''Linaria'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and

herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition ...

perennials A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

, and the largest genus in the

Antirrhineae The Antirrhineae are one of the 12 tribes of the family Plantaginaceae. It contains the toadflax relatives, such as snapdragons. Description and uses Most Antirrhineae are herbaceous, short-lived, perennial or annual plants growing at most ...

tribe of the plantain family

Plantaginaceae Plantaginaceae, the plantain family, is a large, diverse family of flowering plants in the order Lamiales that includes common flowers such as snapdragon and foxglove. It is unrelated to the banana-like fruit also called "plantain." In older ...

Taxonomy

''Linaria'' was traditionally placed in the family

Scrophulariaceae The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral ( zygomorphic) or rarely radial ( actinomorphic) symmetry. The S ...

Phylogenetic analysis In biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that pro ...

has now placed it in the vastly expanded family

. Closely related genera include ''

Nuttallanthus ''Nuttallanthus'' is a genus of four species of herbaceous annuals and perennials that was traditionally placed in the foxglove family Scrophulariaceae. Due to new genetic research, it has now been placed in the vastly expanded family Planta ...

'' (American toadflaxes, recently split from ''Linaria''), ''

Antirrhinum ''Antirrhinum'' is a genus of plants commonly known as dragon flowers, snapdragons and dog flower because of the flowers' fancied resemblance to the face of a dragon that opens and closes its mouth when laterally squeezed. They are native to ...

'' (snapdragons) and ''

Cymbalaria ''Cymbalaria'' is a genus of about 10 species of herbaceous perennial plants previously placed in the family Scrophulariaceae, but recently shown by genetic research to be in the much enlarged family Plantaginaceae. The genus is native to southe ...

'' (ivy-leaved toadflaxes).

Cultivation

Several ''Linaria'' species are cultivated as garden plants, and some are regarded as having a weedy habit. *Common toadflax or butter-and-eggs (''

Linaria vulgaris ''Linaria vulgaris'', the common toadflax,Blamey, M. & Grey-Wilson, C. (1989). ''Flora of Britain and Northern Europe''. yellow toadflax or butter-and-eggs, is a species of flowering plant in the family Plantaginaceae, native to Europe, Siberia ...

''), a European species which is widely introduced elsewhere and grows as a common weed in some areas. *Broomleaf toadflax or Dalmatian toadflax (''

Linaria genistifolia ''Linaria genistifolia'' is a species of flowering plant belonging to the family Plantaginaceae. Its native range is from Europe to southwestern Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is a ...

'', syn. ''L. dalmatica''), a native of southeast Europe that has become a weed in parts of North America.Dalmatian Toadflax (''Linaria dalmatica'').
National Invasive Species Information Center,

United States National Agricultural Library The United States National Agricultural Library (NAL) is one of the world's largest agricultural research libraries, and serves as a national library of the United States and as the library of the United States Department of Agriculture. Located ...

. *Purple toadflax ('' Linaria purpurea''), a species native to the

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on th ...

region grown as a

garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...

plant for its dark purple or pink flowers. The version with purple flowers can be mistaken for lavender. Spreads readily. *Pale toadflax ('' Linaria repens''), a species from western Europe similar to ''L. purpurea'', but with paler flowers. *Alpine toadflax (''

Linaria alpina ''Linaria alpina'', sometimes called alpine toadflax, is a purple-flowered plant native to mountainous areas of southern and central Europe. It belongs to the family Plantaginaceae (plantain family; unrelated to the fruit). It is found in many m ...

''), purple flowers with orange (or purple) lobes in the center. *Moroccan toadflax (''

Linaria maroccana ''Linaria maroccana'' is a species of flowering plant in the plantain family known by the common names Moroccan toadflax and annual toadflax. It is native to Morocco, but it can be found elsewhere growing wild as an introduced species, such as Ca ...

''), the flower has five lobes arranged into two lips with a spur at the end, often purple with white.

Species

The following species are recognised in the genus ''Linaria'': *''

Linaria accitensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria acutiloba ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria aeruginea ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria afghanica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria alaica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria albifrons ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria algarviana ''Linaria algarviana'' is a species of toadflax in the plantain family Plantaginaceae, endemic to southern Portugal, specifically the western portion of the Algarve The Algarve (, , ; from ) is the southernmost NUTS II region of continental P ...

