Grimmen Formation
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The Grimmen Formation is a Lower
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
(
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
)
geological formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics (lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock expo ...
in northeastern Germany, primarily exposed in the Grimmen and Klein Lehmhagen
clay pit A clay pit is a quarry or Mining, mine for the extraction of clay, which is generally used for manufacturing pottery, bricks or Portland cement. Quarries where clay is mined to make bricks are sometimes called brick pits. A brickyard or brickwor ...
s and documented in wells such as Reinberg 1E. Formally established in 2025, it was previously part of the informal “Green Series” of the
Ciechocinek Formation The Ciechocinek Formation (also known as the Gryfice Formation at Suliszewo, Choszczno County, Suliszewo) is a Jurassic (lower Toarcian) geological formation extending across the Baltic coast, primarily in Poland, with minor occurrences in Lithua ...
. It represents a prodelta to
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
- marine depositional system in the eastern
North German Basin North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north'' is ...
(NGB), shaped by sea-level changes and the Toarcian anoxic event.


History

Studies of
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
strata near Grimmen began in 1874 with the discovery of fossiliferous
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s in a railway cutting near Schönenwalde, 5 km north of Grimmen, initially misidentified as
Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
due to
ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
finds. In 1909, the succession was reclassified as Lower
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
(Lias epsilon), distinct from the
Posidonia Shale Formation The Posidonia Shale (, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Early to Late Toarcian) geological formation in Germany, northern Switzerland, northwestern Austria, souther ...
. From 1959 to 1995, the Grimmen
clay pit A clay pit is a quarry or Mining, mine for the extraction of clay, which is generally used for manufacturing pottery, bricks or Portland cement. Quarries where clay is mined to make bricks are sometimes called brick pits. A brickyard or brickwor ...
was excavated, revealing a glacially dislocated raft of Liassic
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s and
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
s deformed by
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
ice advances. Exploration wells (e.g., Reinberg 1E, Kb Barth 10, Kb Grambow 5) since the 1950s provided extensive core data, with Reinberg 1E designated as the reference section in 2025. The
clay pit A clay pit is a quarry or Mining, mine for the extraction of clay, which is generally used for manufacturing pottery, bricks or Portland cement. Quarries where clay is mined to make bricks are sometimes called brick pits. A brickyard or brickwor ...
, abandoned and water-filled since 1995, was established as the type section in 2025, with 2016 and 2020 excavations refining an 18 m composite log spanning the
Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 192.9 ±0.3 Ma and 184.2 ±0.3 Ma (million years ago). The Plie ...
-
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
boundary to the elegans Subzone.


