Scarce Swallowtail
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The scarce swallowtail (''Iphiclides podalirius'') is a species of butterfly belonging to the family
Papilionidae Swallowtail butterflies are large, colorful Butterfly, butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includ ...
. It is also called the sail swallowtail or pear-tree swallowtail.


Subspecies

Subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
include:"''Iphiclides'' Hübner, [1819]"
at Markku Savela's ''Lepidoptera and Some Other Life Forms''
*''Iphiclides podalirius centralasiae'' (Rosen, 1929) *''Iphiclides podalirius persica'' Verity, 1911 *''Iphiclides podalirius podalirius'' (Central and Southern Europe) *''Iphiclides podalirius virgatus'' (Butler, 1865) *''Iphiclides podalirius xinyuanensis '' Huang & Murayama, 1992 China (Xinyuan) Else see: *''Iphiclides feisthamelii">Iphiclides podalirius feisthamelii'' (North Africa, Spain and southwest France), often treated as a valid species, '' Iphiclides feisthamelii''.


Distribution

Despite the name "scarce swallowtail", this species is quite common. The scarcity of United Kingdom migrants is responsible for the English vernacular name. This species is widespread in the East
Palearctic realm The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. The ...
and in most of Europe with the exception of the northern parts. Its range extends northwards to
Lower Lusatia Lower Lusatia (; ; ; ; ) is a historical region in Central Europe, stretching from the southeast of the Germany, German state of Brandenburg to the southwest of Lubusz Voivodeship in Poland. Like adjacent Upper Lusatia in the south, Lower Lusa ...
and central Poland and eastwards across
Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
and
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and West Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Armenia, ...
as far as the
Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...
, Pakistan, India and western China. A few specimens of the scarce swallowtail have been reported from central Sweden and the United Kingdom but they were probably only strays and not migrants.


Habitat

These swallowtail butterflies inhabit
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s, towns as well as the countryside, in
fields Fields may refer to: Music *Fields (band), an indie rock band formed in 2006 * Fields (progressive rock band), a progressive rock band formed in 1971 * ''Fields'' (album), an LP by Swedish-based indie rock band Junip (2010) * "Fields", a song by ...
and open
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s. They are found in places with sloe thickets and particularly
orchard An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit tree, fruit- or nut (fruit), nut-producing trees that are generally grown for commercial production. Orchards are also so ...
s. In the
Alps The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia. ...
they can be found up to altitudes of 2000 m, but usually they prefer foothills and lower levels.Euro Butterflies by Matt Rowlings
/ref> The presence of ''Iphiclides podalirius'' in the floodplain of the Morava River in the
Slovak Republic Slovakia, officially the Slovak Republic, is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the west, and the Czech Republic to the northwest. Slovakia's ...
have been found to be a good indicator of relatively well preserved xerothermic
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
habitats with forest-steppe vegetation, which have no cutting history.


Status

In some years the scarce swallowtail is quite abundant. The scarce swallowtail is getting rarer as blackthorn bushes are being cleared. The butterfly is now protected by law in the
Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
,
Slovakia Slovakia, officially the Slovak Republic, is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the west, and the Czech Republic to the northwest. Slovakia's m ...
,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
,
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
,
Luxembourg Luxembourg, officially the Grand Duchy of Luxembourg, is a landlocked country in Western Europe. It is bordered by Belgium to the west and north, Germany to the east, and France on the south. Its capital and most populous city, Luxembour ...
,
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
,
Ukraine Ukraine is a country in Eastern Europe. It is the List of European countries by area, second-largest country in Europe after Russia, which Russia–Ukraine border, borders it to the east and northeast. Ukraine also borders Belarus to the nor ...
,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
and
Macedonia Macedonia (, , , ), most commonly refers to: * North Macedonia, a country in southeastern Europe, known until 2019 as the Republic of Macedonia * Macedonia (ancient kingdom), a kingdom in Greek antiquity * Macedonia (Greece), a former administr ...
. It is considered rare and endangered and protected in some provinces of Austria and of indeterminate status throughout Europe. In Armenia the species demonstrates stable population trend and is evaluated as Least Concern. Though referred by some authorities to be of status "vulnerable", it is however unlisted in the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
.


Description

''Iphiclides podalirius'' has a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
of in males, of in females. It is a very large distinctive butterfly. The background color of the wings is creamy white or pale yellow. On the front wings there are six tiger stripes and wedge-shaped markings. At the outer edge of the hind wings there are blue crescent markings, with an oblong, orange spot at the back corner and a relatively long tail. This species is rather similar to ''
Papilio machaon ''Papilio machaon'', the Old World swallowtail, is a butterfly of the family Papilionidae. The butterfly is also known as the common yellow swallowtail or simply the swallowtail (a common name applied to all members of the family, but this speci ...
'', '' Papilio hospiton'', '' Papilio alexanor'' and '' Protographium marcellus''. Iphiclides podalirius MHNT CUT 2013 3 9 Male Dos Cahors.jpg, Male Iphiclides podalirius MHNT CUT 2013 3 9 Male Ventre Cahors.jpg, Male underside Iphiclides podalirius MHNT CUT 2013 3 9 Female Dos Cahors.jpg, Female Iphiclides podalirius MHNT CUT 2013 3 9 Female Ventre Cahors.jpg, Female underside


