Martin Andreas Nowak (born April 7, 1965)
is an Austrian-born professor of mathematics and biology at
Harvard University
Harvard University is a Private university, private Ivy League research university in Cambridge, Massachusetts, United States. Founded in 1636 and named for its first benefactor, the History of the Puritans in North America, Puritan clergyma ...
. He is a researcher in
evolutionary dynamics,
and has made contributions to the fields of
evolutionary theory and
viral dynamics.
He held faculty positions at
Oxford University
The University of Oxford is a collegiate research university in Oxford, England. There is evidence of teaching as early as 1096, making it the oldest university in the English-speaking world and the second-oldest continuously operating u ...
and the
Institute for Advanced Study
The Institute for Advanced Study (IAS) is an independent center for theoretical research and intellectual inquiry located in Princeton, New Jersey. It has served as the academic home of internationally preeminent scholars, including Albert Ein ...
in Princeton, before being recruited by Harvard in 2003. During this time,
Jeffrey Epstein
Jeffrey Edward Epstein ( , ; January 20, 1953August 10, 2019) was an American financier and child sex offender. Born and raised in New York City, Epstein began his professional career as a teacher at the Dalton School, despite lacking a col ...
funded a portion of Nowak's work, helping to set up a center for studying cooperation in evolution. He was the director of Harvard's Program for Evolutionary Dynamics (PED) from 2003 until 2020. He was suspended from supervising undergraduate research for two years and his institute permanently closed due to Epstein’s continued use of a personal office in the PED building for over ten years after his conviction for
sex crimes.
Early life and education
Nowak was born in
Vienna
Vienna ( ; ; ) is the capital city, capital, List of largest cities in Austria, most populous city, and one of Federal states of Austria, nine federal states of Austria. It is Austria's primate city, with just over two million inhabitants. ...
,
Austria
Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
, on April 7, 1965.
He studied at
Albertus Magnus Gymnasium and the
University of Vienna
The University of Vienna (, ) is a public university, public research university in Vienna, Austria. Founded by Rudolf IV, Duke of Austria, Duke Rudolph IV in 1365, it is the oldest university in the German-speaking world and among the largest ...
, where he earned a doctorate in biochemistry and mathematics in 1989. During his studies, he collaborated with
Peter Schuster on
quasispecies theory, and with
Karl Sigmund on
evolution
Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
of cooperation. Nowak received the highest Austrian honors (
Sub auspiciis Praesidentis) upon receiving his degree.
In 1993, he completed his
habilitation
Habilitation is the highest university degree, or the procedure by which it is achieved, in Germany, France, Italy, Poland and some other European and non-English-speaking countries. The candidate fulfills a university's set criteria of excelle ...
at the Institute of Mathematics, University of Vienna. Later, in 2001, Nowak was elected as a member of the
Austrian Academy of Sciences.
Career
From 1989 to 1998, Nowak worked at the
University of Oxford
The University of Oxford is a collegiate university, collegiate research university in Oxford, England. There is evidence of teaching as early as 1096, making it the oldest university in the English-speaking world and the List of oldest un ...
with
Robert May. First, he was an
Erwin Schrödinger
Erwin Rudolf Josef Alexander Schrödinger ( ; ; 12 August 1887 – 4 January 1961), sometimes written as or , was an Austrian-Irish theoretical physicist who developed fundamental results in quantum field theory, quantum theory. In particul ...
Postdoctoral Scholar, then a
Junior Research Fellow at
Wolfson College and
Keble College. From 1992, he was a
Wellcome Trust
The Wellcome Trust is a charitable foundation focused on health research based in London, United Kingdom. It was established in 1936 with legacies from the pharmaceutical magnate Henry Wellcome (founder of Burroughs Wellcome, one of the predec ...
Senior Research Fellow.
From 1997 to 1998, Nowak was a professor of mathematical biology.
In 1998, Martin Nowak was recruited by the Institute for Advanced Study in Princeton, and was Head of the Institute's first Initiative in Theoretical Biology from 1998 until 2003. In 2003, Nowak joined
Harvard University
Harvard University is a Private university, private Ivy League research university in Cambridge, Massachusetts, United States. Founded in 1636 and named for its first benefactor, the History of the Puritans in North America, Puritan clergyma ...
as a Professor of Mathematics and Biology.
