Myanmar Amber
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Burmese amber, also known as Burmite or Kachin amber, is
amber Amber is fossilized tree resin. Examples of it have been appreciated for its color and natural beauty since the Neolithic times, and worked as a gemstone since antiquity."Amber" (2004). In Maxine N. Lurie and Marc Mappen (eds.) ''Encyclopedia ...
from the Hukawng Valley in northern
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
. The amber is dated to around 100 million years ago, during the latest
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
to earliest
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
ages of the mid-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to the potential role of the amber trade in funding
internal conflict in Myanmar Myanmar has been embroiled in armed conflict since 1948, when the country, then known as Burma, Burmese Declaration of Independence, gained independence from the United Kingdom. The conflict has largely been Ethnic conflict, ethnic-based, wit ...
and hazardous working conditions in the mines where it is collected.


Geological context, depositional environment and age

The amber is found in the Hukawng Basin, a large Cretaceous-
Cenozoic The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
sedimentary basin Sedimentary basins are region-scale depressions of the Earth's crust where subsidence has occurred and a thick sequence of sediments have accumulated to form a large three-dimensional body of sedimentary rock They form when long-term subsidence ...
within northern Myanmar. The strata have undergone folding and
faulting In geology, a fault is a planar fracture or discontinuity in a volume of rock across which there has been significant displacement as a result of rock-mass movements. Large faults within Earth's crust result from the action of plate tectonic ...
. The Hukawng basin is part of the larger Myanmar Central Basin, a N-S orientated
synclinal Synclinal may refer to: * Syncline, in structural geology, a syncline is a fold, with younger layers closer to the center of the structure. *Synclinal, in alkane stereochemistry, a torsion angle between 30° to 90° and –30° to –90° See als ...
basin extending to the Gulf of Martaban to the south. The basin is considered to be a part of the West Burma block (a.k.a. Burma
terrane In geology, a terrane (; in full, a tectonostratigraphic terrane) is a crust fragment formed on a tectonic plate (or broken off from it) and accreted or " sutured" to crust lying on another plate. The crustal block or fragment preserves its d ...
), which has a debated tectonic history, it is considered to be associated with the concepts of the Cimmeria and Sibumasu terranes. The block was part of
Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...
during at least the Early
Paleozoic The Paleozoic ( , , ; or Palaeozoic) Era is the first of three Era (geology), geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma a ...
, but the timing of rifting is very uncertain, with estimates ranging from the
Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...
to Early Cretaceous. It is also disputed whether the block had accreted onto the Asian
continental margin A continental margin is the outer edge of continental crust abutting oceanic crust under coastal waters. It is one of the three major zones of the ocean floor, the other two being deep-ocean basins and mid-ocean ridges. The continental marg ...
by the time of the amber deposition. A 2025 study found that the West Burma block had separated from what is now the northwest coast of Australia during the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
period. Paleomagnetic reconstructions by Westerweel et al, have suggested that the Burma terrane formed an island land mass in the
Tethys Ocean The Tethys Ocean ( ; ), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean during much of the Mesozoic Era and early-mid Cenozoic Era. It was the predecessor to the modern Indian Ocean, the Mediterranean Sea, and the Eurasia ...
during the Mid Cretaceous at a
latitude In geography, latitude is a geographic coordinate system, geographic coordinate that specifies the north-south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from −90° at t ...
around 5-10 degrees south of the equator, considerably northeast of the combined Insular India (comprising the modern Indian subcontinent)-
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
landmass. Some members of the Burmese amber flora and fauna have Gondwanan affinities, while others have Laurasian affinities. At Noije Bum, located on a ridge, amber is found within fine grained
clastic rock Clastic rocks are composed of fragments, or clasts, of pre-existing minerals and rock. A clast is a fragment of geological detritus,Essentials of Geology, 3rd Ed, Stephen Marshak, p. G-3 chunks, and smaller grains of rock broken off other rocks b ...
s, typically medium to greyish green in colour, resulting from the constituent grains being black, yellow, grey and light green. The fine grained rocks are primarily fine to very fine grained
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
, with beds of
silt Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension (chemistry), suspension with water. Silt usually ...
and
shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of Clay mineral, clay minerals (hydrous aluminium phyllosilicates, e.g., Kaolinite, kaolin, aluminium, Al2Silicon, Si2Oxygen, O5(hydroxide, OH)4) and tiny f ...
and laterally persistent thin (1–2 mm thick)
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal i ...
horizons. Massive micritic
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
interbeds of 6-8 centimetre thickness, often containing coalified plant material also occur. This
facies In geology, a facies ( , ; same pronunciation and spelling in the plural) is a body of rock with distinctive characteristics. The characteristics can be any observable attribute of rocks (such as their overall appearance, composition, or con ...
association is typically around 1 metre thick and typically thinly bedded and laminated. Associated with the fine grained facies is a set of medium facies primarily consisting of medium to fine grained sandstones also containing thin beds of siltstone, shale and conglomerate, alongside a persistent conglomerate horizon. A specimen of the
ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
'' Mortoniceras'' has been found in a sandstone bed 2 metres above the amber horizon, alongside indeterminate gastropods and bivalves. Lead-uranium dating of
zircon Zircon () is a mineral belonging to the group of nesosilicates and is a source of the metal zirconium. Its chemical name is zirconium(IV) silicate, and its corresponding chemical formula is Zr SiO4. An empirical formula showing some of th ...
crystals of volcanic clasts within the amber bearing horizons has given a maximum age of 98.79 ± 0.62  million years ago ( Ma), making the deposit earliest
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
in age. Unpublished data by Wang Bo on other layers suggests an age range of deposition of at least 5 million years. The amber does not appear to have undergone significant transport since hardening or be redeposited. The strata at the site are younging upwards, striking north north-east and dipping 50-70 degrees E and SE north of the ridge and striking between south south-east and south-east and dipping 35-60 degrees south-west south of the ridge, suggesting the site is on the northwest limb of a syncline plunging to the northeast. A minor fault with a conspicuous gouge zone was noted as present, though it appeared to have no significant displacement. Several other localities are known, including the colonial Khanjamaw and the more recent Inzutzut, Angbamo, and Xipiugong sites, within the vicinity of Tanai. The Hkamti site SW of the Hukawng basin has been determined to be significantly older, dating to the early
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
around ca. 110 Ma and is therefore considered distinct.