Sedimentology/Lithology

The Grimmen Formation comprises greenish to bluish
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone and shale. Most of the particles of which the stone is composed are less than and are too small to ...
s,
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
s, and fine-grained
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
s, with
siderite Siderite is a mineral composed of iron(II) carbonate (FeCO3). Its name comes from the Ancient Greek word (), meaning "iron". A valuable iron ore, it consists of 48% iron and lacks sulfur and phosphorus. Zinc, magnesium, and manganese commonly ...
and carbonate
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s, reaching up to 100 m thick in the eastern
North German Basin North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north'' is ...
. The type section in the Grimmen
clay pit A clay pit is a quarry or Mining, mine for the extraction of clay, which is generally used for manufacturing pottery, bricks or Portland cement. Quarries where clay is mined to make bricks are sometimes called brick pits. A brickyard or brickwor ...
forms an 18 m composite log, transitioning from
Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 192.9 ±0.3 Ma and 184.2 ±0.3 Ma (million years ago). The Plie ...
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
y deposits of the
Wolgast Formation Wolgast () is a town in the district of Vorpommern-Greifswald, in Mecklenburg-Vorpommern, Germany. It is situated on the bank of the river (or strait) Peenestrom, vis-a-vis the island of Usedom on the Baltic coast that can be accessed by road and ...
to
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
organo-detrital
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s and heteroliths of the Lehmhagen Member (
Posidonia Shale Formation The Posidonia Shale (, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Early to Late Toarcian) geological formation in Germany, northern Switzerland, northwestern Austria, souther ...
), followed by the Reinberg Member of the Grimmen Formation. The Reinberg Member (~10 m thick) starts with bluish
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s, grading to greenish, pellet-laminated
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s with exaratum and lower elegans marker beds (marly
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
s with fecal pellets). Pellet laminae, up to 5 mm thick, decrease upsection, while silty/
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
y intercalations and
siderite Siderite is a mineral composed of iron(II) carbonate (FeCO3). Its name comes from the Ancient Greek word (), meaning "iron". A valuable iron ore, it consists of 48% iron and lacks sulfur and phosphorus. Zinc, magnesium, and manganese commonly ...
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s increase, forming a coarsening-upward trend with
pyrite The mineral pyrite ( ), or iron pyrite, also known as fool's gold, is an iron sulfide with the chemical formula Fe S2 (iron (II) disulfide). Pyrite is the most abundant sulfide mineral. Pyrite's metallic luster and pale brass-yellow hue ...
-filled burrows. The upper Grimmen Formation features greenish
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s with thin
silt Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension (chemistry), suspension with water. Silt usually ...
/
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...
laminae in 8–15 m symmetric cycles, capped by the
Glashütte Formation Glashütte [] is a town in Saxony, Germany, known as the birthplace of the German watchmaking industry and has a population of about 7,000. Historically, it was first mentioned in a document circa 1445. In January 2008, the former municipality R ...
’s
deltaic A river delta is a landform, archetypically triangular, created by the deposition of the sediments that are carried by the waters of a river, where the river merges with a body of slow-moving water or with a body of stagnant water. The creati ...
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
s. Heavy minerals (
