The belligerent sculpin (''Megalocottus platycephalus''), or flathead sculpin, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...

, the typical sculpins. This species occurs in the northern

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...

Taxonomy

The belligerent sculpin was first formally described in 1814 as ''Cottus platycephalus'' by the German

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...

with its type locality given as

Kamchatka The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively. Immediately offshore along the Pacific ...

and the

Sea of Okhotsk The Sea of Okhotsk; Historically also known as , or as ; ) is a marginal sea of the western Pacific Ocean. It is located between Russia's Kamchatka Peninsula on the east, the Kuril Islands on the southeast, Japan's island of Hokkaido on the sou ...

. In 1861 the American

biologist A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual Cell (biology), cell, a multicellular organism, or a Community (ecology), community of Biological inter ...

Theodore Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist, and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural hist ...

classified this species in the

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

Megalocottus ''Megalocottus'' is a small genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the western Pacific Ocean. Taxonomy ''Megalocottus'' was first proposed as a monospecific genus in 1 ...

''. Some authorities consider the belligerent sculpin to be the still the only species in its genus, either recognising the Southern flathead sculpin (''M. taeniopterus'') as a

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

of ''M. platycephalus''. Fishbase still recognises two species within the genus ''Megalocottus''. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''playcephalus'' means "flat head", a reference to the wide, flat head with a projecting lower jaw.

Description

The belligerent sculpin has a broad, strongly flattened head, much wider than it is deep with a projecting lower jaw. The top and the sides of the head are very warty and there are large spines or bumps behind the eye and on the nape, these may have short cirri on them. The first and second spines on the

preoperculum This glossary of ichthyology is a list of definitions of terms and concepts used in ichthyology, the study of fishes. A B C ...

are curved, the third and fourth are poorly developed. There are irregular rows of spiny plates over and under the

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

. The colour is grey-brown marked with yellow or white spotting and blotching. The fins have spots and bars. This species reaches a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of .

Distribution and habitat

The belligerent sculpin is found in the northern Pacific Ocean. Its range extends from the

Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, it ...

, through the Sea of Okhotsk and the northern

Kuril Islands The Kuril Islands or Kurile Islands are a volcanic archipelago administered as part of Sakhalin Oblast in the Russian Far East. The islands stretch approximately northeast from Hokkaido in Japan to Kamchatka Peninsula in Russia, separating the ...

and the southeastern

Kamchatka Peninsula The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively. Immediately offshore along the Pacific ...

east to the

Alaska Peninsula The Alaska Peninsula (also called Aleut Peninsula or Aleutian Peninsula, ; Sugpiaq language, Sugpiaq: ''Aluuwiq'', ''Al'uwiq'') is a peninsula extending about to the southwest from the mainland of Alaska and ending in the Aleutian Islands. T ...

and north along the

Bering Sea The Bering Sea ( , ; rus, Бе́рингово мо́ре, r=Béringovo móre, p=ˈbʲerʲɪnɡəvə ˈmorʲe) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasse ...

coasts of Alaska and Russia to the

Chukchi Sea The Chukchi Sea (, ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west by the Long Strait, off Wrangel Island, and in the east by Point Barrow, Alaska, ...

. It is a

demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...

, typically found in the subtidal zone down to depths of , although has been claimed. It is found on substrates of sand, mud and gravel, especially where there are beds of ''

Fucus ''Fucus'' is a genus of brown algae found in the intertidal zones of rocky seashores almost throughout the world. Description and life cycle The thallus is perennial with an irregular or disc-shaped holdfast or with haptera. The erect portion ...

'' and ''

Zostera ''Zostera'' is a small genus of widely distributed seagrasses, commonly called marine eelgrass, or simply seagrass or eelgrass. The genus ''Zostera'' contains 15 species. Ecology '' Zostera marina'' is found on sandy substrates or in estuarie ...

''. It is generally found in brackish water close to river mouths, near to the shore and in shallow water.

Biology

The belligerent sculpin's biology is not well known. They are oviparous. In the Spring they move closer inshore and into estuaries, even into the lower reaches of rivers. The juveniles feed on algae, and crustaceans such as

copepod Copepods (; meaning 'oar-feet') are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (living in the water column), some are benthos, benthic (living on the sedimen ...

cumacea Cumacea is an order (biology), order of small marine crustaceans of the superorder Peracarida, occasionally called hooded shrimp or comma shrimp. Their unique appearance and uniform body plan makes them easy to distinguish from other crustaceans. ...

ns,

amphipod Amphipoda () is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods () range in size from and are mostly detritivores or scavengers. There are more than 10,700 amphipod species cur ...

s, and

mysid Mysida is an order of small, shrimp-like crustaceans in the malacostracan superorder Peracarida. Their common name opossum shrimps stems from the presence of a brood pouch or "marsupium" in females. The fact that the larvae are reared in thi ...

s while the adults have a similar diet but they also eat small fishes.

References

{{Taxonbar, from=Q2044901 Belligerent sculpin Fish described in 1814 Taxa named by Peter Simon Pallas