Percy Alexander Hulley (born 23 February 1941) is a South African

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

and

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...

. He is a research associate at the

South African Museum The Iziko South African Museum, formerly the South African Museum (Afrikaans: Suid-Afrikaanse Museum), is a South African national museum located in Cape Town. The museum was founded in 1825, the first in the country. It has been on its present ...

and has described many

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of fish including the taillight shark.

Biography

Hulley completed his MSc in 1967 from the

University of Cape Town The University of Cape Town (UCT) (, ) is a public university, public research university in Cape Town, South Africa. Established in 1829 as the South African College, it was granted full university status in 1918, making it the oldest univer ...

with a thesis entitled: ''Studies on the anatomy of some South African Mytilidae (Bivalvia) with notes on their ecology and distribution''. He submitted his PhD thesis in 1971 entitled: ''The origin, interrelationship and distribution of Southern African Rajidae (Chondrichthyes, Batoidei)'' He has worked at the

since 1965, where he was deputy director (research) and curator of fishes at the

until 2005. From 2006, Hulley has worked at the South African Museum in an honorary capacity as a research associate. The comprehensive

Mesopelagic Fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that live on or near the bottom, and reef fish that are associated with coral reefs. ...

Collection of the South African museum was assembled primarily by Hulley. The collection contains fish that live between 200m and 1 000m below the surface. Mesopelagic fish constitutes 95% of the world’s fish

biomass Biomass is a term used in several contexts: in the context of ecology it means living organisms, and in the context of bioenergy it means matter from recently living (but now dead) organisms. In the latter context, there are variations in how ...

Selected publications

Hulley has authored or co-authored more than 100 articles, reports and chapters for academic works, including: * * * Amblyraja hyperborea1

Taxa named in his honor

Hulley is honoured in the naming of a legskate and other fishes: * ''

Eustomias ''Eustomias'' is a genus of barbeled dragonfishes native to the oceanic depths of the Indian, Atlantic and Pacific oceans. Species There are currently 121 recognized species in this genus: * '' Eustomias achirus'' Parin & Pokhil'skaya, 1974 (P ...

hulleyi'' - Goman & Gibbs 1985 * '' Antigonia hulleyi'' - Parin & Borodulina 2005 * '' Argyripnus hulleyi'' - Quéro,

Spitz A spitz (; , in reference to the pointed muzzle) is a Dog type, type of domestic dog consisting of between 50 and 70 breeds depending on classification. There is no precise definition of 'spitz' but typically most spitz breeds have pricked ears ...

& Vayne, 2009 * Roughnose Legskate ''

Cruriraja ''Cruriraja'' is a genus of skates in the family Gurgesiellidae.Weigmann, S., Séret, B., Last, P.R. & McEachran, J.D. 2016. Pygmy Skates, Family Gurgesiellidae. Pp. 473–493 in Last, P.R., White, W.T., Carvalho, M.R., Séret, B., Stehmann, M. ...

hulleyi'' - Aschliman, Ebert & Compagno 2010 * ''

Diogenichthys ''Diogenichthys'' is a genus of lanternfishes. Species There are currently three recognized species in this genus: * '' Diogenichthys atlanticus'' ( Tåning, 1928) (Longfin lanternfish) * '' Diogenichthys laternatus'' (Garman Garman is a surname ...

hulleyi'' - Schwarzhans 2013

Taxa described by him

Hulley discovered and named several other sharks and fish, amongst others: * Taillight Shark

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Euprotomicroides The taillight shark (''Euprotomicroides zantedeschia'') is a little-known species of shark in the family Dalatiidae and the only member of its genus.Stehmann, M.F.W., Van Oijen, M. & Kamminga, P. (2016): Re-description of the rare taillight s ...

'' - Hulley 1966 * Bottleneck Skate genus ''

Rostroraja ''Rostroraja'' is a genus of skate belonging to the family Rajidae, native to the Atlantic Ocean and East Pacific, including the Mediterranean, Caribbean, Gulf of Mexico and Gulf of California. The genus was considered monotypic in the past (c ...

