Zemaciinae
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Borsoniidae is a
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of small to medium-sized
sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
s, marine
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s in the superfamily
Conoidea Conoidea is a Taxonomic rank, superfamily of predatory sea snails, marine (ocean), marine gastropod Mollusc, mollusks within the suborder Hypsogastropoda. This superfamily is a very large group of marine mollusks, estimated at 340 recent valid gen ...
.Bouchet, P. (2011). Borsoniidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=153870 on 2011-08-12Bouchet P. & Rocroi J.-P. (Ed.) (2005). "Classification and nomenclator of gastropod families". ''
Malacologia ''Malacologia'' is a peer-reviewed scientific journal in the field of malacology, the study of mollusks. The journal publishes articles in the fields of molluscan systematics, ecology, population ecology, genetics, molecular genetics, evolution, an ...
'' 47(1-2). . 397 pp.
In 2011, Bouchet, Kantor ''et al''. brought genera from the subfamilies
Clathurellinae Clathurellinae was a subfamily of small to quite large sea snails, marine gastropod mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant specie ...
and
Raphitominae Raphitominae was a Family (biology), subfamily of small to quite large sea snails, marine (ocean), marine gastropod mollusks in the superfamily Conoidea. Philippe Bouchet, Bouchet, Kantor ''et al''. elevated in 2011 the subfamily Raphitominae (wh ...
(they were previously placed in the family Conidae), as well as genera from the subfamily Zemaciinae (at that point belonging to the family
Turridae Turridae is a Taxonomy (biology), taxonomic family (biology), family name for a number of predatory sea snails, Marine (ocean), marine gastropod mollusks in the superfamily Conoidea.MolluscaBase (2018). Turridae H. Adams & A. Adams, 1853 (1838). ...
), together to form the family Borsoniidae. This re-arrangement was based on anatomical characters and a dataset of molecular sequences of three
gene In biology, the word gene has two meanings. The Mendelian gene is a basic unit of heredity. The molecular gene is a sequence of nucleotides in DNA that is transcribed to produce a functional RNA. There are two types of molecular genes: protei ...
fragmentsBouchet, P.; Kantor, Y. I.; Sysoev, A.; Puillandre, N. (2011). ''A new operational classification of the Conoidea''. Journal of Molluscan Studies. 77, 273-308
/ref>


Description

This family is a rather heterogenous group, with wide-ranging varieties in their properties. The medium- to large-sized shells are fusiform to biconic in shape. Their size has a wide range (between 5 mm and 80 mm). The shell can be longitudinally coarsely ribbed, but axial ribs are sometimes obsolete to absent. The smooth
columella Lucius Junius Moderatus Columella (, Arabic: ) was a prominent Roman writer on agriculture in the Roman Empire. His in twelve volumes has been completely preserved and forms an important source on Roman agriculture and ancient Roman cuisin ...
has a strong (one to three plicae) to obsolete plication upon the middle. This plication varies within genera. In the genus '' Cordiera'' there are two columellar plaits. The
aperture In optics, the aperture of an optical system (including a system consisting of a single lens) is the hole or opening that primarily limits light propagated through the system. More specifically, the entrance pupil as the front side image o ...
is elliptical to oval in shape and it has a short to moderately long (e.g. '' Zemacies excelsa'')
siphonal canal The siphonal canal is an anatomical feature of the shells of certain groups of sea snails within the clade Neogastropoda. Some sea marine gastropods have a soft tubular anterior extension of the mantle called a siphon through which water i ...
and a deep anal sinus. The anterior canal has a moderate length and is slightly twisted to the left. The thin outer lip is arcuate. The vestigial or fully developed operculum has a terminal nucleus, and may be missing in some species. The
radula The radula (; : radulae or radulas) is an anatomical structure used by mollusks for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food enters ...
is absent in the genus ''
Zemacies ''Zemacies'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae Most species in this genus are extinct. Their age range is 55.8 to 11.608 Ma. Species Species within the genus ''Zemacies'' include: * † '' Zemacies a ...
''. McLean, J.H. (1971) ''A revised classification of the family Turridae, with the proposal of new subfamilies, genera, and subgenera from the Eastern Pacific''. The Veliger, 14, 114–130
/ref>


