the Trans-Saharan Seaway during the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

and early

Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...

, provide the first formal description of and nomenclature for the Upper Cretaceous and lower Paleogene geological formations of this region, and revise fossil flora and fauna of this region. * Zeebe & Lourens (2019) provide a new absolute

astrochronology Astrochronology is the dating of sedimentary units by calibration with astronomically tuned timescales, such as Milankovic cycles, or even sunspot cycles. When used in concert with radiometric dating Radiometric dating, radioactive dating or r ...

up to 58 Ma and a new Paleocene–Eocene boundary age. * A study on

stoma In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...

ta of fossil specimens of members of the family

Lauraceae Lauraceae, or the laurels, is a plant family that includes the true laurel and its closest relatives. This family comprises about 2850 known species in about 45 genera worldwide (Christenhusz & Byng 2016 ). They are dicotyledons, and occur ma ...

from the

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

of

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

and

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

, evaluating their implications for reconstructions of Eocene ''p''CO₂ levels, is published by Steinthorsdottir ''et al.'' (2019). * Climate simulations capturing major climatic features of the Early Eocene and the

Paleocene–Eocene Thermal Maximum The Paleocene–Eocene thermal maximum (PETM), alternatively (ETM1), and formerly known as the "Initial Eocene" or "", was a time period with a more than 5–8 °C global average temperature rise across the event. This climate event o ...

in a state-of-the-art Earth system model are presented by Zhu, Poulsen & Tierney (2019). * A study evaluating the utility of membrane

lipid Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids include ...

s of members of Thaumarchaeota (now

Nitrososphaerota The Nitrososphaerota (syn. Thaumarchaeota) are a phylum of the Archaea proposed in 2008 after the genome of ''Cenarchaeum symbiosum'' was sequenced and found to differ significantly from other members of the hyperthermophilic phylum Thermoprote ...

) as proxies for the carbon isotope excursion and surface ocean warming, and assessing their implications for the knowledge of the source and size of carbon emissions during the Paleocene–Eocene Thermal Maximum, is published by Elling ''et al.'' (2019). * A study on abundant black charcoal shards from Paleogene sites of Wilson Lake B (

New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delaware ...

) and Randall's Farm (

Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...

) is published by Fung ''et al.'' (2019), who interpret these shards as most likely to be evidence of widespread wildfires at the Paleocene-Eocene boundary caused by extraterrestrial impact. * A study on the impact of carbon-based greenhouse gas fluxes associated with the

North Atlantic Igneous Province The North Atlantic Igneous Province (NAIP) is a large igneous province in the North Atlantic, centered on Iceland. In the Paleogene, the province formed the Thulean Plateau, a large basaltic lava plain, which extended over at least in area and in ...

on the onset of the Paleocene–Eocene Thermal Maximum is published by Jones ''et al.'' (2019). * Evidence from the Deep Ivorian Basin offshore West Africa (equatorial

Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...

), indicating that peak warming during the Middle Eocene Climatic Optimum was associated with upper-ocean

stratification Stratification may refer to: Mathematics * Stratification (mathematics), any consistent assignment of numbers to predicate symbols * Data stratification in statistics Earth sciences * Stable and unstable stratification * Stratification, or st ...

, decreased

export production In the deep ocean, marine snow (also known as "ocean dandruff") is a continuous shower of mostly organic detritus falling from the upper layers of the water column. It is a significant means of exporting energy from the light-rich photic zone t ...

, and possibly harmful algal blooms, is presented by Cramwinckel ''et al.'' (2019). * New stable isotopes record of the Middle Eocene Climatic Optimum event is reported from eastern

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...

by Giorgioni ''et al.'' (2019). * A study on variations of ocean circulation and marine bioproductivity related to the beginnings of the formation of the

Antarctic Circumpolar Current The Antarctic Circumpolar Current (ACC) is an ocean current that flows clockwise (as seen from the South Pole) from west to east around Antarctica. An alternative name for the ACC is the West Wind Drift. The ACC is the dominant circulation feat ...

, based on data from Eocene and Oligocene sedimentary drift deposits east of

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

, is published by Sarkar ''et al.'' (2019). * A study on changes in surface water temperature in the eastern

North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian S ...

