Mopane (other)
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''Colophospermum mopane'', commonly called mopane, mopani, butterfly tree, turpentine tree, or balsam tree, is a tree in the
legume Legumes are plants in the pea family Fabaceae (or Leguminosae), or the fruit or seeds of such plants. When used as a dry grain for human consumption, the seeds are also called pulses. Legumes are grown agriculturally, primarily for human consum ...
family (
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
), that grows in hot, dry, low-lying areas, in elevation, in parts of
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
. The tree only occurs in Africa and is the only
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
in
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Colophospermum''. Its distinctive
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
-shaped (bifoliate) leaf and thin kidney-shaped/nearly semi-circular
seed pod This page provides a glossary of plant morphology. Botanists and other biologists who study plant morphology use a number of different terms to classify and identify plant organs and parts that can be observed using no more than a handheld magnify ...
make it easy to identify. Though hard and dense and difficult to work with, it is valued timber in all sorts of construction due to its termite tolerance. Together with
camel thorn Camelthorn or camel thorn is a common name for several plants and may refer to: *''Acacia erioloba'', a tree native to southern Africa *''Alhagi ''Alhagi'' is a genus of Old World plants in the family Fabaceae. They are commonly called came ...
and leadwood, one of the three regionally important
firewood Firewood is any wooden material that is gathered and used for fuel. Generally, firewood is not heavily processed, and is in some sort of firelog, recognizable log or branch form, compared to other forms of wood fuel like pellet fuel, pellets. ...
trees, due to the enduring heat, and these woods are also some of the preferred use culinarily for
braai Barbecue varies by the type of meat, sauce, Spice rub, rub, or other flavorings used, the point in barbecuing at which they are added, the role smoke plays, the equipment and fuel used, cooking temperature, and cooking time. The meat may be wh ...
.


Range and habitat

The mopane is found on low-lying ground from (Mozambique) to (Zimbabwe).


Mopane ecoregions

Native to
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
, its habitat is divided crudely into two regions: the "Angola region", i.e.,
Angolan mopane woodlands Angolan mopane woodlands are situated in southwestern Angola, extending into northern Namibia. This ecosystem surrounds Etosha Pan, which is considered a separate ecoregion. The mopane trees are the main type of vegetation. Location and descript ...
ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
, which includes Southern
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
and northwestern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
, and the "Zambezian region", i.e. the
Zambezian and mopane woodlands The Zambezian and mopane woodlands is a tropical and subtropical grasslands, savannas, and shrublands ecoregion of southeastern Africa. The ecoregion is characterized by the mopane tree ''( Colophospermum mopane)'', and extends across portion ...
ecoregion extending over the lowlands of the
Zambezi River The Zambezi (also spelled Zambeze and Zambesi) is the fourth-longest river in Africa, the longest east-flowing river in Africa and the largest flowing into the Indian Ocean from Africa. Its drainage basin covers , slightly less than half of t ...
and its tributaries across the countries of
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
, Southern
Malawi Malawi, officially the Republic of Malawi, is a landlocked country in Southeastern Africa. It is bordered by Zambia to the west, Tanzania to the north and northeast, and Mozambique to the east, south, and southwest. Malawi spans over and ...
, other parts of Namibia,
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
,
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
,
Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
,
Eswatini Eswatini, formally the Kingdom of Eswatini, also known by its former official names Swaziland and the Kingdom of Swaziland, is a landlocked country in Southern Africa. It is bordered by South Africa on all sides except the northeast, where i ...
/Swaziland and northern
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
.