Biology

Adults of ''Iphiclides podalirius'' fly from March to October. There are one, two, or three generations a year. Caterpillars are
polyphagous Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffixes -vore, -vory, or -vorous from Latin ''vorare'', meaning "to devour", or -phage, -phagy, or -phagous from Greek φαγε ...
feeding on ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' (Finland), ''
Crataegus monogyna ''Crataegus monogyna'', known as common hawthorn, whitethorn, one-seed hawthorn, or single-seeded hawthorn, is a species of flowering plant in the rose family, Rosaceae. It grows to about tall, producing plant sexuality, hermaphrodite flowers i ...
'' (Palaearctic), ''
Malus pumila An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are cultivated worldwide. The tree originated in Central A ...
'' and ''
Malus domestica An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...
'' (Palaearctic), ''Prunus'' (Finland, Palaearctic), ''
Prunus armeniaca ''Prunus armeniaca'' is the most commonly cultivated apricot species. The native range is somewhat uncertain due to its extensive prehistoric cultivation. Genetic studies indicate Central Asia is the center of origin. It is extensively cultivat ...
'' (Palaearctic, Spain), ''
Prunus avium ''Prunus avium'', commonly called wild cherry, sweet cherry or gean is a species of Prunus subg. Cerasus, cherry, a flowering plant in the rose family, Rosaceae. It is native to Eurasia and naturalized elsewhere. It is an ancestor of ''Prunus c ...
'' (Palaearctic, Spain) ''
Prunus cerasus ''Prunus cerasus'' (sour cherry, tart cherry, or dwarf cherry) is an Old World species of ''Prunus'' in the subgenus ''Prunus subg. Cerasus, Cerasus'' (cherries). It has two main groups of cultivars: the dark-red Morello cherry and the lighter- ...
'' (Palaearctic, Russia), ''
Prunus domestica ''Prunus domestica'' is a species of flowering plant in the family Rosaceae. A deciduous tree, it includes many varieties of the fruit trees known as plums in English, though not all plums belong to this species. The greengages and damsons als ...
'' (Palaearctic), ''
Prunus dulcis The almond (''Prunus amygdalus'', syn. ''Prunus dulcis'') is a species of tree from the genus ''Prunus''. Along with the peach, it is classified in the subgenus ''Amygdalus'', distinguished from the other subgenera by corrugations on the sh ...
'' (Palaearctic, Spain), ''
Prunus padus ''Prunus padus'', known as bird cherry, hackberry (unrelated to the genus ''Celtis''), hagberry, or Mayday tree, is a flowering plant in the Rosaceae, rose family. It is a species of cherry, a deciduous small tree or large shrub up to tall. It ...
'' (Palaearctic), ''
Prunus persica The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in China. It bears edible juicy fruits with various characteristics, most called peaches and the glossy-skinned, non-fuzzy varieties called nectarines. Peach ...
'' (Palaearctic), ''
Prunus spinosa ''Prunus spinosa'', called blackthorn or sloe, is an Old World species of flowering plant in the rose family, Rosaceae. It is locally naturalized in parts of the New World. The fruits are used to make sloe gin in Britain and patxaran in B ...
'' (Palaearctic), ''
Pyrus Pears are fruits produced and consumed around the world, growing on a tree and harvested in late summer into mid-autumn. The pear tree and shrub are a species of genus ''Pyrus'' , in the family Rosaceae, bearing the pomaceous fruit of the sa ...
'' (Palaearctic) and ''
Sorbus aucuparia ''Sorbus aucuparia'', commonly called rowan (, also ) and mountain-ash, is a species of deciduous tree or shrub in the rose family. The tree has a slender trunk with smooth bark, a loose and roundish crown, and its leaves are pinnate in pairs o ...
'' (Palaearctic). The caterpillars spin little pads on leaves and grip them firmly. The newly hatched caterpillar is dark in colour with two smaller and two bigger greenish patches on the dorsal side; later they are greenish with yellowish dorsal and side stripes. The summer chrysalids are green as a rule, the hibernating ones are brown. A number of hibernating chrysalids fall prey to various enemies. The caterpillars of the scarce swallowtail have been noted to leave silk trails from the permanent resting sites to feeding sites. This has been seen in both solitary and territorial larvae with larvae having the ability to discern their trails from those of others. Research on pupae of ''Iphiclides podalirius'' in Spain indicates that the pupae manifest in two colours, green and brown, for the purpose of camouflage. The green pupae develop on host plants and develop directly while brown pupae enter into
diapause In animal dormancy, diapause is the delay in development in response to regular and recurring periods of adverse environmental conditions.Tauber, M.J., Tauber, C.A., Masaki, S. (1986) ''Seasonal Adaptations of Insects''. Oxford University Press It ...
in the
leaf litter Plant litter (also leaf litter, tree litter, soil litter, litterfall, or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that has fallen to the ground. This detritus or dead organic material and its constituen ...
. Pupating larvae tend to form green pupae before August while after August they tend to form brown pupae. Duration of the photophase or light period appears to be the mechanism which dictates the path of development of the pupa. The results suggest that the green pupa develop on food plants to avoid predation by small mammals and visual avian predators while the brown pupa develop on leaf litter to avoid avian predators.


Life cycle

File: Iphiclides podalirius 1.jpg, Caterpillar File:Iphiclides podalirius pupating.jpg, Pupating caterpillar File:SailSwallowtail.jpg, Imago


References


External links


Scarce swallowtail
on Butterflycorner.net
Lepiforum.de
* Paolo Mazzei, Daniel Morel, Raniero Panfil

* ttps://www.guypadfield.com/scarceswallowtail.html ''Iphiclides podalirius'' on Guy Padfield's Butterfly Page* Butterfly Conservation Armeni

{{Taxonbar, from=Q31080 Iphiclides Butterflies of Asia Butterflies of Europe Butterflies described in 1758 Taxa named by Carl Linnaeus Butterflies of Africa