Basin during the late

Priabonian The Priabonian is, in the ICS's geologic timescale, the latest age or the upper stage of the Eocene Epoch or Series. It spans the time between . The Priabonian is preceded by the Bartonian and is followed by the Rupelian, the lowest stage of t ...

to earliest

Rupelian The Rupelian is, in the geologic timescale, the older of two ages or the lower of two stages of the Oligocene Epoch/Series. It spans the time between . It is preceded by the Priabonian Stage (part of the Eocene) and is followed by the Chattian ...

is published by Śliwińska ''et al.'' (2019). * A study linking the onset or strengthening of an

Atlantic meridional overturning circulation The Atlantic meridional overturning circulation (AMOC) is part of a global thermohaline circulation in the oceans and is the zonally integrated component of surface and deep currents in the Atlantic Ocean. It is characterized by a northward fl ...

to the closure of the Arctic–Atlantic gateway at the Eocene–Oligocene transition is published by Hutchinson ''et al.'' (2019). * A study on the timing of the uplift of the

Tibetan Plateau The Tibetan Plateau (, also known as the Qinghai–Tibet Plateau or the Qing–Zang Plateau () or as the Himalayan Plateau in India, is a vast elevated plateau located at the intersection of Central, South and East Asia covering most of the Ti ...

, as indicated by the discovery of the

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...

fossils in the Lunpola Basin in Tibet, is published by Su ''et al.'' (2019). * A review of vertebrate fossils from the Tibetan Plateau, evaluating their implications for inferring the course of the uplift of the Tibetan Plateau, is published by Deng ''et al.'' (2019). * A study on the impact of changing Eocene paleogeography and climate on the utility of stable isotope paleoaltimetry methods in the studies aiming to reconstruct the elevation history of the Tibetan Plateau is published by Botsyun ''et al.'' (2019). * A study on the causes of the long-term climate cooling during the

Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...

is published by Rugenstein, Ibarra & von Blanckenburg (2019). * A study on the climatic and environmental conditions in the Loperot site (

Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...

) in the early

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

is published by Liutkus-Pierce ''et al.'' (2019). * A study on the timing and course of the separation of the

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by th ...

and the

Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ea ...

in the Miocene is published by Bialik ''et al.'' (2019). * A study comparing changes of the export of intermediate-depth Pacific waters to the western North Atlantic prior to the closure of the

Central American Seaway The Central American Seaway (also known as the Panamanic Seaway, Inter-American Seaway and Proto-Caribbean Seaway) was a body of water that once separated North America from South America. It formed during the Jurassic (200–154 Ma) during the br ...

with records of strength of the

Atlantic meridional overturning circulation The Atlantic meridional overturning circulation (AMOC) is part of a global thermohaline circulation in the oceans and is the zonally integrated component of surface and deep currents in the Atlantic Ocean. It is characterized by a northward fl ...

, evaluating the implications of this data for the knowledge of the timing of closure of the Central American Seaway, is published by Kirillova ''et al.'' (2019). * A study on climatic and environmental changes in central

Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...

during the late Miocene is published by Carrapa, Clementz & Feng (2019). * A study on the exact age of the marine fauna from the Miocene Chilcatay and

Pisco Pisco is a colorless or yellowish-to-amber colored brandy produced in winemaking regions of Peru and Chile. Made by distilling fermented grape juice into a high-proof spirit, it was developed by 16th-century Spanish settlers as an alternative t ...

formations (

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

), and on its implications for reconstructions of local paleoenvironment, is published online by Bosio ''et al.'' (2019). * A study on the origin of the African C₄ savannah grasslands is published by Polissar ''et al.'' (2019). * A study on the anatomical traits of teeth and inferred diet of

bovids The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, the ...

, suids and rhinocerotids from Kanapoi, and on their implications for reconstructing the environments of this site, is published online by Dumouchel & Bobe (2019). * New spatial data on the Plio-Pleistocene Bolt's Farm pits from the

Cradle of Humankind The Cradle of Humankind is a paleoanthropological site and is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest concentration of ...

site (

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

) is presented by Edwards ''et al.'' (2019), who also attempt to provide key biochronological ages for the Bolt's Farm deposits. * A study on the global mean sea level during the Pliocene mid-

Piacenzian The Piacenzian is in the international geologic time scale the upper stage (stratigraphy), stage or latest age (geology), age of the Pliocene. It spans the time between 3.6 ± 0.005 year#SI prefix multipliers, Ma and 2.588 ± 0.005 Ma (million yea ...