Soil and morphology

While it prefers slightly acidic deep clay soil that are and , it also grows in
alkaline In chemistry, an alkali (; from the Arabic word , ) is a basic salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as a base that dissolves in water. A solution of a soluble base has a pH greater than 7.0. The ...
(high lime content) soils which are shallow and not well drained. It also grows in
alluvial soil Alluvium (, ) is loose clay, silt, sand, or gravel that has been deposited by running water in a stream bed, on a floodplain, in an alluvial fan or beach, or in similar settings. Alluvium is also sometimes called alluvial deposit. Alluvium is ...
s (soil deposited by rivers). Where it occurs, it is often the dominant tree species, frequently forming homogeneous stands. Within the Zambezian region described above, the trees' height varying considerably, and larger, single-stemmed trees of are found in open savannah woodlands on the (sandy) alluvial soil, forming
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
woodlands, and even reaching 18m in height for the so-called "cathedral mopane" of Zambia. The same tree is also found stunted as (multi-stemmed)
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
bery about in clayey soil or impermeable alkaline soil. Thus water or drainage seems to be a cause factor for stunting, though other causes have been hypothesized. These stunted growths are sometimes called "mopane scrub" (for shrub). The shrubs will not transition into tall woods. This tree will not grow well outside hot, mostly frost-free areas with low to moderate rainfall (as low as 100mm per year, or averaging 400–500mm per year).


Uses

Mopane wood is one of southern Africa's heaviest, with an average
density Density (volumetric mass density or specific mass) is the ratio of a substance's mass to its volume. The symbol most often used for density is ''ρ'' (the lower case Greek letter rho), although the Latin letter ''D'' (or ''d'') can also be u ...
range of at 12% moisture. and is difficult to work because of its hardness.Meier, Eric (2016)
The Wood Database
' Mopane. Retrieved 24 June 2013.
However it is also
termite Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...
resistant making it one of the most preferred local timbers for construction, and for this reason it has long been used for building houses and fences, as railway sleepers and as pit props. The termite-resistance and rich, reddish colouring also make it popular for
parquet Parquet (; French for "a small compartment") is a geometric mosaic of wood pieces used for decorative effect in flooring. Parquet patterns are often entirely geometrical and angular—squares, triangles, lozenges—but may contain curves. T ...
flooring. Outside Africa, mopane is gaining popularity as a heavy, decorative wood, its uses including
aquarium An aquarium (: aquariums or aquaria) is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. fishkeeping, Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aquati ...
bogwood ornaments, similarly as
driftwood Driftwood is a wood that has been washed onto a shore or beach of a sea, lake, or river by the action of winds, tides or waves. It is part of beach wrack. In some waterfront areas, driftwood is a major nuisance. However, the driftwood provides ...
decor for
terrarium A terrarium ( terraria or terrariums) is a glass container containing soil and plants in an environment different from the surroundings. It is usually a sealable container that can be opened for maintenance or to access the plants inside; howeve ...
s or gardens, or bases for lamps or sculptures. It is also increasingly being used in the construction of musical instruments, particularly
woodwind Woodwind instruments are a family of musical instruments within the greater category of wind instruments. Common examples include flute, clarinet, oboe, bassoon, and saxophone. There are two main types of woodwind instruments: flutes and Ree ...
. Suitable quality African blackwood (''
Dalbergia melanoxylon ''Dalbergia melanoxylon'' (African blackwood, grenadilla, or mpingo) in French ''Granadille d'Afrique'' is a flowering plant in the family Fabaceae, native to seasonally dry regions of Africa from Senegal east to Eritrea, to southern regions of ...
''), traditionally used for
clarinet The clarinet is a Single-reed instrument, single-reed musical instrument in the woodwind family, with a nearly cylindrical bore (wind instruments), bore and a flared bell. Clarinets comprise a Family (musical instruments), family of instrume ...
s, is becoming harder to find. Mopane is fairly oily, seasons very well with few splits or shakes, and produces instruments of a warm, rich tone. Clarinets made of mopane are offered by the manufacturers
Schwenk & Seggelke Seggelke Klarinetten (GmbH & Co. KG), is a German clarinet manufacturer based in Bamberg in the Bavarian Upper Franconia. The company manufactures clarinets of a variety of fingering systems. A specialty of the company is the reproduction o ...
,
Wurlitzer The Rudolph Wurlitzer Company, usually referred to as simply Wurlitzer, is an American company started in Cincinnati in 1853 by German immigrant (Franz) Rudolph Wurlitzer. The company initially imported stringed, woodwind and brass instruments ...
,
F. Arthur Uebel The company F. Arthur Uebel GmbH (FAU) is a German manufacturer of clarinets with headquarters in Wiesbaden and production facilities in Markneukirchen (Saxony).Gisbert König, ''Eine Reise durch die deutsche Klarinettenbaulandschaft'' (''A journe ...
and
Buffet Crampon Buffet Crampon SAS is a French manufacturer of wind instruments based in Mantes-la-Ville, in the Yvelines department. The company is the world market leader in the production of Boehm system clarinets. Its subsidiary, Buffet Crampon Deuts ...
. Mopane twigs been traditionally used as a sort of tooth brush (chewing stick to clean teeth), and the leaves for
folk remedy Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
in healing the wound after
ritual A ritual is a repeated, structured sequence of actions or behaviors that alters the internal or external state of an individual, group, or environment, regardless of conscious understanding, emotional context, or symbolic meaning. Traditionally ...
extraction of teeth. Various parts of the tree (seed, leaf, bark, root) are used in the
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
of Southern African for a wide range of symptoms. The bark to make twine or rope and for tanning leather, The wood is also used to make charcoal and for
braai Barbecue varies by the type of meat, sauce, Spice rub, rub, or other flavorings used, the point in barbecuing at which they are added, the role smoke plays, the equipment and fuel used, cooking temperature, and cooking time. The meat may be wh ...
wood.