zircon Zircon () is a mineral belonging to the group of nesosilicates and is a source of the metal zirconium. Its chemical name is zirconium(IV) silicate, and its corresponding chemical formula is Zr SiO4. An empirical formula showing some of th ...
,
rutile Rutile is an oxide mineral composed of titanium dioxide (TiO2), the most common natural form of TiO2. Rarer polymorphs of TiO2 are known, including anatase, akaogiite, and brookite. Rutile has one of the highest refractive indices at vis ...
,
tourmaline Tourmaline ( ) is a crystalline silicate mineral, silicate mineral group in which boron is chemical compound, compounded with chemical element, elements such as aluminium, iron, magnesium, sodium, lithium, or potassium. This gemstone comes in a ...
) and
smectite A smectite (; ; ) is a mineral mixture of various swelling sheet silicates (phyllosilicates), which have a three-layer 2:1 (TOT) structure and belong to the clay minerals. Smectites mainly consist of montmorillonite, but can often contain secon ...
-group
clay mineral Clay minerals are hydrous aluminium phyllosilicates (e.g. kaolin, Al2 Si2 O5( OH)4), sometimes with variable amounts of iron, magnesium, alkali metals, alkaline earths, and other cations found on or near some planetary surfaces. Clay minera ...
s indicate volcanogenic input, while
kaolinite Kaolinite ( ; also called kaolin) is a clay mineral, with the chemical composition Al2 Si2 O5( OH)4. It is a layered silicate mineral, with one tetrahedral sheet of silica () linked through oxygen atoms to one octahedral sheet of alumina () ...
and
chlorite The chlorite ion, or chlorine dioxide anion, is the halite (oxyanion), halite with the chemical formula of . A chlorite (compound) is a compound that contains this group, with chlorine in the oxidation state of +3. Chlorites are also known as s ...
are common. Glacially dislocated
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
greensand Greensand or green sand is a sand or sandstone which has a greenish color. This term is specifically applied to shallow marine sediment that contains noticeable quantities of rounded greenish grains. These grains are called ''glauconies'' and co ...
and
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
are locally intercalated. Organic-rich layers contain charcoal fragments, suggesting
wildfire A wildfire, forest fire, or a bushfire is an unplanned and uncontrolled fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identified as a ...
s. Toarcian material found in glacial Erratics in
Ahrensburg Ahrensburg () is a town in the district of Stormarn, Schleswig-Holstein, Germany. It is located northeast of Hamburg and is part of the Hamburg Metropolitan Region. Its population is around 31,000. ''Schloss Ahrensburg'', the town's symbol, is ...
and the Hagen Forest have been in controversy due to its dubious origin, being linked with the Rya Formation and
Sorthat Formation The Sorthat Formation is a geologic formation on Bornholm, Denmark, and the Rønne Graben, Baltic Sea, from the Latest Pliensbachian to Late Toarcian. It holds plant fossils and invertebrate traces, overlain by Fluvial processes, fluvial and Lacu ...
, as well this unit. They were originally considered or local or Baltic in derivation, but that changued with the recovery of erratic concretions in the Baltic sea cliffs near
Lübeck Lübeck (; or ; Latin: ), officially the Hanseatic League, Hanseatic City of Lübeck (), is a city in Northern Germany. With around 220,000 inhabitants, it is the second-largest city on the German Baltic Sea, Baltic coast and the second-larg ...
, being found as part of the Weichselian Glacial Maximum. Liassic–Cretaceous sediments in the assemblage are most probably associated with the tectonic Sorgenfrei-Tornquist Zone. The origin of this erratics from southwestern Baltic, Poland or Danish archipelago is unlikely, as those zones are dominated by Late Cretaceous–Paleocene strata, suggesting that this Toarcian assamblages should come from south/SW between STZ–TTZ and the German Baltic coast. The most clear hint link this deposits with the Grimmen Fm, as they're identical in fauna and facies composition of Grimmen and Dobbertin, also affected by subglacial erosion and thrusting, suggesting a close stratigraphic and palaeogeographical origin.