'' *''

Linaria altaica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria amethystea ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria amoi ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

'' *''

Linaria angustissima ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria antilibanotica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria arcusangeli ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria arenaria ''Linaria arenaria'', sometimes called sand toadflax or French toadflax, is a yellow-flowered member of the genus ''Linaria''. It is found in sand-dunes and sandy grassland in western France, north-west Spain and Portugal and is thus a psammophyt ...

'' *''

Linaria arenicola ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria argillicola ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria armeniaca ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria arvensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *'' Linaria atlantica'' *''

Linaria azerbaijanensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria badachschanica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria badalii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria bamianica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria becerrae ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria bessarabica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria biebersteinii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria bipartita ''Linaria bipartita'' is a species of flowering plant in the plantain family known by the common name clovenlip toadflax. It is native to Morocco, but it can be found elsewhere as an introduced species and it is cultivated as an ornamental plant. ...

'' *''

Linaria bipunctata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria bordiana ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria boushehrensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria brachyphylla ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria bubanii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria bungei ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria buriatica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

'' *''

Linaria caesia ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria capraria ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria cavanillesii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria chalepensis ''Linaria chalepensis'' is a white-flowered plant found in southern Europe, belonging to the family Plantaginaceae (plantain family; unrelated to the fruit). Sources References {{Taxonbar, from=Q17735634 chalepensis Flora of Malta ...

'' *''

Linaria clementei ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria confertiflora ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria corifolia ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria × cornubiensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria cossoniana ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria cossonii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria cretacea ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria dalmatica ''Linaria dalmatica'' is a herbaceous, short-lived perennial plant native to western Asia and southeastern Europe that has become a weed in other areas. The family this plant now belongs to is the Plantaginaceae Family. Previously, it belonge ...

'' *''

Linaria damascena ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria decipiens ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria depauperata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria diffusa ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria dissita ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria × dominii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

'' *''

Linaria dumanii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria elegans ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria elymaitica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

'' *''

Linaria fallax ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria farsensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria fastigiata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria faucicola ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria fedorovii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria ficalhoana ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria flava ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria gharbensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria glacialis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria glauca ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria golestanensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria grandiflora ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria griffithii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria grjunerae ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria guilanensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria haelava ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria hepatica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

'' *''

Linaria heratensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria hirta ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria hohenackeri ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria huteri ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria × hybrida ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria iconia ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria ikonnikovii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria imzica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria incarnata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria incompleta ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria intricata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

'' *''

Linaria iranica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria × jalancina ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *'' Linaria japonica'' *''

Linaria jaxartica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

'' *''

Linaria joppensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria kavirensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''

Linaria khalkhalensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria khorasanensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria × kocianovichii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria kokanica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria kulabensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria kurdica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria latifolia ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria laxiflora ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

'' *''

Linaria leptoceras ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria lineolata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria loeselii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are Annual plant, annuals and herbaceous Perennial plant, perennials, and the largest genus in the Antirrhineae tribe ...

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Linaria longicalcarata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria macrophylla ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria macroura ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria maymanica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria mazandaranensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria melampyroides ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

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Linaria melanogramma ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria meyeri ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

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Linaria michauxii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria micrantha ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria microsepala ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria multicaulis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria munbyana ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria musilii ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria nachitschevanica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria nigricans ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria nivea ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria nurensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria nuristanica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria oblongifolia ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

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Linaria odora ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria oligantha ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria × oligotricha ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria onubensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plan ...

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Linaria orbensis ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria ordubadica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria pamirica ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

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Linaria paradoxa ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *'' Linaria parviracemosa'' *'' Linaria pedicellata'' *''

Linaria pedunculata ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Plantag ...

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Linaria pelisseriana ''Linaria'' is a genus of almost 200 species of flowering plants, one of several related groups commonly called toadflax. They are annuals and herbaceous perennials, and the largest genus in the Antirrhineae tribe of the plantain family Planta ...

