The Limacoidei is a
taxonomic infraorder
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
of air-breathing
land snails,
semislug
Semi-slugs, also spelled semislugs, are land gastropods whose shells are too small for them to retract into, but not quite vestigial. The shell of some semi-slugs may not be easily visible on casual inspection, because the shell may be covered ...
s and
slugs,
terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod molluscs
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...
in the suborder
Helicina
''Helicina'' is a genus of tropical and subtropical land snails with an operculum, terrestrial gastropod mollusks.
This is the type genus of its family ( Helicinidae) and subfamily ( Helicinae), as well as the superfamily Helicinoidea. Thi ...
Distribution
The original ancestral area of limacoid families is thought to be the
Palearctic
The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa.
The realm consists of several bioregions: the Euro-Si ...
region and south-eastern Asia.
Etymology
The word "limacoid" means "resembling a slug".
Typography of the name
In 1998, for the same taxon, Hausdorf
used the name Limacoidea
sensu lato
''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...
.
The name of this
taxon
In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
, the limacoid clade, was written by Bouchet & Rocroi (2005)
with
quotation mark
Quotation marks (also known as quotes, quote marks, speech marks, inverted commas, or talking marks) are punctuation marks used in pairs in various writing systems to set off direct speech, a quotation, or a phrase. The pair consists of an ...
s like this: "limacoid clade".
Other typographical variants are used by various other authors, for example capitalizing and restricting the use of the quote marks thus: "Limacoid" clade and "Limacoid clade".
2003 taxonomy by Schileyko
The study of A. Schileyko in this group, published in parts 8–11 of his monograph between 2002 and 2003, did not discuss the system that Hausdorf had proposed.
However, Schileyko refers to the work of Hausdorf, where this system was proposed,
in part 10 (2003) on p. 1390.
So he was acquainted with this system before 2003.
Alternative taxonomy is as follows (subfamilies listed only for Helicarionidae and Zonitidae):
*Superfamily Helicarionoidea
**Family Euconulidae
**Family Trochomorphidae
**Family Helicarionidae
***Subfamily Geotrochinae
***Subfamily Helicarioninae
***Subfamily Papuarioninae
***Subfamily Urocyclinae
**Family Gymnarionidae
**Family Rhysotinidae
**Family Ariophantidae
**Family Ostracolethidae
**Family Ryssotidae
**Family Milacidae
*Superfamily Dyakioidea
**Family Dyakiidae
**Family Staffordiidae
*Superfamily Gastrodontoidea
**Family Gastrodontidae
*Superfamily Zonitoidea
**Family Zonitidae
***Subfamily Pristilomatinae
***Subfamily Godwiniinae
***Subfamily Zonitinae
***Subfamily Oxychilinae
**Family Daudebardiidae
**Family Parmacellidae
*Superfamily Trigonochlamydoidea
**Family Trigonochlamydidae
*Superfamily Vitrinoidea
**Family Vitrinidae
*Superfamily Limacoidea
**Family Limacidae
**Family Agriolimacidae
**Family Boettgerillidae
Moreover, after these groups, in the same infraorder Limacoinei, Schileyko listed six more superfamilies: Camaenoidea, Xanthonychoidea, Helicoidea, Polygyroidea, Hygromioidea and Arionoidea.
2005 taxonomy by Bouchet & Rocroi
Explanatory note
For this particular group of families,
Bouchet & Rocroi adapted the taxonomic approach which was originally published by German malacologist Bernhard Hausdorf in 1998 in the ''Journal of Molluscan Studies''.
[ http://mollus.oxfordjournals.org/cgi/content/abstract/64/1/35] Bouchet & Rocroi clearly state, on p. 283:
However, in their paper, Bouchet & Rocroi (apparently accidentally) failed to show, graphically or typographically, ''any'' coverage of the limacoid clade in the taxonomic section of their paper, on pp. 268–269.
Because of this omission, some authors have subsequently (mistakenly) included in the "limacoid clade" the two following superfamilies:
Arionoidea
Arionoidea is a taxonomic group, superfamily of air-breathing land slugs, shell-less terrestrial pulmonate gastropod mollusks.
Families
Families within the superfamily Arionoidea include:
* Arionidae
* Anadenidae
* Ariolimacidae
* Binneyidae ...
and
Helicoidea
Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora.
Taxonomy 2005 taxonomy
There are 19 families within the superfamily Helicoidea according to the taxonom ...