Paleoenvironment

The Burmese amber paleoforest is considered to have been a
tropical rainforest Tropical rainforests are dense and warm rainforests with high rainfall typically found between 10° north and south of the Equator. They are a subset of the tropical forest biome that occurs roughly within the 28° latitudes (in the torrid zo ...
, situated near the coast, where resin was subsequently transported into a shallow marine environment. The shell of a dead juvenile ''Puzosia'' (''Bhimaites'') ammonite, four marine
gastropod shell The gastropod shell is part of the body of many gastropods, including snails, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium ...
s (including '' Mathilda'') and littoral or supralittoral
isopods Isopoda is an Order (biology), order of crustaceans. Members of this group are called isopods and include both Aquatic animal, aquatic species and Terrestrial animal, terrestrial species such as woodlice. All have rigid, segmented exoskeletons ...
entombed in a piece of amber with shell sand, along with growth of Isocrinid
crinoid Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that remain attached to the sea floor by a stalk in their adult form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are ...
s,
coral Corals are colonial marine invertebrates within the subphylum Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important Coral ...
s and
oyster Oyster is the common name for a number of different families of salt-water bivalve molluscs that live in marine or brackish habitats. In some species, the valves are highly calcified, and many are somewhat irregular in shape. Many, but no ...
s on the surface of some amber pieces indicate marine conditions for final deposition. Additionally pholadid (piddock) bivalve borings into amber specimens along with at least one pholadid which became trapped was interpreted to show that the resin was still fresh and unhardened when it was being moved into the tidal areas. However, the phloladids in question, belonging to the extinct genus '' Palaeolignopholas,'' were later interpreted as a freshwater species, and the presence of numerous freshwater insects suggests that the initial environment of deposition was a downstream estuarine to freshwater section of a river, with the forests extending across coastal rivers, river deltas, lakes, lagoons, and coastal bays. The forest environment may have been prone to fire, similar to modern tropical peat swamps, based on the presence of fire adapted plants and burned plant remains found in the amber. The amber itself is primarily disc-shaped and flattened along the bedding plane, and is typically reddish brown, with the colour ranging from shades of yellow to red. The opacity of the amber ranges from clear to opaque. Many amber pieces have thin
calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...
veins Veins () are blood vessels in the circulatory system of humans and most other animals that carry blood towards the heart. Most veins carry deoxygenated blood from the tissues back to the heart; exceptions are those of the pulmonary and fetal c ...
that are typically less than , but up to thick. The number and proportion of veins in a piece of amber varies significantly, in some pieces veins are virtually absent, while others are described as being "packed with veinlets". The amber is considered to be of
coniferous Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
origin, with a likely araucarian source tree, based on spectroscopic analysis and wood fragment inclusions, though a
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
origin has also been suggested.