'' - Hulley 1972 * Taillight Shark ''Euprotomicroides zantesdeschia'' - Hulley & Penrith 1966 * Roberts Bigmouth Skate '' Amblyraja robertsi'' - Hulley 1970 * Yellow-spotted Skate ''

Leucoraja ''Leucoraja'' is a genus of hardnose skates in the family Rajidae, commonly known as the rough skates. They occur mostly on continental shelves and slopes in the north-western and eastern Atlantic Ocean, the Mediterranean Sea, the south-wester ...

wallacei'' - Hulley 1970 * Ghost Skate ''

Rajella dissimilis ''Rajella'' is a genus of skate found deeper than in the Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on ...

'' - Hulley 1970 * Smoothback Skate ''

Rajella ravidula ''Rajella'' is a genus of skate found deeper than in the Pacific, Atlantic and Indian oceans. Species There are currently 18 recognized species in this genus: * '' Rajella annandalei'' M. C. W. Weber, 1913 (Annandale's skate) * '' Rajella bar ...

'' - Hulley 1972 * African Pygmy Skate ''

Neoraja stehmanni The African dwarf skate (''Neoraja stehmanni''), or South African pygmy skate, is a species of fish in the family Rajidae. It is endemic to South Africa. Its natural habitat In ecology, habitat refers to the array of resources, biotic f ...

'' - Hulley 1972 * Lanternfish genus ''

Krefftichthys ''Krefftichthys anderssoni'' is a species of lanternfish found circumglobally in the southern oceans. This species grows to a length of fish measurement, SL. References

* Myctophidae Monotypic fish genera Taxa named by Einar Lönnberg ...

'' - Hulley 1981 * False-Midas lanternfish ''

Gymnoscopelus ''Gymnoscopelus'' is a genus of lanternfishes. The name is from the Greek ''gymnos'', "naked" and ''skopelos'', "lanternfish." They can live in extremely cold water; two species, '' Gymnoscopelus nicholsi'' and '' Gymnoscopelus braueri'', have b ...

hintonoides'' - Hulley 1981 * Minispotted lanternfish ''

microlampas'' - Hulley 1981 * Nacreous lanternfish ''

Lampanyctus vadulus ''Lampanyctus vadulus'' is a species of lanternfish Lanternfish (or myctophids, from the Greek language, Greek μυκτήρ ''myktḗr'', "nose" and ''ophis'', "serpent") are small mesopelagic fish of the large family (biology), family Myctop ...

'' - Hulley 1981 * Herwig lanternfish ''

Metelectrona ''Metelectrona'' is a genus of lanternfishes. Species There are currently three recognized species in this genus: * '' Metelectrona ahlstromi'' Wisner, 1963 * '' Metelectrona herwigi'' Hulley, 1981 (Herwig's lanternfish) * '' Metelectrona vent ...

herwigi'' - Hulley 1981 * Gaptooth lanternfish ''

Protomyctophum ''Protomyctophum'' is a genus of lanternfishes. Species There are currently 16 recognized species in this genus: * '' Protomyctophum andriashevi'' Becker, 1963 * '' Protomyctophum arcticum'' ( Lütken, 1892) (Arctic telescope) * '' Protomyct ...

choriodon'' - Hulley 1981 * Jewelled lanternfish ''

gemmatum'' - Hulley 1981 * Damsel lanternfish ''

luciferum'' - Hulley 1981 * Krefft's lanternfish ''

Symbolophorus kreffti ''Symbolophorus kreffti'' is a species of fish in the family Myctophidae. Etymology The fish is named in honor of Gerhard Krefft (1912-1993) of the Institut für Seefischerei in Hamburg, he being the first to recognize the specimens of ''Symbo ...

'' - Hulley 1981 * '' Astronesthes illuminatus'' - Parin, Borodulina & Hulley 1999 * '' Astronesthes galapagensis'' - Parin, Borodulina & Hulley 1999 *See :Taxa named by P. Alexander Hulley

References

External references

*
Publications by P. Alexander Hulley
on

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{{DEFAULTSORT:Hulley, Percy Alexander South African zoologists South African ichthyologists Living people 1941 births University of Cape Town alumni