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Prosomapoda
Prosomapoda is a clade of Chelicerata, euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising Bunodidae, bunodids, Pseudoniscidae, pseudoniscids, Chasmataspidida, chasmataspidids, Eurypterida, eurypterids and Arachnida, arachnids), as well as several basal Synziphosurina, synziphosurid genera. The clade is defined by the lack of exopods (outer branches) of prosomal appendage II-V in the adult instar, where in contrast the exopods of appendage II-V are well-developed in the non-prosomapod euchelicerates ''Offacolus'' and ''Dibasterium''. References

Prosomapoda, Middle Ordovician first appearances {{Chelicerata-stub ...
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Borchgrevinkium
''Borchgrevinkium'' is an extinct genus of chelicerate arthropod. A fossil of the single and type species, ''B. taimyrensis'', has been discovered in Deposition (geology), deposits of the Early Devonian, Devonian period (Lochkovian, Lochkovian epoch) in the Krasnoyarsk Krai, Siberia, Russia. The name of the genus honors Carsten Borchgrevink, an Anglo-Norwegian explorer who participated in many expeditions to Antarctica. ''Borchgrevinkium'' represents a poorly known genus whose Affinity (taxonomy), affinities are uncertain. It had several unique characteristics that differentiated it from many other arthropods, such as its long parabolic (nearly U-shaped) prosoma (head), its elongated first and second Segmentation (biology), segments and the presence of paired "ridges" in the surface of its third to tenth tergites (Dorsal (anatomy), dorsal halves of the segments). Furthermore, the opisthosoma (the "trunk") of ''Borchgrevinkium'' was triangular, and its telson (the posteriormost di ...
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Prosomapoda
Prosomapoda is a clade of Chelicerata, euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising Bunodidae, bunodids, Pseudoniscidae, pseudoniscids, Chasmataspidida, chasmataspidids, Eurypterida, eurypterids and Arachnida, arachnids), as well as several basal Synziphosurina, synziphosurid genera. The clade is defined by the lack of exopods (outer branches) of prosomal appendage II-V in the adult instar, where in contrast the exopods of appendage II-V are well-developed in the non-prosomapod euchelicerates ''Offacolus'' and ''Dibasterium''. References

Prosomapoda, Middle Ordovician first appearances {{Chelicerata-stub ...
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Chelicerata
The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and '' Mollisonia'' present by this time. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered in a cuti ...
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Offacolus
''Offacolus'' is an extinct genus of euchelicerate, a group of chelicerate arthropods. Its only species, ''O. kingi'', has been found in deposits from the Silurian period ( Homerian epoch) in the Wenlock Series Lagerstätte of Herefordshire, England. The genus is named after Offa, a king from the ancient kingdom of Mercia, and ''colus'', a person who dwelled among (this time referring to) the Offa's Dyke. The species name honors Robert Joseph King, a British mineralogist who found the fossils of ''Offacolus''. Similar to ''Dibasterium'', ''Offacolus'' possess limb-like exopods (outer limb branches) on appendage II to V, a character suggest to be plesiomorphic (observable in the putative stem-chelicerate taxon Habeliida) and lost within the prosomapod clade. Classification ''Offacolus'' was originally described as an arthropod with chelicerate affinities, with detailed redescription done by Sutton ''et al.'' 2002 further suggested it to be unambiguously a chelicerate arthro ...
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Eurypterida
Eurypterids, often informally called sea scorpions, are a group of extinct marine arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period, 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction e ...
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Arachnid
Arachnids are arthropods in the Class (biology), class Arachnida () of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, opiliones, harvestmen, Solifugae, camel spiders, Amblypygi, whip spiders and Uropygi, vinegaroons. Adult arachnids have eight Arthropod leg, legs attached to the cephalothorax. In some species the frontmost pair of legs has converted to a sensory function, while in others, different appendages can grow large enough to take on the appearance of extra pairs of legs. Almost all Extant taxon, extant arachnids are terrestrial animal, terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 110,000 named species, of which 51,000 are species of spiders. The term is derived from the Ancient Greek, Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, w ...