Fauna and flora

The list of taxa is extraordinarily diverse, with 50 classes (or equivalent), 133 orders (or equivalent), 726 families, 1,757 genera and 2,770 species described as of 2023. The vast majority of the species are arthropods, mostly insects.


Arthropods

Over 2500 species of arthropods (with over 2000 of these species beings insects) are known from the deposit, notably including the oldest members of the arachnid orders Palpigradi ('' Electrokoenenia'') and
Schizomida Schizomida, also known as sprickets or short-tailed whip-scorpions, is an order of arachnids, generally less than in length. The order is not yet widely studied. E. O. Wilson has identified schizomids as among the "groups of organisms that des ...
and the only known fossil members of Ricinulei since the Paleozoic. '' Chimerarachne'' is a unique
stem-group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...
spider still possessing a tail, with similar forms only known from the Paleozoic, with over 360 other species of spiders (including the only non Paleozoic fossils of
Mesothelae The Mesothelae are a suborder of spiders (order Araneae). , two extant families were accepted by the World Spider Catalog, Liphistiidae and Heptathelidae. Alternatively, the Heptathelidae can be treated as a subfamily of a more broadly circum ...
) being known from Burmese amber, along with over 40 species of scorpions. Other significant arachnid fossils include some of the oldest
tick Ticks are parasitic arachnids of the order Ixodida. They are part of the mite superorder Parasitiformes. Adult ticks are approximately 3 to 5 mm in length depending on age, sex, and species, but can become larger when engorged. Ticks a ...
s, as well as the oldest
mesostigmata Mesostigmata is an order of mites belonging to the Parasitiformes. They are by far the largest group of Parasitiformes, with over 8,000 species in 130 families. Mesostigmata includes parasitic as well as free-living and predatory forms. They ca ...
n and opilioacarid mites.
Beetles Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
(Coleoptera), reflecting their modern diversity, are represented by over 100 families and over 500 species, with
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typi ...
(bees, wasps, ants and sawflies) represented by 70 families and 350 species including some of the oldest known
ants Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of ...
.
Flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...
(Diptera) are represented by over 50 families and over 250 species and true bugs (
Hemiptera Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from ...
) are represented by over 80 families and over 220 species. Dictyoptera (
cockroaches Cockroaches (or roaches) are insects belonging to the Order (biology), order Blattodea (Blattaria). About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known Pest (organism), pests. Modern cockro ...
,
termite Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...
s and mantises, among others), are represented by over 100 species in 28 families.
Odonata Odonata is an order of predatory flying insects that includes the dragonflies and damselflies (as well as the '' Epiophlebia'' damsel-dragonflies). The two major groups are distinguished with dragonflies (Anisoptera) usually being bulkier with ...
(which contains
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
and
damselflies Damselflies are flying insects of the suborder Zygoptera in the order Odonata. They are similar to dragonflies (which constitute the other odonatan suborder, Epiprocta) but are usually smaller and have slimmer bodies. Most species fold the win ...
, among other extinct groups), is represented by over 19 families and over 40 species, far higher than in other Cretaceous amber deposits. Among the oldest members of the insect order
Zoraptera The insect order (biology), order Zoraptera, commonly known as angel insects, contains small and soft bodied insects with two forms: winged with wings sheddable as in termites, dark and with eyes (compound) and ocelli (simple); or wingless, pale ...
are known from Burmese amber, as well as the oldest parasitic
lice Louse (: lice) is the common name for any member of the infraorder Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera was previously recognized as an order, until a 2021 genetic study determined th ...
and the oldest members of the parasitic insect order
Strepsiptera The Strepsiptera () are an order of insects with eleven extant families that include about 600 described species. They are endoparasites of other insects, such as bees, wasps, leafhoppers, Zygentoma, silverfish, and cockroaches. Females of most s ...
. The winged insect '' Aristovia'' is transitional between the extinct " Grylloblattodea" and modern wingless Grylloblattidae (ice crawlers). Over a dozen species each of
Myriapoda Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. Although molecular evidence and similar fossils suggests a diversifi ...
(millipedes and centipedes, among others) and
Entognatha The Entognatha, from Ancient Greek ἐντός (''entós''), meaning "inside", and γνάθος (''gnáthos''), meaning "jaw", are a class (biology), class of wingless and Ametabolism, ametabolous arthropods, which, together with the insects, ...
(
springtail Springtails (class Collembola) form the largest of the three lineages of modern Hexapoda, hexapods that are no longer considered insects. Although the three lineages are sometimes grouped together in a class called Entognatha because they have in ...
s, among others) have also been reported, along with a number of
woodlice Woodlice are terrestrial isopods in the suborder Oniscidea. Their name is derived from being often found in old wood, and from louse, a parasitic insect, although woodlice are neither parasitic nor insects. Woodlice evolved from marine isopods ...
(representing some of the oldest records of the group) and various aquatic crustaceans.