Paleoenvironment

The Grimmen Formation was deposited in a prodelta to
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
- marine environment in the eastern
North German Basin North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''north'' is ...
, influenced by sea-level changes and the Toarcian anoxic event. The Reinberg Member reflects a transition from organo-detrital
Posidonia Shale Formation The Posidonia Shale (, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Early to Late Toarcian) geological formation in Germany, northern Switzerland, northwestern Austria, souther ...
to
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
- marine
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s, with high
total organic carbon Total organic carbon (TOC) is an analytical parameter representing the concentration of organic carbon in a sample. TOC determinations are made in a variety of application areas. For example, TOC may be used as a non-specific indicator of wa ...
(TOC, ~3 wt%) at the base decreasing to <1 wt% upsection, indicating a shift to dysoxic conditions. Basal bluish
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s (exaratum Subzone) represent a restricted marine basin, followed by greenish
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s with pellet laminae, suggesting increased
fluvial A river is a natural stream of fresh water that flows on land or inside caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of its course if it ru ...
input and suspension-load plumes during wetter phases. The coarsening-upward trend and onset of
bioturbation Bioturbation is defined as the reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains. Bioturbating activities have a profound effect on the environment and are thought to be a ...
(
pyrite The mineral pyrite ( ), or iron pyrite, also known as fool's gold, is an iron sulfide with the chemical formula Fe S2 (iron (II) disulfide). Pyrite is the most abundant sulfide mineral. Pyrite's metallic luster and pale brass-yellow hue ...
-filled burrows) reflect prodelta progradation from
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and ; ), or the Fennoscandian Peninsula, is a peninsula in Europe which includes the Scandinavian Peninsula, Scandinavian and Kola Peninsula, Kola peninsulas, mainland ...
, culminating in the
Glashütte Formation Glashütte [] is a town in Saxony, Germany, known as the birthplace of the German watchmaking industry and has a population of about 7,000. Historically, it was first mentioned in a document circa 1445. In January 2008, the former municipality R ...
’s
deltaic A river delta is a landform, archetypically triangular, created by the deposition of the sediments that are carried by the waters of a river, where the river merges with a body of slow-moving water or with a body of stagnant water. The creati ...
systems. High
kaolinite Kaolinite ( ; also called kaolin) is a clay mineral, with the chemical composition Al2 Si2 O5( OH)4. It is a layered silicate mineral, with one tetrahedral sheet of silica () linked through oxygen atoms to one octahedral sheet of alumina () ...
and
smectite A smectite (; ; ) is a mineral mixture of various swelling sheet silicates (phyllosilicates), which have a three-layer 2:1 (TOT) structure and belong to the clay minerals. Smectites mainly consist of montmorillonite, but can often contain secon ...
content indicate a warm, humid
climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...
with biochemical
weathering Weathering is the deterioration of rocks, soils and minerals (as well as wood and artificial materials) through contact with water, atmospheric gases, sunlight, and biological organisms. It occurs '' in situ'' (on-site, with little or no move ...
and volcanogenic influence, akin to modern
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
settings. The absence of major ripples suggests minimal
wave In physics, mathematics, engineering, and related fields, a wave is a propagating dynamic disturbance (change from List of types of equilibrium, equilibrium) of one or more quantities. ''Periodic waves'' oscillate repeatedly about an equilibrium ...
action, with sedimentation driven by
fluvial A river is a natural stream of fresh water that flows on land or inside caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of its course if it ru ...
input and
storm A storm is any disturbed state of the natural environment or the atmosphere of an astronomical body. It may be marked by significant disruptions to normal conditions such as strong wind, tornadoes, hail, thunder and lightning (a thunderstor ...
events. The formation extends from western
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
to the Ringkøbing-Fyn-Møn-Arkona High, interfingering with the
Ciechocinek Formation The Ciechocinek Formation (also known as the Gryfice Formation at Suliszewo, Choszczno County, Suliszewo) is a Jurassic (lower Toarcian) geological formation extending across the Baltic coast, primarily in Poland, with minor occurrences in Lithua ...
in the east. The Grimmen Formation’s
biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. "Biostratigraphy." ''Oxford Reference: Dictionary of Biology ...
is primarily based on
ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