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Planaterga
Planaterga is a clade of prosomapod euchelicerates including several synziphosurid genera (mainly bunodids and pseudoniscids) and the group Dekatriata (which in turn includes arachnids, chasmataspidids and eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct marine arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period, 467.3 Myr, mil ...s). Planaterga is defined by the opisthosoma with tergites broadest at third or fourth and lacking enlarged axial nodes, carapace (prosomal dorsal shield) with reduced genal spines, as well as somite VII (first opisthosomal segment) with reduced appendages and microtergite. References Middle Ordovician first appearances {{Chelicerata-stub ...
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Arachnida
Arachnids are arthropods in the class Arachnida () of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Adult arachnids have eight legs attached to the cephalothorax. In some species the frontmost pair of legs has converted to a sensory function, while in others, different appendages can grow large enough to take on the appearance of extra pairs of legs. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 110,000 named species, of which 51,000 are species of spiders. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Morphology Almost all adult arachnids have eight legs, unlike adult insects whic ...
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Cryptocellus Goodnighti
''Cryptocellus goodnighti'' is an arachnid species in the genus '' Cryptocellus''. It occurs in Costa Rica.On the female of Cryptocellus goodnighti Arachnida: Ricinulei. Norman I. Platnick, The Journal of Arachnology, 1993, Volume 21, no. 1, pages 79-80article Retrieved 26 March 2016 References External links * * Arthropods of Central America Ricinulei Animals described in 1981 {{Arachnid-stub ...
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Ricinulei
Ricinulei is a small order of arachnids. Like most arachnids, they are predatory; eating small arthropods. They occur today in west-central Africa ('' Ricinoides'') and the Americas ('' Cryptocellus'' and '' Pseudocellus'') from Brazil to southern Texas, where they inhabit either leaf-litter or caves. As of 2022, 103 extant species of ricinuleids have been described worldwide, all in the single family Ricinoididae. Due to their obscurity they do not have a proper common-name, though in academic literature they are occasionally referred to as hooded tickspiders. In addition to the three living genera, Ricinulei has a fossil-record spanning over 300 million years, including fossils from the Late Carboniferous of Euramerica and the Cretaceous Burmese amber. Anatomy and physiology The most important general account of ricinuleid anatomy remains the 1904 monograph by Hans Jacob Hansen and William Sørensen. Useful further studies can be found in, e.g., the work of Pittard and Mitche ...
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Synziphosurina
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded from the monophyletic Xiphosura ''sensu stricto'' and only regarded as horseshoe crabs under a broader sense ('Xiphosura' ''sensu lato''). Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata. File:20200920 Synziphosurina dorsal segmentation.png, Dorsal morphology and variation of opisthosomal segmentation across synziphosurines. While ''Weinbergina'' and '' Willwerathia'' being exceptionally large, most synziphosurines are small arthropods with body length ranging only about a few centimeters long. The body of synziphosurine composed of a prosoma covered by a dome-like carapace (prosomal dorsal shield) and an opisthosoma with usually unfused 9-11 segments expr ...
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Xiphosura
Xiphosura (; , in reference to its sword-like telson) is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs (a name applied more specifically to the only extant family, Horseshoe crab, Limulidae). They first appeared in the Hirnantian (Late Ordovician). Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera. The group has hardly changed in appearance in hundreds of millions of years; the modern horseshoe crabs look almost identical to prehistoric genera and are considered to be living fossils. The most notable difference between ancient and modern forms is that the abdominal segments in present species are fused into a single unit in adults. Xiphosura were historically placed in the class Merostomata, although this term was intended to encompass also the Eurypterida, eurypterids, whence it denoted what is now thought to be an unnatural (paraphyletic) group (although this is a gr ...
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