Parornix Anglicella
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''Parornix anglicella'' is a
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
of the family
Gracillariidae Gracillariidae is an important family of insects in the order Lepidoptera and the principal family of leaf miners that includes several economic, horticultural or recently invasive pest species such as the horse-chestnut leaf miner, ''Cameraria ...
found in Asia and Europe. It was described in 1850, by the English
entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...
Henry Tibbats Stainton Henry Tibbats Stainton (13 August 1822 – 2 December 1892) was an England, English entomologist. He served as an editor for two popular entomology periodicals of his period, ''The Entomologist's Annual'' and ''The Entomologist's Weekly Intellig ...
, from a
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...
from
Lewisham Lewisham ( ) is an area of southeast London, England, south of Charing Cross. It is the principal area of the London Borough of Lewisham, and was within the Historic counties of England, historic county of Kent until 1889. It is identified in ...
,
Kent Kent is a Ceremonial counties of England, ceremonial county in South East England. It is bordered by Essex across the Thames Estuary to the north, the Strait of Dover to the south-east, East Sussex to the south-west, Surrey to the west, and Gr ...
.


Description

The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
is 9–11 mm. The head is ochreous-whitish mixed with fuscous. Palpi white, apex of second joint and median band of terminal dark fuscous. Forewings are dark fuscous, irrorated with whitish; numerous costal strigulae, a spot in middle of disc and another posteriorly, and suffused dorsal strigidae interrupted by two blackish spots white; a black apical dot; cilia with three entire dark fuscous lines. Hindwings are grey. The larva is pale greenish-grey; dorsal line darker; spots pale; head pale greenish-brown; segment 2 with four black spots.


Biology

Adults are on the wing in two generations in April and May and again in August. ;Ovum Eggs are laid on the underside of the leaves of hawthorns (''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' species), but have also been recorded on medlar (''
Mespilus germanica ''Mespilus germanica'', known as the medlar or common medlar, is a large shrub or small tree in the rose family Rosaceae. When the genus ''Mespilus'' is included in the genus ''Crataegus'', the correct name for this species is ''Crataegus german ...
''), rowan (''
Sorbus aucuparia ''Sorbus aucuparia'', commonly called rowan (, also ) and mountain-ash, is a species of deciduous tree or shrub in the rose family. The tree has a slender trunk with smooth bark, a loose and roundish crown, and its leaves are pinnate in pairs o ...
''), wild service tree (''
Sorbus torminalis ''Torminalis'' is a genus of plants in the rose family, Rosaceae. The genus was formerly included within the genus ''Sorbus'' as the Section (biology), section ''Torminaria'', but the simple-leafed species traditionally classified in ''Sorbus'' ...
'') and strawberries (''
Fragaria ''Fragaria'' () is a genus of flowering plants in the rose family, Rosaceae, commonly known as strawberries for their edible fruits. There are more than 20 described species and many Hybrid plant, hybrids and cultivars. The most common strawberr ...
'' species). ;Larvae Feeding starts in a gallery in the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and Subcutaneous tissue, hypodermis. The epidermal layer provides a barrier to infection from environmental pathogens and regulates the ...
, which leads to a leaf margin and develops into a blotch in the second instar. In the next phase the larva eats through to the upper epidermis, puckers the leaf and the epidermis turns brown and the frass is at one end of the mine. It is distinguished from a ''
Phyllonorycter oxyacanthae ''Phyllonorycter oxyacanthae'' is a moth of the family Gracillariidae found in all of Europe except the Balkan Peninsula. It was Species description, described by the German people, German-born Swiss people, Swiss entomologist, Heinrich Frey in 1 ...
'' mine, by the smaller size and the lower epidermis stays green. After leaving the mine two or more cones are made by folding the leaves downwards in the manner of a ''
Caloptilia ''Caloptilia'' is a genus of moths in the family Gracillariidae. Species *'' Caloptilia acericola'' Kumata, 1966 *'' Caloptilia acericolella'' Kuznetzov, 1981 *'' Caloptilia aceriella'' (Chambers, 1881) *'' Caloptilia acerifoliella'' (Chambers, ...
'' rather than other ''Parornix''. Occupied mines can be found from July to September. ;Pupa The pupa is pale-yellow-brown in a whitish cocoon, sometimes spun under a narrowly folded edge of a leaf, or in leaf-litter or an old bird's nest. Pupation takes place in July–August and September to late-April.