Other invertebrates

A wide variety of other invertebrates have been reported. These include
gastropods Gastropods (), commonly known as slugs and snails, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and from the land. Ther ...
, including
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...
and
land snails A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
, the oldest modern onychophoran (also known as velvet worms) '' Cretoperipatus,'' as well as
nematode The nematodes ( or ; ; ), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (h ...
s, nematomorphs,
annelid The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to vario ...
s and
flatworm Platyhelminthes (from the Greek language, Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") is a Phylum (biology), phylum of relatively simple bilaterian, Segmentation (biology), ...
s.


Vertebrates

Only a handful of vertebrates have been described from Burmese amber, these include the albanerpetontid (a group of extinct salamander-like amphibians) '' Yaksha'', the frog '' Electrorana'', a number of primitive toothed birds belonging to the extinct clade
Enantiornithes The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct Avialae, avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teet ...
, including the named species '' Elektorornis'' and '' Fortipesavis'', the lizards '' Protodraco'' (suggested to be the oldest agamid)'', Cretaceogekko'' (suggested to be the oldest modern
gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards ...
)'', Electroscincus'' (the oldest known
skink Skinks are a type of lizard belonging to the family (biology), family Scincidae, a family in the Taxonomic rank, infraorder Scincomorpha. With more than 1,500 described species across 100 different taxonomic genera, the family Scincidae is one o ...
) '' Barlochersaurus'', '' Oculudentavis'' (formerly erroneously considered to be a bird), and the snake '' Xiaophis.'' An indeterminate
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur is known from a tail with preserved feathers.


Flora

A wide variety of plants have been reported from the deposit. These include
flowering plants Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
,
conifers Conifers () are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All e ...
,
ferns The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...
,
lycophyte The lycophytes, when broadly circumscribed, are a group of vascular plants that include the clubmosses. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest lineag ...
s and
bryophytes Bryophytes () are a group of land plants ( embryophytes), sometimes treated as a taxonomic division referred to as Bryophyta '' sensu lato'', that contains three groups of non-vascular land plants: the liverworts, hornworts, and mosses. In t ...
.