Food source

The tree is a major food source for the mopane worm, the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
of the
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
''
Gonimbrasia belina ''Gonimbrasia belina'' is a species of emperor moth which is native to the warmer parts of southern Africa. Its large edible caterpillar, known as the mopane worm, madora, amacimbi “pigeon moth”, masonja or Seboko sa Mongana, feeds primari ...
''. The caterpillars are rich in protein and are eaten by people. The mopane worm is rich in crude fats and contains vitamins and minerals, such as iron, calcium and phosphorus. Another edible caterpillar is that of ''
Gynanisa maja Kibembe ''Gynanisa maja'', the speckled emperor (; ) is a moth of the family Saturniidae. The species was first described by Johann Christoph Friedrich Klug in 1836. It is known from South Africa to eastern Africa (up to Angola and Zambia). ''Gyn ...
'' (speckled emperor moth; in
Bemba Bemba may refer to: Languages and ethnic groups * Bemba language, or Chibemba, a Bantu language spoken in Zambia * Bemba people, or AbaBemba, of Zambia * Bemba, a dialect of the Buyu language of the Democratic Republic of the Congo People * Jea ...
) which also uses mopane as host tree. And when infested with the "mopane fly" or "mopane
psyllid Psyllidae, the jumping plant lice or psyllids, are a family of small plant-feeding insects that tend to be very host-specific, i.e. each plant-louse species only feeds on one plant species (monophagous) or feeds on a few closely related plants ( ...
" ('' Retroacizzia mopani'' ''
Arytaina ''Arytaina'' is a genus of mostly Palaearctic Psylloidea, plant lice belonging to the subfamily Psyllinae (it was previously placed in a subfamily Arytaininae); this genus was erected by Arnold Förster in 1848.Foerster A (1848) Uebersicht der G ...
mopani'' ) that feeds on the tree, the tree produces edible wax-like gums on its leaves called "mopane manna" which are collected and eaten by humans and monkeys. The tree also acts as a foodplant for the
wild silk Wild silks have been known and used in many countries from early times, although the scale of production is far smaller than that from cultivated silkworms. Silk cocoons and nests often resemble paper or cloth, and their use has arisen independe ...
moth, ''
Gonometa rufobrunnea ''Gonometa'' is a genus of moths in the family Lasiocampidae. The genus was erected by Francis Walker (entomologist), Francis Walker in 1855.
''. Cocoons of the moth are harvested as wild
silk Silk is a natural fiber, natural protein fiber, some forms of which can be weaving, woven into textiles. The protein fiber of silk is composed mainly of fibroin and is most commonly produced by certain insect larvae to form cocoon (silk), c ...
, to make cloth. The Mopane worm creates employment and serves as a source of income for the majority of rural women. Harvesters sell it in villages, towns or to the trader. The tree is also considered an important food resource for
animal husbandry Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, animal fiber, fibre, milk, or other products. It includes day-to-day care, management, production, nutrition, selective breeding, and the raising ...
(
goat herding Goat farming involves the raising and breeding of domestic goats (''Capra aegagrus hircus'') as a branch of animal husbandry. People farm goats principally for their meat, milk, fibre and skins. Goat farming can be very suited to production ...
), as goats can be allowed to browse on the tree's leaves (cf. fig. above). They are also browsed by
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
and by
game animals Game or quarry is any wild animal hunted for animal products (primarily meat), for recreation (" sporting"), or for trophies. The species of animals hunted as game varies in different parts of the world and by different local jurisdictions, thou ...
such as
elephant Elephants are the largest living land animals. Three living species are currently recognised: the African bush elephant ('' Loxodonta africana''), the African forest elephant (''L. cyclotis''), and the Asian elephant ('' Elephas maximus ...
s,
giraffe The giraffe is a large Fauna of Africa, African even-toed ungulate, hoofed mammal belonging to the genus ''Giraffa.'' It is the Largest mammals#Even-toed Ungulates (Artiodactyla), tallest living terrestrial animal and the largest ruminant on ...
s,
water buffalo The water buffalo (''Bubalus bubalis''), also called domestic water buffalo, Asian water buffalo and Asiatic water buffalo, is a large bovid originating in the Indian subcontinent and Southeast Asia. Today, it is also kept in Italy, the Balkans ...
s and
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
s in
game farm A game farm is a place where game animals are raised to stock wildlife areas for hunting. The term also includes places where such animals are raised to be sold as food or for photography. Their existence has been exemplified within the South Afr ...
s and
nature reserve A nature reserve (also known as a wildlife refuge, wildlife sanctuary, biosphere reserve or bioreserve, natural or nature preserve, or nature conservation area) is a protected area of importance for flora, fauna, funga, or features of geologic ...
s.