zones, with the falciferum Zone (elegantulum, exaratum, elegans, falciferum subzones) for the lower
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
. The type section spans the
Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 192.9 ±0.3 Ma and 184.2 ±0.3 Ma (million years ago). The Plie ...
-
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
boundary (tenuicostatum Subzone) to the elegans Subzone, with key fossils preserved in
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s and
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
s.
Ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
s define subzones, with ''
Dactylioceras ''Dactylioceras'' was a widespread genus of ammonites from the Lower Jurassic period, approximately 180 million years ago ( mya).Like many other ammonites, the genus ''Dactylioceras'' is extremely important in biostratigraphy, being a key index ...
semicelatum'' and possible ''D. tenuicostatum'' at the boundary, followed by '' Lobolytoceras siemensi'' and '' Tiltoniceras antiquum'' in the Lehmhagen Member (semicelatum Subzone). Elegantulum
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s yield abundant '' Eleganticeras elegantulum'' and rare '' Hildaites murleyi'', while exaratum
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s in the Reinberg Member contain '' Cleviceras exaratum'', '' Phylloceras heterophyllum'', and ''
Lytoceras ''Lytoceras'' is an ammonite genus that was extant during most of the Jurassic and Cretaceous periods, and is the type genus for the family Lytoceratidae. These cephalopods were fast-moving nektonic carnivores. Description Shells of ''Lytoceras ...
crenatum''.
Belemnite Belemnitida (or belemnites) is an extinct order (biology), order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous (And possibly the Eocene). Unlike squid, belemnites had an internal skeleton that made up the cone ...
s ('' Passaloteuthis bisulcata'') are common in the semicelatum Subzone but absent from the elegantulum Subzone due to anoxia. Teuthoids and
coleoid Coleoidea or Dibranchiata is one of the two subclasses of cephalopod molluscs containing all the various taxa popularly thought of as "soft-bodied" or "shell-less" (i.e. octopus, squid and cuttlefish). Unlike its extant sister group Nautiloide ...
s occur in elegantulum and exaratum
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s.
Holoplankton Holoplankton are organisms that are planktic (they live in the water column and cannot swim against a current) for their entire life cycle. Holoplankton can be contrasted with meroplankton, which are planktic organisms that spend part of their l ...
ic
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
''
Coelodiscus ''Ceolodiscus'' is an extinct genus of gastropod from the Lower and Middle Jurassic of Europe, mostly on Germany, France and United Kingdom. Other posible records include specimens from Hungary of Earliest Jurassic (Hettangian) age. As well there ...
minutus'' and larval ''‘
Inoceramus ''Inoceramus'' (Greek: translation "fibrous shell" for the fibrous structure of the mineral crystals in the shell) is an extinct genus of fossil marine pteriomorphian bivalves that superficially resembled the related winged pearly oysters of th ...
’ dubius'' are abundant, with adults in elegantulum and exaratum
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s, often in
regurgitalite Regurgitalites, or sometimes Regurgitaliths,
are the
Ostracod Ostracods, or ostracodes, are a Class (biology), class of the crustacean, Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 33,000 species (only 13,000 of which are extant taxon, extant) have been identified,Brandão, S.N.; Antoni ...
s and
foraminifera Foraminifera ( ; Latin for "hole bearers"; informally called "forams") are unicellular organism, single-celled organisms, members of a phylum or class (biology), class of Rhizarian protists characterized by streaming granular Ectoplasm (cell bio ...
are absent in
black shale Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g., kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especiall ...
s but occur above the Dörnten Member. Calcareous nannoplankton (e.g., ''Rhombolithion'') are diverse in the elegantulum Subzone, correlating with NJ5b–NJ6 zones.
Palynofacies Palynology is the study of microorganisms and microscopic fragments of mega-organisms that are composed of acid-resistant organic material and occur in sediments, sedimentary rocks, and even some metasedimentary rocks. Palynomorphs are the mic ...
show a dinoflagellate cyst blackout in the elegantulum-exaratum Subzones, with amorphous organic matter (AOM) dominance and micro-charcoal.
Megaspore Megaspores, also called macrospores, are a type of spore that is present in heterosporous plants. These plants have two spore types, megaspores and microspores. Generally speaking, the megaspore, or large spore, germinates into a female gametoph ...
s (e.g., '' Paxillitriletes phyllicus'') from
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and ; ), or the Fennoscandian Peninsula, is a peninsula in Europe which includes the Scandinavian Peninsula, Scandinavian and Kola Peninsula, Kola peninsulas, mainland ...
indicate a humid
climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...
.