Other

A number of fungi species have been reported, as well as various microorganisms.


History

The amber is recorded as originating from
Yunnan Province Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
as early as the first century AD according to the ''
Book of the Later Han The ''Book of the Later Han'', also known as the ''History of the Later Han'' and by its Chinese name ''Hou Hanshu'' (), is one of the Twenty-Four Histories and covers the history of the Han dynasty from 6 to 189 CE, a period known as the Lat ...
.'' The trade of Burmese amber into China during the
Han Dynasty The Han dynasty was an Dynasties of China, imperial dynasty of China (202 BC9 AD, 25–220 AD) established by Liu Bang and ruled by the House of Liu. The dynasty was preceded by the short-lived Qin dynasty (221–206 BC ...
has been confirmed by Fourier-transform infrared spectroscopy (FTIR) analysis of amber artifacts from the Eastern Han Dynasty (25 - 220 CE). It was first mentioned in European sources by the
Jesuit The Society of Jesus (; abbreviation: S.J. or SJ), also known as the Jesuit Order or the Jesuits ( ; ), is a religious order (Catholic), religious order of clerics regular of pontifical right for men in the Catholic Church headquartered in Rom ...
Priest Álvaro Semedo who visited China in 1613, it was described as being "digged out of mines, and sometimes in great pieces, it is redder than our amber though not so cleane".Zherikhin, V.V., Ross, A.J., 2000
A review of the history, geology and age of Burmese amber (Burmite).
Bulletin of the Natural History Museum, London (Geology) 56 (1), 3–10.
The locality itself has been known to European explorers since the 1800s with visitation to the Hukawng Valley by Simon Fraser Hannay in 1836–1837. At that time the principle products of the valley mines were salt, gold, and amber, with the majority of gold and amber being bought by Chinese traders. Hannay visited the amber mines themselves on March 21, 1836, and he noted that the last three miles to the mines were marked with numerous abandoned pits, up to in depth, where amber had been dug in the past. The mining had moved over the hill to a series of 10 pits but no visible amber was seen, suggesting that miners possibly hid the amber found that day before the party arrived. Mining was being performed manually at the time through the use of sharpened
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
rods and small wooden shovels. Finer pieces of amber were recovered from the deeper pits, with clear yellow being recovered from depths of The recovered amber was bought with silver or often exchanged for jackets, hats, copper pots, or
opium Opium (also known as poppy tears, or Lachryma papaveris) is the dried latex obtained from the seed Capsule (fruit), capsules of the opium poppy ''Papaver somniferum''. Approximately 12 percent of opium is made up of the analgesic alkaloid mor ...
among other goods. mixed and lower quality amber was sold from around ticals to 4
rupee Rupee (, ) is the common name for the currency, currencies of Indian rupee, India, Mauritian rupee, Mauritius, Nepalese rupee, Nepal, Pakistani rupee, Pakistan, Seychellois rupee, Seychelles, and Sri Lankan rupee, Sri Lanka, and of former cu ...
s per seer. Pieces that were considered high quality or fit for use as ornamentation were described as expensive, and price varied depending on the clarity and color of the amber. Women of the valley were noted to wear amber earrings as part of their jewelry. In 1885 the
Konbaung dynasty The Konbaung dynasty (), also known as the Third Burmese Empire (တတိယမြန်မာနိုင်ငံတော်), was the last dynasty that ruled Burma from 1752 to 1885. It created the second-largest empire in history of Mya ...
was annexed to the
British Raj The British Raj ( ; from Hindustani language, Hindustani , 'reign', 'rule' or 'government') was the colonial rule of the British The Crown, Crown on the Indian subcontinent, * * lasting from 1858 to 1947. * * It is also called Crown rule ...
and a survey of the area was conducted by Dr. Fritz Noetling on behalf of the Geological Survey of India. The final research before Burmese independence in 1947 was conducted by Dr. H.L. Chhibber in 1934, who provided the most detailed description of Burmite occurrences.