Etymology

The genus name ''Colophospermum'' is a compound from Greek "resin" + Latin ''spermum'' "seed". The former derives from Colophon", which was the birthplace of
Homer Homer (; , ; possibly born ) was an Ancient Greece, Ancient Greek poet who is credited as the author of the ''Iliad'' and the ''Odyssey'', two epic poems that are foundational works of ancient Greek literature. Despite doubts about his autho ...
in
Ionia Ionia ( ) was an ancient region encompassing the central part of the western coast of Anatolia. It consisted of the northernmost territories of the Ionian League of Greek settlements. Never a unified state, it was named after the Ionians who ...
, famous for its
rosin Rosin (), also known as colophony or Greek pitch (), is a resinous material obtained from pine trees and other plants, mostly conifers. The primary components of rosin are diterpenoids, i.e., C20 carboxylic acids. Rosin consists mainly of r ...
, and is a reference to the seed's
aromatic In organic chemistry, aromaticity is a chemical property describing the way in which a conjugated system, conjugated ring of unsaturated bonds, lone pairs, or empty orbitals exhibits a stabilization stronger than would be expected from conjugati ...
(turpentine-smelling) component. The species name ''mopane'' is from its common name in
Bantu language The Bantu languages (English: , Proto-Bantu language, Proto-Bantu: *bantʊ̀), or Ntu languages are a language family of about 600 languages of Central Africa, Central, Southern Africa, Southern, East Africa, Eastern and Southeast Africa, South ...
s for the tree throughout most of its range. Thus the nomenclature, starting with areas in or around South Africa are: in
Northern Sotho Sepedi, also known as Northern Sotho, is one of South Africa’s twelve official languages and belongs to the Bantu language family, specifically the Sotho-Tswana group. The language is spoken mainly in Limpopo Province, and to a lesser exten ...
/Sepedi (northern
Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name ''Transvaal''. * South African Republic (1856–1902; ...
, current
Limpopo Limpopo () is the northernmost Provinces of South Africa, province of South Africa. It is named after the Limpopo River, which forms the province's western and northern borders. The term Limpopo is derived from Rivombo (Livombo/Lebombo), a ...
province), in Tsuwana (western Transvaal, northern
Cape A cape is a clothing accessory or a sleeveless outer garment of any length that hangs loosely and connects either at the neck or shoulders. They usually cover the back, shoulders, and arms. They come in a variety of styles and have been used th ...
, Botswana) but in
Tsonga Tsonga may refer to: * Tsonga language, a Bantu language spoken in southern Africa * Tsonga people, a large group of people living mainly in southern Mozambique and South Africa. * Jo-Wilfried Tsonga Jo-Wilfried Tsonga (; born 17 April 1985) ...
(eastern Transvaal). It is known as in
Chichewa Chewa ( ; also known as Nyanja ) is a Bantu language spoken in Malawi and a recognised minority in Zambia and Mozambique. The noun class prefix ''chi-'' is used for languages, so the language is often called or Chinyanja. In Malawi, the name wa ...
