''Ixinandria steinbachi'' is the only

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

in the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Ixinandria'' of

catfish Catfish (or catfishes; order (biology), order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Catfish are common name, named for their prominent barbel (anatomy), barbels, which resemble a cat's whiskers, though not ...

(

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Siluriformes) of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Loricariidae Loricariidae is the largest family (biology), family of catfish (order Siluriformes), with over 90 genus, genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South A ...

Taxonomy

''Ixinandria'' is part of the ''Rineloricaria'' group of the Loricariini tribe within the subfamily

Loricariinae Loricariinae is a subfamily of the family Loricariidae of catfish (order Siluriformes). This subfamily is divided into two tribes and about 30 genera. They are mainly native to freshwater habitats in South America, but there are also several spec ...

. The phylogenetic position of ''Ixinandria'' within the tribe Loricariini remains uncertain. It has been suggested that ''Ixinandria'' could be synonym of ''

Rineloricaria ''Rineloricaria'' (from the Greek, ''rhinos'' meaning ''nose'', and the Latin, ''lorica'' meaning ''cuirass of leather'') is a genus of freshwater tropical catfish (order Siluriformes) belonging to the family Loricariidae. They are commonly calle ...

''. Previously there were two species in this genus, but it was found that ''I. montebelloi'' was a

synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

of ''I. steinbachi''.

Distribution

The distribution of ''Ixinandria'' includes rivers of the

Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...

slope of the

Andes The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

and

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

. ''I. steinbachi'' occurs in the Salado River basin. ''I. steinbachi'' occurs in

Salta Province Salta () is a Provinces of Argentina, province of Argentina, located in the northwest of the country. Neighboring provinces are from the east clockwise Formosa Province, Formosa, Chaco Province, Chaco, Santiago del Estero Province, Santiago del Es ...

in mountainous areas at high altitudes ranging from around 1000–2900 metres (3300–9500 ft) above

sea level Mean sea level (MSL, often shortened to sea level) is an mean, average surface level of one or more among Earth's coastal Body of water, bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical ...

Appearance and anatomy

Sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

includes hypertrophied

odontode Odontodes, or dermal teeth, are hard structures found on the external surfaces of animals or near internal openings. They consist of a soft pulp surrounded by dentine and covered by a mineralized substance such as enamel, a structure similar to t ...

s around the head margin and on the

pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...

spines of mature males. These fish reach a length of SL.

Habitat ecology

''Ixinandria steinbachi'' is a

rheophilic A rheophile is an animal that prefers to live in fast-moving water. Examples of rheophilic animals Insects *Many aquatic insects living in riffles require current to survive. *'' Epeorus sylvicola'', a rheophilic mayfly species ( Ephemeroptera ...

species that lives in fast flowing and very oxygenated waters. Its color pattern reflects mimicry with stoned bottoms. Fertilized eggs have been found on the hidden surface of a stone, suggesting that ''I. steinbachi'' could be a cavity spawner.

References

Loricariini Catfish of South America Freshwater fish of Argentina Fish of Bolivia Monotypic freshwater fish genera Catfish genera Taxa named by Isaäc J. H. Isbrücker Taxa named by Han Nijssen Taxa named by Charles Tate Regan {{Loricariinae-stub