History of research

While a 1892 study considered the amber likely to be
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
in age, a study by
Theodore Dru Alison Cockerell Theodore Dru Alison Cockerell (22 August 1866 – 26 January 1948) was an American entomologist and Systematics, systematic biologist who published nearly 4,000 papers, some of them only a few lines long. Cockerell's speciality was the insect or ...
in 1916 noted archaic nature of the insects, and concluded that the amber must be older, and possibly as old as the Late Cretaceous. Various later authors during the mid-late 20th century suggested either a
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
-
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
or an Late Cretaceous age. A Cretaceous age was confirmed during the early 2000s. While during the 20th century and the first decade of the 21st century research on the deposit was low intensity, during the mid-2010s there was a great increase number of papers published on the deposit, numbering hundreds every year, with a plurality coming from Chinese researchers, rising from a few dozen species described per year prior to 2014 to over 350 in 2020, though this number had fallen to just over 250 by 2023. Both the great increase in the amber volume coming out of the mines, as well as a 2013 conference and a 2015 Burmese amber special issue in the journal '' Cretaceous Research'' have been cited as factors in the boom of publications.


Modern exploitation and controversy

Leeward Capital Corp, a small Canadian mining firm, began exporting amber from Myanmar during the 1990s, resulting in the description of a number of fossil species from the deposit. During the late 2000s, Burmese amber production began to increase as a result of the exhaustion of Chinese amber mines and greater trade between Myanmar and China in general, with the amber trade being initially unaffected by the outbreak of renewed conflict between the Kachin Independence Army, an armed rebel group seeking to scede from Myanmar, and the Burmese armed forces (
Tatmadaw The Tatmadaw, also known as the Sit-Tat, is the armed forces of Myanmar (formerly Burma). It is administered by the Ministry of Defence and composed of the Myanmar Army, the Myanmar Navy and the Myanmar Air Force. Auxiliary services include ...
) in 2011. Despite the conflict between them, there was apparently a tacit agreement between the two groups to keep the amber trade flowing, with the KIA controlling the mines, while the Tatmadaw controlled their export to China, with both collecting taxes on the trade. In 2016, the KIA took full control of the amber mines, and controlled amber export via "various licenses, taxes, restrictions on the movement of labor and enforced auctions". In June 2017 the
Tatmadaw The Tatmadaw, also known as the Sit-Tat, is the armed forces of Myanmar (formerly Burma). It is administered by the Ministry of Defence and composed of the Myanmar Army, the Myanmar Navy and the Myanmar Air Force. Auxiliary services include ...
seized control of the mines from the KIA. The main amber market in Myanmar is
Myitkyina Myitkyina (, ; Jingpho language, Jinghpaw: ''Myitkyina'', ; , ''Sèna'') is the capital city of Kachin State in Myanmar (Burma), located from Yangon, and from Mandalay. In Burmese language, Burmese it means "near the big river", and Myitkyina i ...
. Most Burmese amber is exported (primarily smuggled) into China, with the primary Chinese market being in Tengchong,
Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
, with an estimated 100 tonnes of Burmese amber passing into the city in 2015, with a then estimated value between five and seven billion yuan, where it is primarily used for jewelry. Burmese amber was estimated to make up 30% of Tenchong's gemstone market (the rest being Myanmar Jade), and was declared one of the city's eight main industries by the local government. The presence of calcite veins are a major factor in determining the gem quality of pieces, with pieces with a large number of veins having significantly lower value. The working conditions at the mines have been described as extremely unsafe, down deep pits barely wide enough to crawl through, with no accident compensation. Sales of amber were alleged to help fund the Kachin conflict by various news organisations in 2019. Interest in this discussion rose in March 2020 after the highly publicised description of ''Oculudentavis,'' which made the cover of ''Nature''. On April 21, 2020, the
Society of Vertebrate Paleontology The Society of Vertebrate Paleontology (SVP) is a professional organization that was founded in the United States in 1940 to advance the science of vertebrate paleontology around the world. Mission and activities SVP has about 2,300 members inte ...
(SVP) published a letter of recommendation to journal editors asking for "a moratorium on publication for any fossil specimens purchased from sources in Myanmar after June 2017 when the Myanmar military began its campaign to seize control of the amber mining". On April 23, 2020 '' Acta Palaeontologica Polonica'' stated that it would not accept papers on Burmese amber material collected from 2017 onwards, after the Burmese military took control of the deposit, requiring "certification or other demonstrable evidence, that they were acquired before the date both legally and ethically". On May 13, 2020, the '' Journal of Systematic Palaeontology'' published an editorial stating that it would no longer consider papers based whole or in part on Burmese amber material, regardless of whether in historic collections or not. On 30 June 2020, a statement from the International Palaeoentomological Society was published in response to the SVP, criticising the proposal to ban publishing on Burmese amber material. In August 2020, a comment from over 50 authors was published in '' PalZ'' responding to the SVP statement. The authors disagreed with the proposal of a moratorium, describing the focus on the Burmese amber as "arbitrary" and that "The SVP's recommendation for a moratorium on Burmese amber affects fossil non-vertebrate research much more than fossil vertebrate research and clearly does not represent this part of the palaeontological community." The conclusion that Burmese amber funded the Tatmadaw was disputed by George Poinar and Sieghard Ellenberger, who found that the supply of amber collapsed after the 2017 takeover of the mines by the Tatmadaw, and that most of the current circulation of amber in Chinese markets was extracted prior to 2017. A story in ''Science'' in 2019 stated: "Two former mine owners, speaking through an interpreter in phone interviews, say taxes have been even steeper since government troops took control of the area. Both shut their mines when they became unprofitable after the government takeover, and almost all deep mines are now out of business, dealers here corroborate. Only shallow mines and perhaps a few secret operations are still running." There were around 200,000 miners working in the Hukawng valley mines prior to the takeover by the Tatmadaw, which shrunk to 20,000 or less after the military operations. Adolf Peretti, a gemologist who owns a museum with Burmese amber specimens, noted that the 2017 cutoff suggested by the SVP does not take into account that the export of Burmese amber prior to 2017 was also funding internal conflict in Myanmar due to the control by the KIA. Much of the amber cutting since 2017 has been done in
internally displaced person An internally displaced person (IDP) is someone who is forced to leave their home but who remains within their country's borders. They are often referred to as refugees, although they do not fall within the legal definitions of a refugee. I ...
camps, under humanitarian and non-conflict conditions.