Nowak was also co-director, with
Sarah Coakley, of the Evolution and Theology of Cooperation project at
Harvard University
Harvard University is a Private university, private Ivy League research university in Cambridge, Massachusetts, United States. Founded in 1636 and named for its first benefactor, the History of the Puritans in North America, Puritan clergyma ...
, sponsored by the
Templeton Foundation, where he was also a member of their Board of Advisors. He was appointed Director of the Program for Evolutionary Dynamics (PED). The PED was funded with a large sum of money from the
Jeffrey Epstein VI Foundation. In 2003, Epstein had introduced himself as a science philanthropist, cementing the initial interaction with a large donation to Harvard.
Nowak's team had received US$6.5 million between 2003 and 2006, but they did not receive any additional funds after 2007.
After Epstein's 2008 conviction, Harvard president
Drew Faust decided that the university would no longer accept his donations.
report commissioned by the university, found that Nowak allowed Epstein to visit the PED offices more than 40 times after his conviction, to maintain an office with a phone line and webpage, and to interact with students at PED. In 2020, the university placed Nowak on paid academic leave for violation of campus policies including professional conduct and campus access.
In 2021, Harvard decided a proportionate response to the severity of Nowak's failure to follow Harvard policies, closing the institute founded with Epstein's money, and donating the remaining money to a foundation helping victims of sexual assault. Harvard also imposed a two year ban on Nowak that barred him from supervising undergraduate research or further supervising new graduate students or postdoctoral fellows. He was also banned from acting as the
principal investigator of new grants.
Nowak said he would "take the lessons from this time with me as I move forward".
The sanctions against Nowak were lifted in 2023.
Academic research
Martin Nowak contributed to various scientific disciplines, including evolutionary game theory, virology, cancer dynamics, the evolution of cooperation, and the origins of language. Throughout his career, Nowak has collaborated with scientists such as
Robert May,
Karl Sigmund, and
John Maynard Smith.
Nowak has authored books and scientific papers on topics in
evolutionary game theory,
cancer
Cancer is a group of diseases involving Cell growth#Disorders, abnormal cell growth with the potential to Invasion (cancer), invade or Metastasis, spread to other parts of the body. These contrast with benign tumors, which do not spread. Po ...
,
virus
A virus is a submicroscopic infectious agent that replicates only inside the living Cell (biology), cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Viruses are ...
es,
infectious disease
An infection is the invasion of tissue (biology), tissues by pathogens, their multiplication, and the reaction of host (biology), host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmis ...
, the
evolution of language, and the
evolution of cooperation.
At Oxford, he helped to establish the fields of virus dynamics and spatial games (which later became evolutionary graph theory). He maintained a collaboration with
Karl Sigmund in game theory.
He collaborated with
John Maynard Smith on genetic redundancy, with
Baruch Blumberg on hepatitis B virus,
and with
Robert May as well as George Shaw and
Andrew McMichael on HIV.
In 1990, Nowak and
Robert May proposed a mathematical model which aimed to explain the delay between
HIV infection and
AIDS
The HIV, human immunodeficiency virus (HIV) is a retrovirus that attacks the immune system. Without treatment, it can lead to a spectrum of conditions including acquired immunodeficiency syndrome (AIDS). It is a Preventive healthcare, pr ...
. in terms of the evolution of different strains of the virus during individual infections, to the point where the genetic diversity of the virus reaches a threshold whereby the immune system can no longer control it.
At Harvard, Nowak continued his work on virus dynamics, cancer dynamics, and evolutionary game theory. In 2004, he established evolutionary game dynamics in finite populations. In 2005 and 2006 he wrote papers establishing evolutionary graph theory. In 2006, he suggested that cooperation was a third fundamental principle of evolution beside mutation and selection. In 2007, he proposed a theory for the origin of life that views it through competition with its chemical precursors. In a 2008 paper, he suggested that punishment behavior does not encourage the evolution of cooperation, and describes that behavior as maladaptive.