. This misunderstanding was copied by Poppe & Tagaro (2006)
[Poppe G. T. & Tagaro S. (23 February 2006)]
''The New Classification of Gastropods According to Bouchet & Rocroi, 2005''
Visaya Net. who attempted to explain the changes within this taxonomy to the public. This same error is reproduced in various other internet sources.
Taxonomic list
The
taxonomy of Bouchet and Rocroi shows the "limacoid clade" as follows:
(Families that are exclusively fossil are indicated with a dagger †)
*Superfamily
Staffordioidea
**Family
Staffordiidae
*Superfamily
Dyakioidea
**Family
Dyakiidae
*Superfamily
Gastrodontoidea
Gastrodontoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the limacoid clade.
Taxonomy
According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), families in this superfamily ...
**Family
Gastrodontidae
Gastrodontidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Gastrodontoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). MolluscaBase eds. (2020). Mollusca ...
**Family
Chronidae
Chronidae is a family of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea within the superorder Eupulmonata. MolluscaBase eds. (2021). MolluscaBase. Chronidae Thiele, 1931. Accessed through ...
**Family
Euconulidae
**Family
Oxychilidae
Oxychilidae is a taxonomic family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Gastrodontoidea.
Distribution
The distribution of Oxychilidae includes the Nearctic, western-Palearctic, eastern-Pal ...
**Family
Pristilomatidae
Pristilomatidae is a taxonomic family of air-breathing land snails, terrestrial gastropod mollusks in the superfamily Gastrodontoidea.
According to the 2005 taxonomy of the Gastropoda by Bouchet & Rocroi these snails belong to the " limacoi ...
**Family
Trochomorphidae
**Fossil taxa probably belonging to the Gastrodontoidea
***Subfamily † Archaeozonitinae
***Subfamily † Grandipatulinae
***Subfamily † Palaeoxestininae
*Superfamily
Parmacelloidea
**Family
Parmacellidae
**Family
Milacidae
**Family
Trigonochlamydidae
Trigonochlamydidae is a family of air-breathing land slugs, terrestrial gastropod molluscs in the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).
Distribution
The native distribution of Trigonochlamy ...
*Superfamily
Zonitoidea
**Family
Zonitidae
*Superfamily
Helicarionoidea
Helicarionoidea is a superfamily of air-breathing land snails and semi-slugs, terrestrial pulmonate gastropod mollusks in the infraorder Limacoidei.
Families
There are three families within the superfamily Helicarionoidea:
* Helicarionidae
* ...
**Family
Helicarionidae
Helicarionidae is a family of air-breathing land snails or semi-slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea.
Distribution
The distribution of Helicarionidae includes the eastern Palearctic, Malagasy, ...
**Family
Ariophantidae
**Family
Urocyclidae
*Superfamily
Limacoidea
**Family
Limacidae
Limacidae, also known by their common name the keelback slugs, are a taxonomic family of medium-sized to very large, air-breathing land slugs, terrestrial pulmonate gastropod molluscs in the superfamily Limacoidea.
Distribution
The distributi ...
**Family
Agriolimacidae
**Family
Boettgerillidae
**Family
Vitrinidae
Vitrinidae is a family of small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Limacoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005). MolluscaBase eds. (2021). MolluscaBas ...
Cladogram
A
cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
showing phylogenic relations of families in the limacoid clade:
[Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". '']Journal of Biogeography
The ''Journal of Biogeography'' is a Peer review, peer-reviewed scientific journal in biogeography that was established in 1974. It covers aspects of spatial, ecological, and historical biogeography. The founding editor-in-chief was David Watts (b ...
'' 27(2): 379–390.
JSTOR
As explained in the previous section, the superfamilies
Arionoidea
Arionoidea is a taxonomic group, superfamily of air-breathing land slugs, shell-less terrestrial pulmonate gastropod mollusks.
Families
Families within the superfamily Arionoidea include:
* Arionidae
* Anadenidae
* Ariolimacidae
* Binneyidae ...
and
Helicoidea
Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the clade Stylommatophora.
Taxonomy 2005 taxonomy
There are 19 families within the superfamily Helicoidea according to the taxonom ...
do not belong to the limacoid clade. However, they are sometimes mistakenly included in this taxon.
References
* Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526
{{Taxonbar, from1=Q57262978, from2=Q11247992
Stylommatophora