Other Burmese ambers

Other deposits of amber are known from several regions in Myanmar, with noted deposits in the Shwebo District of the
Sagaing Region Sagaing Region (, ; formerly Sagaing Division) is an administrative divisions of Myanmar, administrative region of Myanmar, located in the north-western part of the country between latitude 21° 30' north and longitude 94° 97' east. It is border ...
, from the
Pakokku Pakokku (, ) is the largest city in the Magway Region of Myanmar. It is situated about 30 km northeast of Bagan on the Irrawaddy River. It is the administration seat of Pakokku Township, Pakokku District and Gangaw District. Pakokku Bridge ...
and Thayet districts of
Magway Region Magway Region (, ; formerly Magway Division) is an administrative divisions of Myanmar, administrative division in central Myanmar. It is the second largest of Myanmar's seven divisions, with an area of . Pa-de Dam (ပဒဲဆည်) is one of ...
and the
Bago District Bago District () is a Districts of Burma, district of the Bago Region in central Burma, Burma (Myanmar). The region's capital, Bago, Burma, Bago, is located in the district. Bago District is bordered by Mon State to the southeast, Kayin State to th ...
of the
Bago Region Bago Region (, ; formerly Pegu Division and Bago Division) is an administrative divisions of Myanmar, administrative region of Myanmar, located in the southern central part of the country. It is bordered by Magway Region and Mandalay Region to ...
.