In 2006, Nowak enunciated and unified the mathematical rules for the five understood bases of the evolution of cooperation (
kin selection
Kin selection is a process whereby natural selection favours a trait due to its positive effects on the reproductive success of an organism's relatives, even when at a cost to the organism's own survival and reproduction. Kin selection can lead ...
,
direct reciprocity,
indirect reciprocity,
network reciprocity, and
group selection
Group selection is a proposed mechanism of evolution in which natural selection acts at the level of the group, instead of at the level of the individual or gene.
Early authors such as V. C. Wynne-Edwards and Konrad Lorenz argued that the beha ...
) which he published as a paper in ''Science''. Nowak suggests that evolution is constructive because of cooperation, and that scientists might add “natural cooperation” as a third fundamental principle of evolution beside mutation and natural selection.
In 2007, Nowak and colleagues demonstrated that the transition of irregular verbs to regular verbs in English over time obeys a simple
inverse-square law, thus providing one of the first quantitative laws in the evolution of language
. This demonstration was featured on the front cover of ''
Nature
Nature is an inherent character or constitution, particularly of the Ecosphere (planetary), ecosphere or the universe as a whole. In this general sense nature refers to the Scientific law, laws, elements and phenomenon, phenomena of the physic ...
''.
In 2010, Nowak, with
E. O. Wilson, and
Corina Tarnita, in ''Nature'', argued that standard natural selection theory represents a simpler and superior approach to
kin selection
Kin selection is a process whereby natural selection favours a trait due to its positive effects on the reproductive success of an organism's relatives, even when at a cost to the organism's own survival and reproduction. Kin selection can lead ...
theory in the evolution of
eusociality
Eusociality ( Greek 'good' and social) is the highest level of organization of sociality. It is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations wit ...
. This work has led to many comments including strong criticism from proponents of
inclusive fitness theory. Nowak maintains that the findings of the paper are conclusive and that the field of social evolution should move beyond inclusive fitness theory.
He has over 300 scientific publications, of which 40 are in ''
Nature
Nature is an inherent character or constitution, particularly of the Ecosphere (planetary), ecosphere or the universe as a whole. In this general sense nature refers to the Scientific law, laws, elements and phenomenon, phenomena of the physic ...
'' and 15 in ''
Science
Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
''.
Nowak's research interests include:
* Somatic evolution of
cancer
Cancer is a group of diseases involving Cell growth#Disorders, abnormal cell growth with the potential to Invasion (cancer), invade or Metastasis, spread to other parts of the body. These contrast with benign tumors, which do not spread. Po ...
,
genetic instability,
tumor
A neoplasm () is a type of abnormal and excessive growth of tissue. The process that occurs to form or produce a neoplasm is called neoplasia. The growth of a neoplasm is uncoordinated with that of the normal surrounding tissue, and persists ...
suppressor genes
*
Stem cells,
tissue architecture
*
Virus
A virus is a submicroscopic infectious agent that replicates only inside the living Cell (biology), cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Viruses are ...
es,
infectious disease
An infection is the invasion of tissue (biology), tissues by pathogens, their multiplication, and the reaction of host (biology), host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmis ...
s,
immunology
Immunology is a branch of biology and medicine that covers the study of Immune system, immune systems in all Organism, organisms.
Immunology charts, measures, and contextualizes the Physiology, physiological functioning of the immune system in ...
* Dynamics of
prion
A prion () is a Proteinopathy, misfolded protein that induces misfolding in normal variants of the same protein, leading to cellular death. Prions are responsible for prion diseases, known as transmissible spongiform encephalopathy (TSEs), w ...
infections
*
Quasispecies
*
Genetic redundancy
*
Evolution of language
*
Evolutionary game theory
*
Evolutionary graph theory
*
Evolution of cooperation
* Prelife and
origins of life
Published books
Aside from his scientific career, Nowak has also authored five books. His 2006 work ''Evolutionary Dynamics: Exploring the Equation of Life'' received praise for its perspective on theoretical biology and won the
R.R. Hawkins Award. In 2011, he co-authored ''SuperCooperators'', Also authored by Roger Highfield which argues for cooperation as a fundamental principle of evolution and garnered positive reviews. Additionally, Nowak has edited books, including ''Evolution, Games, and God'', which examines the relationship between theology and evolutionary theory. Nowak identifies himself as a
Roman Catholic
The Catholic Church (), also known as the Roman Catholic Church, is the largest Christian church, with 1.27 to 1.41 billion baptized Catholics worldwide as of 2025. It is among the world's oldest and largest international institut ...