(Malawi), in Lozi (fmr.
Barotseland Barotseland (Lozi language, Lozi: ''Mubuso Bulozi'') is a region between Namibia, Angola, Botswana, Zimbabwe including half of north-western province, southern province, and parts of Lusaka Province, Lusaka, Central Province, Zambia, Central, ...
, western Zambia), in Thimbukushu (Namibia), in
Herero Herero may refer to: * Herero people, a people belonging to the Bantu group, with about 240,000 members alive today * Herero language, a language of the Bantu family (Niger-Congo group) * Herero and Nama genocide * Herero chat, a species of bird ...
(Namibia, Botswana), in
Ovambo Ovambo may refer to: *Ovambo language, Bantu language of Namibia **Ovambo people, Bantu people of Namibia *Ovamboland, former Bantustan in South West Africa (now Namibia) *Ovambo sparrowhawk The Ovambo or Ovampo sparrowhawk, also known as Hilge ...
(southern Angola, northern Namibia).: Table 1. Vernacular names of Colophospermum mopame Also spelt ''mupani'', ''mopani''. In Afrikaans it goes by or "mopanie wood" or "turpentine tree". It is called in
Shona Shona often refers to: * Shona people, a Southern African people ** Shona language, a Bantu language spoken by Shona people today ** Shona languages, a wider group of languages defined in the early 20th century ** Kingdom of Zimbabwe, a Shona stat ...
(
Sabi Sabi may refer to: * A member of an ethnoreligious group known as the Mandaeans (Sabians), in Arabic * Sabi (Korea), an ancient capital of Baekje * Sabi (dog), an Australian special forces dog * Save River (Africa), flows through Zimbabwe and Mozam ...
valley, Zimbabwe), ''chanate'' in southern Mozambique, and in
Umbundu Umbundu, or South Mbundu (autonym ''úmbúndú''), one of many Bantu languages, is the most widely-spoken autochthonous language of Angola. Its speakers are known as ''Ovimbundu'' and are an ethnic group constituting a third of Angola's populati ...
(Angola).


Gallery

File:Colophospermum mopane 2122.jpg, Inflorescences produced during mid-summer File:Colophospermum mopane 2120.jpg, The wind-pollinated flower File:Colophospermum mopane MHNT.BOT.2015.2.6.jpg, Dry foliage, seed pods and two seeds (bottom right) - MHNT File:Colophospermum mopane - Seeds.jpg, The fruit are pods containing one seed each File:Colophospermum mopane 50D 0484.jpg, Mopane worms, locally a staple food, consuming the foliage File:Speckled Emperor (Gynanisa maja) caterpillar on mopane twig ... (51867253398).jpg, The likewise edible caterpillar of the speckled emperor moth, defoliating a sprig File:Mopane_seed_and_leaves.JPG, Winter foliage and dry seed pods in Namibia


See also

*
List of Southern African indigenous trees This is a list of Southern African trees, shrubs, suffrutices, geoxyles and Liana, lianes, and is intended to cover Angola, Botswana, Eswatini, Lesotho, Malawi, Mozambique, Namibia, South Africa, Zambia and Zimbabwe. The notion of 'indigenous' i ...


Explanatory notes


References

*


Bibliography

* * {{Authority control Flora of South Tropical Africa Flora of Southern Africa Trees of South Africa Trees of Angola Drought-tolerant trees Tropical and subtropical dry broadleaf forests Detarioideae Monotypic Fabaceae genera Plants described in 1865