Tilin amber

A 2018 study on an amber deposit from Tilin in central Myanmar indicated that deposit to be 27 million years younger than the Hukawng deposit, dating to approximately 72 million years old, placing it in the latest
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
age. The deposit was associated with an overlying tuffaceous layer, and underlying nodules of brown sandstone yielded remains of the ammonite '' Sphenodiscus.'' Within a number of arthropod specimens were described though much more poorly preserved than specimens in the Hukawng amber. These include members of
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typi ...
(
Braconidae The Braconidae are a family of parasitoid wasps. After the closely related Ichneumonidae, braconids make up the second-largest family in the order Hymenoptera, with about 17,000 recognized species and many thousands more undescribed. One analysis ...
,
Diapriidae The Diapriidae are a family of parasitoid wasps. These tiny insects have an average length of 2–4 mm and never exceed 8 mm. They typically attack larvae and pupae of a wide range of insects, especially flies. The about 2,300 described ...
, Scelionidae)
Diptera Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advance ...
(
Ceratopogonidae Ceratopogonidae is a family of flies commonly known as no-see-ums, sand flies or biting midges, generally in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic. A 2025 study fro ...
,
Chironomidae Chironomidae , commonly known as non-biting midges or chironomids , are a family of Nematoceran flies with a global distribution. They are closely related to the families Ceratopogonidae, Simuliidae, and Thaumaleidae. Although many chironomid ...
) Dictyoptera ( Blattaria,
Mantodea Mantises are an Order (biology), order (Mantodea) of insects that contains List of mantis genera and species, over 2,400 species in about 460 genera in 33 families. The largest family is the Mantidae ("mantids") ...
)
planthopper A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment ...
s, Berothidae and bark lice ( Lepidopsocidae) as well as extant ant subfamilies
Dolichoderinae Dolichoderinae is a subfamily of ants, which includes species such as the Argentine ant (''Linepithema humile''), the erratic ant, the odorous house ant, and the cone ant. The subfamily presents a great diversity of species throughout the wor ...
and tentatively
Ponerinae Ponerinae, the ponerine ants, is a subfamily of ants in the Poneromorph subfamilies group, with about 1,600 species in 47 extant genera, including '' Dinoponera gigantea'' - one of the world's largest species of ant. Mated workers have replac ...
, as well as fragments of
moss Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
.


Hkamti amber

The Hkamti site is located ca. 90 km southwest of the Angbamo site and predominantly consists of limestone, interbedded with mudstone and
tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock co ...
, the amber is found in the unconsolidated mudstone/tuff layers. A crinoid was found attached to one amber specimen, alongside marine plant remains in the surrounding sediment, indicating deposition in a shallow marine setting. The amber is generally red-brown, and yellow colouration is rare, the amber is generally found as angular clasts, indicating short transport distance and is more brittle than other northern Myanmar ambers. Zircon dating has constrained the age of the deposit to the early Albian, c. 110 Ma, significantly older than the dates obtained from other deposits. Fauna found in the amber include:
Archaeognatha The Archaeognatha are an order of apterygotes, known by various common names such as jumping bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the sa ...
,
Diplopoda Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...
,
Coleoptera Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
,
Araneae Spiders ( order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
,
Trichoptera The caddisflies (order Trichoptera) are a group of insects with aquatic larvae and terrestrial adults. There are approximately 14,500 described species, most of which can be divided into the suborders Integripalpia and Annulipalpia on the basis ...
,
Neuroptera The insect order (biology), order Neuroptera, or net-winged insects, includes the lacewings, mantidflies, antlions, and their relatives. The order consists of some 6,000 species. Neuroptera is grouped together with the Megaloptera (alderflies, f ...
,
Psocodea Psocodea is a taxonomic group of insects comprising the bark lice, book lice and parasitic lice. It was formerly considered a superorder, but is now generally considered by entomologists as an order. Despite the greatly differing appearance of ...
, Isoptera
Diptera Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advance ...
,
Orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – gras ...
, Pseudoscorpionida,
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typi ...
and Thysanoptera. A lizard, '' Retinosaurus,'' has also been described from the locality.


See also

*


References

{{Reflist


External links


Blood Amber Military resource grab clears out indigenous peoples in Kachin State's Hugawng Valley ''Kachin Development Networking Group''
Paleontological sites of Asia Paleontology in Myanmar