'' *''Linaria peloponnesiaca'' *''Linaria peltieri'' *''Linaria pinifolia'' *''Linaria platycalyx'' *''Linaria polygalifolia'' *''Linaria popovii'' *''Linaria propinqua'' *''Linaria pseudolaxiflora'' *''Linaria pseudoviscosa'' *'' Linaria purpurea'' *''Linaria pyramidalis'' *''Linaria qartobensis'' *''Linaria quasisessilis'' *''Linaria reflexa'' *''Linaria remotiflora'' *'' Linaria repens'' *''Linaria ricardoi'' *''Linaria riffea'' *''Linaria × rocheri'' *''Linaria rubioides'' *''Linaria sabulosa'' *''Linaria salangensis'' *''Linaria salzmannii'' *''Linaria saposhnikovii'' *''Linaria saturejoides'' *''Linaria saxatilis'' *''Linaria schelkownikowii'' *''Linaria schirvanica'' *''Linaria semialata'' *''Linaria × sepium'' *''Linaria sessilis'' *''Linaria simplex'' *''Linaria spartea'' *''Linaria striatella'' *''Linaria supina'' *''Linaria tarhunensis'' *''Linaria tenuis'' *''Linaria thibetica'' *''Linaria thymifolia'' *''Linaria tingitana'' *''Linaria tonzigii'' *''Linaria triornithophora'' *''Linaria triphylla'' *''Linaria tristis'' *''Linaria tursica'' *''Linaria unaiensis'' *''Linaria × valdesiana'' *''Linaria venosa'' *''Linaria ventricosa'' *''Linaria veratrifolia'' *''Linaria verticillata'' *''Linaria virgata'' *''Linaria viscosa'' *''Linaria volgensis'' *''

'' *''Linaria warionis'' *''Linaria weilleri'' *''Linaria yunnanensis'' *''Linaria yusufeliensis'' *''Linaria × zaborskiana'' *''Linaria zaissanica''

Etymology

The members of this genus are known in English as toadflax, a name shared with several related genera. The 'toad' in toadflax may relate to the plants having historically been used to treat bubonic plague, a false link having been drawn between the words 'bubo' and 'Bufo'. The scientific name ''Linaria'' means "resembling ''linum''" (flax), which the foliage of some species superficially resembles.

Distribution and habitat

The genus is native to temperate regions of Europe, northern Africa and Asia, with the highest species diversity in the Mediterranean region.

Ecology

Some ''Linaria'' are regarded as noxious weeds. They are likely toxic to livestock, but ruminants generally avoid them.Sing, S. E. and R. K. Peterson. (2011)
Assessing environmental risks for established invasive weeds: Dalmatian (''Linaria dalmatica'') and yellow (''L. vulgaris'') toadflax in North America.
''International Journal of Environmental Research and Public Health'' 8(7) 2828-53.

Chemical composition

''Linaria'' species are rich in alkaloids, iridoids, terpenes, phenolic acids and flavonoids. Vasicine, Vasicinone, 7-hyrdoxyvasicine, Linarinic acid, Choline, Linavuline, Luteolin, Acacetin, Apigenin, Chrysin, Quercetin, Myricetin, Linarioside, Aucubin, Linaride, Iridolinaroside A-D, Iridolinarin A-C are some compounds found in plants of this genus.

Uses

Toadflaxes are used as food plants by the larvae of some Lepidoptera species, including the mouse moth (''Amphipyra tragopoginis'') and the Junonia coenia, common buckeye (''Junonia coenia'').

Traditional medicine

''Linaria vulgaris'' has been used as a herbalism, medicinal herb.Duke, J. A
Ethnobotanical uses: ''Linaria vulgaris''.
Dr. Duke's Phytochemical and Ethnobotanical Databases.

References

Bibliography

* A Phylogeny of Toadflaxes (''Linaria'' Mill.) Based on Nuclear Internal Transcribed Spacer Sequences: Systematic and Evolutionary Consequences. Mario Fernández-Mazuecos, José Luis Blanco-Pastor, and Pablo Vargas. ''International Journal of Plant Sciences'', Vol. 174, No. 2 (February 2013), pp. 234–249 Published by: The University of Chicago Press, Article DOI: 10.1086/668790 * Vargas P, JA Rosselló, R Oyama, J Güemes. 2004 Molecular evidence for naturalness of genera in the tribe Antirrhineae (Scrophulariaceae) and three independent evolutionary lineages from the New World and the Old. ''Plant Systematics and Evolution'' 249:151–172. * {{Taxonbar, from=Q156692 Linaria, Plantaginaceae genera Taxa named by Philip Miller