Gallery

Image:Parornix anglicella a hawthorn leaf with a mine of a young larva and two cones formed by adult larvae.JPG, A hawthorn leaf with a
mine Mine, mines, miners or mining may refer to: Extraction or digging *Miner, a person engaged in mining or digging *Mining, extraction of mineral resources from the ground through a mine Grammar *Mine, a first-person English possessive pronoun M ...
of a young larva and two cones formed by adult larvae File:Parornix anglicella, larva. (5093612098).jpg File:15.028 BF303 Parornix anglicella (2534090303).jpg File:Parornix anglicella male, Minera, North Wales, May 2017 (35077145956).jpg


Distribution

It is widespread in:
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
including
Albania Albania ( ; or ), officially the Republic of Albania (), is a country in Southeast Europe. It is located in the Balkans, on the Adriatic Sea, Adriatic and Ionian Seas within the Mediterranean Sea, and shares land borders with Montenegro to ...
,
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
,
Belarus Belarus, officially the Republic of Belarus, is a landlocked country in Eastern Europe. It is bordered by Russia to the east and northeast, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the northwest. Belarus spans an a ...
,
Belgium Belgium, officially the Kingdom of Belgium, is a country in Northwestern Europe. Situated in a coastal lowland region known as the Low Countries, it is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeas ...
,
Bosnia and Herzegovina Bosnia and Herzegovina, sometimes known as Bosnia-Herzegovina and informally as Bosnia, is a country in Southeast Europe. Situated on the Balkans, Balkan Peninsula, it borders Serbia to the east, Montenegro to the southeast, and Croatia to th ...
,
Britain Britain most often refers to: * Great Britain, a large island comprising the countries of England, Scotland and Wales * The United Kingdom of Great Britain and Northern Ireland, a sovereign state in Europe comprising Great Britain and the north-eas ...
,
Bulgaria Bulgaria, officially the Republic of Bulgaria, is a country in Southeast Europe. It is situated on the eastern portion of the Balkans directly south of the Danube river and west of the Black Sea. Bulgaria is bordered by Greece and Turkey t ...
,
Croatia Croatia, officially the Republic of Croatia, is a country in Central Europe, Central and Southeast Europe, on the coast of the Adriatic Sea. It borders Slovenia to the northwest, Hungary to the northeast, Serbia to the east, Bosnia and Herze ...
,
Corsica Corsica ( , , ; ; ) is an island in the Mediterranean Sea and one of the Regions of France, 18 regions of France. It is the List of islands in the Mediterranean#By area, fourth-largest island in the Mediterranean and lies southeast of the Metro ...
,
Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
, Danish mainland,
Estonia Estonia, officially the Republic of Estonia, is a country in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the Baltic Sea across from Sweden, to the south by Latvia, and to the east by Ru ...
,
Finland Finland, officially the Republic of Finland, is a Nordic country in Northern Europe. It borders Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bothnia to the west and the Gulf of Finland to the south, ...
, French mainland,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
,
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
,
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
, Italian mainland,
Latvia Latvia, officially the Republic of Latvia, is a country in the Baltic region of Northern Europe. It is one of the three Baltic states, along with Estonia to the north and Lithuania to the south. It borders Russia to the east and Belarus to t ...
,
Lithuania Lithuania, officially the Republic of Lithuania, is a country in the Baltic region of Europe. It is one of three Baltic states and lies on the eastern shore of the Baltic Sea, bordered by Latvia to the north, Belarus to the east and south, P ...
,
Luxembourg Luxembourg, officially the Grand Duchy of Luxembourg, is a landlocked country in Western Europe. It is bordered by Belgium to the west and north, Germany to the east, and France on the south. Its capital and most populous city, Luxembour ...
,
Montenegro , image_flag = Flag of Montenegro.svg , image_coat = Coat of arms of Montenegro.svg , coa_size = 80 , national_motto = , national_anthem = () , image_map = Europe-Mont ...
,
North Macedonia North Macedonia, officially the Republic of North Macedonia, is a landlocked country in Southeast Europe. It shares land borders with Greece to the south, Albania to the west, Bulgaria to the east, Kosovo to the northwest and Serbia to the n ...
, Norwegian mainland,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
, Portuguese mainland,
Romania Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
, central and northern
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
,
Sardinia Sardinia ( ; ; ) is the Mediterranean islands#By area, second-largest island in the Mediterranean Sea, after Sicily, and one of the Regions of Italy, twenty regions of Italy. It is located west of the Italian Peninsula, north of Tunisia an ...
,
Serbia , image_flag = Flag of Serbia.svg , national_motto = , image_coat = Coat of arms of Serbia.svg , national_anthem = () , image_map = , map_caption = Location of Serbia (gree ...
,
Sicily Sicily (Italian language, Italian and ), officially the Sicilian Region (), is an island in the central Mediterranean Sea, south of the Italian Peninsula in continental Europe and is one of the 20 regions of Italy, regions of Italy. With 4. ...
,
Slovakia Slovakia, officially the Slovak Republic, is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the west, and the Czech Republic to the northwest. Slovakia's m ...
,
Slovenia Slovenia, officially the Republic of Slovenia, is a country in Central Europe. It borders Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the south and southeast, and a short (46.6 km) coastline within the Adriati ...
,
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
,
Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
,
Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
,
Ukraine Ukraine is a country in Eastern Europe. It is the List of European countries by area, second-largest country in Europe after Russia, which Russia–Ukraine border, borders it to the east and northeast. Ukraine also borders Belarus to the nor ...
. Outside Europe it is recorded from the
Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...
and
Nearctic realm The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface. The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America t ...
.