Genera

* '' Antarctospira'' Kantor, Harasewych & Puillandre, 2016 * ''
Apaturris ''Apaturris'' is a genus of sea snails in the family Borsoniidae Borsoniidae is a monophyletic family of small to medium-sized sea snails, marine gastropod mollusks in the superfamily Conoidea.Bouchet, P. (2011). Borsoniidae. Accessed throug ...
'' Iredale, 1917 * ''
Aphanitoma ''Aphanitoma'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae.Bouchet, P.; Gofas, S. (2015). Aphanitoma Bellardi, 1875. In: MolluscaBase (2015). Accessed through: World Register of ...
'' Bellardi, 1875 * † '' Asthenotoma'' Harris & Burrows, 1891 * '' Austroturris'' Laseron, 1954 * '' Awateria'' Suter, 1917 * ''
Bathytoma ''Bathytoma'' is a genus of deep-water sea snails, marine gastropod mollusks in the family (biology), family Borsoniidae. Fossil records This genus is known in the fossil records from the Paleocene to the Quaternary (age range: 55.8 to 0.781 mi ...
'' Harris & Burrows, 1891 * † '' Belatomina'' Powell, 1944 * ''
Belaturricula ''Belaturricula'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae. Species Species within the genus ''Belaturricula'' include: * ''Belaturricula dissimilis'' (Watson, 1886) * ''Bela ...
'' Powell, 1951 * † '' Boettgeriola'' Wenz, 1943 * '' Borsonella'' Dall, 1908 * ''
Borsonellopsis ''Borsonellopsis'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae. Description In contrast with Borsonella, the shells in this genus are larger in size, have a weakly plicate columella and have a vestigial opercu ...
'' McLean, 1971 * '' Borsonia'' Bellardi, 1839 * ''
Cordieria ''Cordieria'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae.Bouchet, P. (2015). Cordieria. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=ta ...
'' Rouault, 1848 * ''
Darbya ''Darbya'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae.Bouchet, P. (2015). Darbya Bartsch, 1934. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at http://marinespecies.org/aphia.php? ...
'' Bartsch, 1934 * ''
Diptychophlia ''Diptychophlia'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae.Rosenberg, G.; Bouchet, P. (2015). Diptychophlia Berry, 1964. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at http://w ...
'' Berry, 1964 * ''
Drilliola ''Drilliola'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae, the Conus, cone snails and their allies. Species Species within the genus ''Drilliola'' include: * ''Drilliola annosa ...
'' Locard, 1897 * ''
Filodrillia ''Filodrillia'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae, the Conus, cone snails and their allies.Bouchet, P. (2015). Filodrillia Hedley, 1922. In: MolluscaBase (2015). Access ...
'' Hedley, 1922 * '' Genota'' H. Adams & A. Adams, 1853 * ''
Glyptaesopus ''Glyptaesopus'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae. Description The small shell is slender and contains 5½ sculptured whorls with weakly incised sutures. The aperture is long and narrow and has a th ...
'' Pilsbry & Olsson, 1941 * ''
Heteroturris ''Heteroturris'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family BorsoniidaeBouchet, P. (2015). Heteroturris Powell, 1967. In: MolluscaBase (2015). Accessed through: World Register of Marine Spec ...
'' Powell, 1967 * '' Maoritomella'' Powell, 1942 * ''
Microdrillia ''Microdrillia'' is a genus of very small sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae.Bouchet, P.; Gofas, S. (2015). Microdrillia Casey, 1903. In: MolluscaBase (2015). Accessed through: World ...
'' Casey, 1903 * '' Ophiodermella'' Bartsch, 1944 * '' Paraborsonia'' Pilsbry, 1922 * '' Phenatoma'' Finlay, 1924 * † '' Pseudotoma'' Bellardi, 1875 * '' Pulsarella'' Laseron, 1954 * ''
Retidrillia ''Retidrillia'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae Borsoniidae is a monophyletic family of small to medium-sized sea snails, marine gastropod mollusks in the superfamily Conoidea.Bouchet, P. (2011) ...