Reinberg Member Fossil Record

The Reinberg Member, notably at Grimmen and Dobbertin, is a Konservatlagerstätte with exceptional marine and
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
fossils in exaratum
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s.
Ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
s (e.g., ''Cleviceras exaratum''), ''Coelodiscus minutus'', and ''‘Inoceramus’ dubius'' are preserved in
calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...
, with predation traces.
Decapod The Decapoda or decapods, from Ancient Greek δεκάς (''dekás''), meaning "ten", and πούς (''poús''), meaning "foot", is a large order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfish, shrimp, and p ...
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s (e.g., ''Palaeoastacus''-like),
coleoid Coleoidea or Dibranchiata is one of the two subclasses of cephalopod molluscs containing all the various taxa popularly thought of as "soft-bodied" or "shell-less" (i.e. octopus, squid and cuttlefish). Unlike its extant sister group Nautiloide ...
gladii, and
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
(leptolepids, ''Grimmenodon aureum'') occur in
calcium phosphate The term calcium phosphate refers to a family of materials and minerals containing calcium ions (Ca2+) together with inorganic phosphate anions. Some so-called calcium phosphates contain oxide and hydroxide as well. Calcium phosphates are white ...
.
Coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name ...
s, likely from
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
,
shark Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the ...
s, and
marine reptile Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. Only about 100 of the 12,000 extant reptile species and subspecies are classed as marine reptiles, including mari ...
s (e.g.,
ichthyosaur Ichthyosauria is an order of large extinct marine reptiles sometimes referred to as "ichthyosaurs", although the term is also used for wider clades in which the order resides. Ichthyosaurians thrived during much of the Mesozoic era; based on fo ...
s,
plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
s), are common.
Terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, especially in exaratum
concretion A concretion is a hard and compact mass formed by the precipitation of mineral cement within the spaces between particles, and is found in sedimentary rock or soil. Concretions are often ovoid or spherical in shape, although irregular shapes a ...
s types 1 and 2, include well-preserved wings and rare 3D specimens, comparable to Dobbertin’s fauna. These fossils, formed in
anoxic Anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts: * Anoxic waters, sea water, fresh water or groundwater that are depleted of dissolved ox ...
, fine-grained
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
below the storm wave base, reflect a nutrient-rich environment fertilized by
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
carcasses.


Biota


Foraminifera


Ichnofossils


Brachiopoda


Bivalvia


Gastropoda


Cephalopoda


Crustacea

Small indeterminate shrimps, sometimes found associated in great numbers, are recovered on several layers at Grimmen.


Arachnida


Insecta

Insects are common terrestrial animals that were probably drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation. In Klein Lehmhagen insects are found as part of calcareous nodules in the exaratum-elegantulum subzones, with specimens also found in living chambers of '' Eleganticeras elegantulum'' macrochonchs and in fish coprolites which are the most frequent fossils at all. In the elegantulum the insect fauna is dominated by beetle elytra, indicating strong fluvial input and a nearshore deltaic complex. On Dobbertin, insects are present in the exaratum nodules, where fluvial input is seen thanks to the phyllopod abundance and whole bedding planes covered by algae substituted by Ca-phosphat, being the layers where insects are most abundant.


Notoptera


Odonatoptera


=Odonata

=


Paraneoptera


Eoblattida


Thysanoptera


Plecoptera


Orthoptera


Phasmatodea


Blattodea


Hemiptera


Hymenoptera


Megaloptera


Neuroptera


Coleoptera


Trichoptera


Lepidoptera


Mecoptera


Diptera


Echinodermata

In Dobbertin, the echinoderm remains are rare in contrast to foraminifera, phyllopods and ostracods, yet in some places they attain a percentage of the total fauna between 0.7 and 26.5%. In the upper layers they're totally absent, as well on the erratics and in the whole Grimmen sequence.


Vertebrates


Fishes


Amniotes


Flora


See Also

*
List of fossiliferous stratigraphic units in Germany See also * Lists of fossiliferous stratigraphic units in Europe * Geology of Germany References * {{DEFAULTSORT:Fossiliferous stratigraphic units in Germany Germany Germany geology-related lists ...
*
Toarcian turnover The Toarcian extinction event, also called the Pliensbachian-Toarcian extinction event, the Early Toarcian mass extinction, the Early Toarcian palaeoenvironmental crisis, or the Jenkyns Event, was an extinction event that occurred during the early ...
*
Posidonia Shale Formation The Posidonia Shale (, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Early to Late Toarcian) geological formation in Germany, northern Switzerland, northwestern Austria, souther ...


References

{{reflist, colwidth=30em Jurassic Germany Toarcian Stage Mudstone formations Deltaic deposits Fossiliferous stratigraphic units of Europe Paleontology in Germany