Taxonomy

The genus ''Paronix'' comes from ''para'', meaning alongside and ''ornix'' named after the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Ornix'' Treitschke, 1833, which refers to ''ornis'', a bird. The genus ''Ornix'' originally included a wide range of feathery-winged
microlepidoptera Microlepidoptera (micromoths) is an artificial (i.e., unranked and not monophyletic) grouping of moth families, commonly known as the "smaller moths" ( micro, Lepidoptera). These generally have wingspans of under 20 mm, so are harder to iden ...
in the
Coleophoridae The Coleophoridae are a family of small moths, belonging to the huge superfamily Gelechioidea. Collectively known as case-bearers, casebearing moths or case moths, this family is represented on all continents, but the majority are found in temp ...
and the
Gracillariidae Gracillariidae is an important family of insects in the order Lepidoptera and the principal family of leaf miners that includes several economic, horticultural or recently invasive pest species such as the horse-chestnut leaf miner, ''Cameraria ...
. ''Ornix'' was an early
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
of the genus ''
Coleophora ''Coleophora'' is a very large genus of moths of the family Coleophoridae. It contains some 1,350 described species. The genus is represented on all continents, but the majority are found in the Nearctic and Palaearctic regions. Many authors have t ...
'' as a number of moths were named after birds. The feathery-winged moths were later restricted to ''Paronix''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''anglicella'' refers to ''Anglicus'' i.e. English; the type locality is from England.


References


External links


Lepiforum de

Larvae on Plant Parasites of Europe

UKflymines
{{Taxonbar, from=Q7139471 Parornix Leaf miners Moths described in 1850 Moths of Asia Moths of Europe Taxa named by Henry Tibbats Stainton