'' McLean, 2000 * '' Suavodrillia'' Dall, 1918 * '' Tomopleura'' Casey, 1904 * '' Tropidoturris'' Kilburn, 1986 * ''
Typhlodaphne ''Typhlodaphne'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae. Description The smooth protoconch is bluntly rounded and paucispiral, consisting of two whorls. The sinus is subsutural, steeply descending and then ...
'' Powell, 1951 * ''
Typhlomangelia ''Typhlomangelia'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae, the Conus, cone snails and their allies. Species Species within the genus ''Typhlomangelia'' include: * ''Typhlom ...
'' G.O. Sars, 1878 * ''
Zemacies ''Zemacies'' is a genus of sea snails, marine gastropod mollusks in the family Borsoniidae Most species in this genus are extinct. Their age range is 55.8 to 11.608 Ma. Species Species within the genus ''Zemacies'' include: * † '' Zemacies a ...
'' Finlay, 1926 ;Genera brought into synonymy : * ''Acamptogenotia'' Rovereto, 1899: synonym of † ''Pseudotoma'' Bellardi, 1875 (junior objective synonym, unnecessary substitute name for ''Pseudotoma'', by Rovereto believed to be a junior homonym of "Pseudotoma Stephens, 1852", obviously an error for ''Pseudotomia'' Stephens, 1829 epidoptera) * ''Acrobela'' Thiele, 1925 : synonym of ''
Microdrillia ''Microdrillia'' is a genus of very small sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae.Bouchet, P.; Gofas, S. (2015). Microdrillia Casey, 1903. In: MolluscaBase (2015). Accessed through: World ...
'' Casey, 1903 * ''Acropota'' F. Nordsieck, 1977: synonym of ''
Microdrillia ''Microdrillia'' is a genus of very small sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae.Bouchet, P.; Gofas, S. (2015). Microdrillia Casey, 1903. In: MolluscaBase (2015). Accessed through: World ...
'' Casey, 1903 * ''Boettgeria'' Peyrot, 1931 : synonym of ''Boettgeriola'' Wenz, 1943 * ''Boettgeriola'' Wenz, 1943 : synonym of '' Borsonia'' Bellardi, 1839 * ''Genotia'' P. Fischer, 1883 : synonym of '' Genota'' H. Adams & A. Adams, 1853 * ''Micantapex'' Iredale, 1936 : synonym of ''
Bathytoma ''Bathytoma'' is a genus of deep-water sea snails, marine gastropod mollusks in the family (biology), family Borsoniidae. Fossil records This genus is known in the fossil records from the Paleocene to the Quaternary (age range: 55.8 to 0.781 mi ...
'' Harris & Burrows, 1891 * ''Narraweena (gastropod)'' Laseron, 1954 : synonym of '' Maoritomella'' Powell, 1942 * ''Oligotoma'' Bellardi, 1875 : synonym of '' Asthenotoma'' Harris & Burrows, 1891 * ''Parabathytoma'' Shuto, 1961 : synonym of ''
Bathytoma ''Bathytoma'' is a genus of deep-water sea snails, marine gastropod mollusks in the family (biology), family Borsoniidae. Fossil records This genus is known in the fossil records from the Paleocene to the Quaternary (age range: 55.8 to 0.781 mi ...
'' Harris & Burrows, 1891 * ''Riuguhdrillia'' Oyama, 1951 : synonym of ''
Bathytoma ''Bathytoma'' is a genus of deep-water sea snails, marine gastropod mollusks in the family (biology), family Borsoniidae. Fossil records This genus is known in the fossil records from the Paleocene to the Quaternary (age range: 55.8 to 0.781 mi ...
'' Harris & Burrows, 1891 * ''Vexithara'' Finlay, 1926 : synonym of ''
Typhlomangelia ''Typhlomangelia'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae, the Conus, cone snails and their allies. Species Species within the genus ''Typhlomangelia'' include: * ''Typhlom ...
'' G.O. Sars, 1878 * ''Viridoturris'' Powell, 1964 : synonym of ''
Typhlomangelia ''Typhlomangelia'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Borsoniidae, the Conus, cone snails and their allies. Species Species within the genus ''Typhlomangelia'' include: * ''Typhlom ...
'' G.O. Sars, 1878


References


External links


Gastropods.com: Borsoniidae; accessed : 12 August 2011
*

{{Taxonbar, from=Q4946684 Conoidea