, advocating for the compatibility of science and religion in the pursuit of truth. His 2024 book, ''Beyond'', is a poetic exploration of the connection between religion and science. In 2015, he received the honorary degree Doctor of Humane Letters from the Dominican School of Philosophy & Theology at
Berkeley.
Nowak's first book ''Virus Dynamics: Mathematical Principles of Immunology and Virology'', written with Robert May, was published by
Oxford University Press
Oxford University Press (OUP) is the publishing house of the University of Oxford. It is the largest university press in the world. Its first book was printed in Oxford in 1478, with the Press officially granted the legal right to print books ...
in 2001. Nowak's 2006 book ''Evolutionary Dynamics: Exploring the Equations of Life'' discusses the evolution of various biological processes. Reviewing ''Evolutionary Dynamics'' in ''Nature'',
Sean Nee called it a "unique book" that "should be on the shelf of anyone who has, or thinks they might have, an interest in theoretical biology." The book received the
Association of American Publishers' R.R. Hawkins Award for the Outstanding Professional, Reference or Scholarly Work of 2006.
Nowak's book ''SuperCooperators: The Mathematics of Evolution, Altruism and Human Behaviour (Or, Why We Need Each Other to Succeed)'', co-authored with
Roger Highfield, was published in 2011. ''SuperCooperators'' is an autobiography of Nowak and a presentation of his work in mathematical biology on the evolution of cooperation, the origin of life, and the evolution of language. In the book, Nowak argues that cooperation is the third fundamental principle of evolution, next to
mutation
In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, ...
and
natural selection
Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
. ''SuperCooperators'' received positive reviews in ''
The New York Times
''The New York Times'' (''NYT'') is an American daily newspaper based in New York City. ''The New York Times'' covers domestic, national, and international news, and publishes opinion pieces, investigative reports, and reviews. As one of ...
'', ''Nature'', and the ''
Financial Times
The ''Financial Times'' (''FT'') is a British daily newspaper printed in broadsheet and also published digitally that focuses on business and economic Current affairs (news format), current affairs. Based in London, the paper is owned by a Jap ...
''.
With Sarah Coakley, Nowak edited the 2013 book ''Evolution, Games, and God: The Principle of Cooperation'', published by
Harvard University Press
Harvard University Press (HUP) is an academic publishing house established on January 13, 1913, as a division of Harvard University. It is a member of the Association of University Presses. Its director since 2017 is George Andreou.
The pres ...
. The volume features articles from experts in multiple fields who explore the interplay between theology and evolutionary theory as pertaining to cooperation and
altruism
Altruism is the concern for the well-being of others, independently of personal benefit or reciprocity.
The word ''altruism'' was popularised (and possibly coined) by the French philosopher Auguste Comte in French, as , for an antonym of egoi ...
.
Awards
Nowak is a corresponding member of the
Austrian Academy of Sciences. He won the
Weldon Memorial Prize, the Albert Wander Prize, the Akira Okubo Prize, the David Starr Jordan Prize
and the Henry Dale Prize.
Personal life
Nowak is a
Roman Catholic
The Catholic Church (), also known as the Roman Catholic Church, is the largest Christian church, with 1.27 to 1.41 billion baptized Catholics worldwide as of 2025. It is among the world's oldest and largest international institut ...
.
In a 2007 lecture at Harvard, he argued that science and religion occupied different but complementary roles in humans' search for meaning, stating: "Science and religion are two essential components in the search for truth. Denying either is a barren approach."
References
External links
*
Martin Nowak: Extended film interview with transcript for the 'Why Are We Here?' documentary series
{{DEFAULTSORT:Nowak, Martin
1965 births
Christian scholars
Living people
Fellows of Wolfson College, Oxford
Evolutionary biologists
University of Vienna alumni
Harvard University faculty
Austrian Roman Catholics
Theistic evolutionists
Austrian mathematicians
Austrian biochemists