Warm Period is published by Dumitru ''et al.'' (2019). * A study on the amplitude of sea-level variations during the Pliocene is published by Grant ''et al.'' (2019). * Simulations of coevolution of climate, ice sheets and

carbon cycle The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and Earth's atmosphere, atmosphere of the Earth. Carbon is the main component of biological compounds as well as ...

over the past 3 million years are presented by Willeit ''et al.'' (2019). * A study on the age of the

Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...

, as indicated by data from Pliocene and Pleistocene

paleosol In the geosciences, paleosol (''palaeosol'' in Great Britain and Australia) is an ancient soil that formed in the past. The precise definition of the term in geology and paleontology is slightly different from its use in soil science. In geolo ...

s from the

Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...

, is published by Muhs ''et al.'' (2019). * A study on the latest

Villafranchian Villafranchian age ( ) is a period of geologic time (3.5–1.0 Ma) spanning the Late Pliocene and Early Pleistocene used more specifically with European Land Mammal Ages. Named by Italian geologist Lorenzo Pareto for a sequence of terrestrial s ...

climate and environment of the area of southern

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

, as indicated by amphibian and reptile fossil record from the Pirro Nord karstic complex, is published by Blain ''et al.'' (2019). * A study on atmospheric gas levels before and after the shift from glacial cycles of 100 thousand years to 40-thousand-year cycles around one million years ago, as inferred from data from ice core samples from the Allan Hills Blue Ice Area (East Antarctica), is published by Yan ''et al.'' (2019). * A study on pCO₂ levels from 2.6 to 0.8 Ma is published by Da ''et al.'' (2019), who find no evidence indicating that the

Mid-Pleistocene Transition The Mid-Pleistocene Transition (MPT), also known as the Mid-Pleistocene Revolution (MPR), is a fundamental change in the behaviour of glacial cycles during the Quaternary glaciations. The transition happened approximately 1.25–0.7 milli ...

was caused by the decline of pCO₂. * A study on changes in winter rainfall in the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

over the past 1.36 million years is published by Wagner ''et al.'' (2019). * Results of stable carbon and oxygen isotope analyses of

tooth enamel Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The ...

samples from

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

mammals from the Yugong Cave and Baxian Cave (

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

) are presented by Sun ''et al.'' (2019), who evaluate the implications of their findings for the knowledge of Pleistocene climatic and environmental changes in South China. * A study on Pleistocene mammal fossils from the Yai Ruak Cave ( Krabi Province,

Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...

), including the southernmost known record of '' Crocuta crocuta ultima'', is published by Suraprasit ''et al.'' (2019), who evaluate the implications of these fossils for reconstructions of the environment in the area of the

Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area ...

in the Pleistocene. * A study on

Acheulean Acheulean (; also Acheulian and Mode II), from the French ''acheuléen'' after the type site of Saint-Acheul, is an archaeological industry of stone tool manufacture characterized by the distinctive oval and pear-shaped "hand axes" associated ...

and Middle Stone Age sites from the

Eastern Desert The Eastern Desert (Archaically known as Arabia or the Arabian Desert) is the part of the Sahara desert that is located east of the Nile river. It spans of North-Eastern Africa and is bordered by the Nile river to the west and the Red Sea and ...

(

Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...

), preserving stone artifacts, is published by Masojć ''et al.'' (2019), who interpret these sites as evidence of green corridor or corridors across Sahara which made early hominin dispersal possible. * Evidence from oxygen isotope data from

Soreq Cave Avshalom Cave ( he, מערת אבשלום, Me'arat Avshalom), known in academic literature as Soreq Cave ( he, מערת שׂורק, Me'arat Soreq), ar, مغارة سوريك, Mghar Suriq) an popularly as Stalactites Cave ( he, מערת הנטי� ...

speleothem A speleothem (; ) is a geological formation by mineral deposits that accumulate over time in natural caves. Speleothems most commonly form in calcareous caves due to carbonate dissolution reactions. They can take a variety of forms, depending on ...

s (

Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...

), indicative of the occurrence of summer monsoon rainfall in the Middle East during recurrent intervals of the last interglacial period (overlapping with archeological indicators of human migration), is presented by Orland ''et al.'' (2019). * A study on the spatial and temporal distribution of ancient

peatland A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...

s in the past 130,000 years is published by Treat ''et al.'' (2019). * A study on the size of fossil rabbits from 14 late Pleistocene and Holocene archaeological sites in

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

, and on its implications for the knowledge of temperatures and environment in the area of Portugal during the last glaciation, is published by Davis (2019). * A study on Pleistocene small mammal remains from Stratigraphic Unit V from El Salt site (

Alcoy, Spain Alcoy ( ca-valencia, Alcoi) is an industrial and university city, region and municipality located in the Valencian Community, Spain. The Serpis river crosses the municipal boundary of Alcoy. The local authority reported a population of 61,135 res ...

), evaluating their implications for the knowledge of climatic conditions in the eastern

Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...

at the time of the disappearance of local

Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While th ...

populations during Marine Isotope Stage 3, is published by Fagoaga ''et al.'' (2019). * A study on the sedimentary sequence from the Pilauco site in

Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...

, evaluating whether evidence from this site is consistent with the

Younger Dryas impact hypothesis The Younger Dryas impact hypothesis (YDIH) or Clovis comet hypothesis is a speculative attempt to explain the onset of the Younger Dryas (YD) as an alternative to the long standing and widely accepted cause due to a significant reduction or shut ...

, is published by

Pino Pino or Piño may refer to: People * Danny Pino (born 1974), American actor * Domenico Pino (1760–1826), Italian general of the Napoleonic Wars * Fernando Solanas (born 1936), aka "Pino" Solanas, Argentine filmmaker * Frank J. Pino (1909–200 ...

''et al.'' (2019). * A study on variations of size of fossil

murine The Old World rats and mice, part of the subfamily Murinae in the family Muridae, comprise at least 519 species. Members of this subfamily are called murines. In terms of species richness, this subfamily is larger than all mammal families excep ...

rodents from

Liang Bua Liang Bua is a limestone cave on the island of Flores, Indonesia, slightly north of the town of Ruteng in Manggarai Regency, East Nusa Tenggara. The cave demonstrated archaeological and paleontological potential in the 1950s and 1960s as describe ...

(

Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Including the Komodo Islands off its west coast (but excluding the Solor Archipelago to the east of Flores), the land area is 15,530.58 km2, and th ...

,

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...

) through time, and on their implications for reconstructions of paleoclimate and paleoenvironment of Flores, is published by Veatch ''et al.'' (2019). * A study on human land use worldwide from 10,000 years before the present to 1850 CE, indicating that Earth was to a large extent transformed by human activity by 3000 years ago, is published by Stephens ''et al.'' (2019). * Evidence for synchronous cyclical changes in monsoon climate, human activity and prehistoric cultural development in the area of northeast

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

throughout the Holocene is presented by Xu ''et al.'' (2019). * A study on Andean

plate tectonics Plate tectonics (from the la, label=Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large ...

since the late Mesozoic is published by Chen, Wu & Suppe (2019). * A study on the course of the

collision In physics, a collision is any event in which two or more bodies exert forces on each other in a relatively short time. Although the most common use of the word ''collision'' refers to incidents in which two or more objects collide with great fo ...

of India and Asia, as indicated by

palaeomagnetic Paleomagnetism (or palaeomagnetismsee ), is the study of magnetic fields recorded in rocks, sediment, or archeological materials. Geophysicists who specialize in paleomagnetism are called ''paleomagnetists.'' Certain magnetic minerals in rocks ...

data from the Burma Terrane, is published by Westerweel ''et al.'' (2019). * A scenario for the genesis of tropical cyclones throughout the Cenozoic is presented by Yan ''et al.'' (2019). * A study on the extent of ice sheets in the Northern Hemisphere throughout the Quaternary is published by Batchelor ''et al.'' (2019). * A new method of concentration of proteins from fossil specimens with high humic content and of removal of humic substances is presented by Schroeter ''et al.'' (2019).

References

* {{reflist 2019 in paleontology,

Flora

Plants

Fungi

Paleomycological research

Sponges

Research

New taxa

Cnidarians

Research

New taxa

Arthropods

Bryozoans

Brachiopods

Molluscs

Echinoderms

Research

New taxa

Conodonts

Research

New taxa

Fishes

Amphibians

Reptiles

Synapsids

Non-mammalian synapsids

Research

New taxa

Mammals

Other animals

New taxa

Research

Foraminifera

Research

New taxa

Other organisms

New taxa

Research

Trace fossils

History of life in general

Other research

References