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Coleoid
Subclass Coleoidea, or Dibranchiata, is the grouping of cephalopods containing all the various taxa popularly thought of as "soft-bodied" or "shell-less" (i.e., octopuses, squid and cuttlefish). Unlike its extant sister group, Nautiloidea, whose members have a rigid outer shell for protection, the coleoids have at most an internal cuttlebone, gladius, or shell that is used for buoyancy or support. Some species have lost their cuttlebone altogether, while in some it has been replaced by a chitinous support structure. A unique trait of the group is the ability to edit their own RNA. The major divisions of Coleoidea are based upon the number of arms or tentacles and their structure. The extinct and most primitive form, the Belemnoidea, presumably had ten equally-sized arms in five pairs numbered dorsal to ventral as I, II, III, IV and V. More modern species either modified or lost a pair of arms. The superorder Decapodiformes has arm pair IV modified into long tentacles with sucker ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda ( Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles ( muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by '' Nautilus'' and '' Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been i ...
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Nautiloidea
Nautiloids are a group of marine cephalopods ( Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and '' Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant re ...
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Belemnitida
Belemnitida (or the belemnite) is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most to the tip: the tongue-shaped pro-ostracum, the conical phragmocone, and the pointy guard. The calcitic guard is the most common belemnite remain. Belemnites, in life, are thought to have had 10 hooked arms and a pair of fins on the guard. The chitinous hooks were usually no bigger than , though a belemnite could have had between 100 and 800 hooks in total, using them to stab and hold onto prey. Belemnites were an important food source for many Mesozoic marine creatures, both the adults and the planktonic juveniles, and likely played an important role in restructuring marine ecosystems after the Triassic–Jurassic extinction event. They may have laid between 100 and 1,000 eggs. Some species may have been adapted to speed and swam in the tu ...
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Neocoleoidea
Neocoleoidea is a large group of marine cephalopods. This cohort contains two extant groups: Decapodiformes (squid, cuttlefish, and relatives) and Octopodiformes (octopuses and the vampire squid). Species within this group exist in all major habitats in the ocean, in both the southern and northern polar regions, and from intertidal zones to great depths. Whilst conventionally held to be monophyletic, the only morphological character for the group is the presence of suckers: although the presence of these features in the belemnites suggests that they do not support the Neocoleoidea, and hence that the group may be paraphyletic. Taxonomy *Superorder Decapodiformes **Order Spirulida: ram's horn squid **Order Sepiida: cuttlefish **Order Sepiolida: bobtail squid **Order Teuthida: squid *Superorder Octopodiformes **Family † Trachyteuthididae (''incertae sedis'') **Order Vampyromorphida Vampyromorphida is an order of cephalopods comprising one known extant species ('' Vampyrote ...
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Boletzkyida
Boletzkyida is a primitive order of teuthid coleoid cephalopod: the boletzkyids are thought to be the earliest forms of coleoid cephalopods, and appear to form a link between nautiloid orthocerids and more advanced coleoids. Boletzkyida was named and described by Bandel, Reitner, and Sturmer in 1983 (B.R.&S) from specimens found in the Lower Devonian black slate (" Hunsrück Slate") in Germany. Taxonomy Bandel, ''et al'' (1983) divided the Boletzkyida into two, not too dissimilar families, each based on a single genus. They are the Boletzkyidae based on the genus ''Boletzkya'' and the Naefiteuthidae based on the genus ''Naefiteuthis''. The main difference between ''Boletzkya'' and ''Naefiteuthis'' is the length of the phragmocone at which time the living chamber becomes gladius-like and dominates the shell. Ontogeny Boletzkyd shells grew to as long as 16 cm. The juvenile portion is a phragmocone that closely resembles that of Silurian orthocerid ''Michelinoceras'' wit ...
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Hematitida
Hematitida is a group of coleoid cephalopods known from the early Carboniferous Period. They are the oldest definite coleoids, although there are controversial claims for even older coleoids from the Devonian. Fossil hematitidans have so far been found only in Arkansas and Utah of the United States. The only family described so far is Hematitidae. Characteristics Some features shared by hematitidans include * a short rostrum, made of aragonite and organic material, and bearing ridges * a short living chamber, only 1.5 to 2 times the length of the last chamber of the phragmocone * a spherical protoconch * a narrow, straight phragmocone * a multi-layered conotheca - the outer wall of the phragmocone * a narrow, ventral siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ... ...
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Phragmoteuthida
Phragmoteuthida is an order of extinct coleoid cephalopods characterized by a fan-like teuthoid pro-ostracum attached to a belemnoid-like phragmocone. Diagnosis Jeletzky characterized phragmoteuthids as having a large tripartite, fanlike pro-ostracum forming the longest portion of the shell, attached to about three-quarters of the circumference of a comparatively small breviconic phragmocone with short camerae and superficially belemnitid-like siphuncle, an absent or much reduced rostrum at the apical part of the phragmocone, belemnite-like arm hooks, an ink sack, beaks resembling those of Recent teuthids, and a muscular mantle. Donovan (2006), gives a similar description for ''Phragmoteuthis'': Phragmocones as having an apical angle of between 20 and 30 degrees, and relatively few chambers compared with belemnoids; a multi-layered conotheca, thick-walled siphuncle, and a long, three-lobed pro-ostracum as in the Triassic species. Arms are short and bear pairs of slightly ...
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Decapodiformes
Decapodiformes is a superorder of Cephalopoda comprising all cephalopod species with ten limbs, specifically eight short arms and two long tentacles. It is hypothesized that the ancestral coleoid had five identical pairs of limbs, and that one branch of descendants evolved a modified arm pair IV to become the Decapodiformes, while another branch of descendants evolved and then eventually lost its arm pair II, becoming the Octopodiformes. The Decapodiformes include: *Order Spirulida: ram's horn squid *Order Sepiida: cuttlefish, pygmy, bobtail and bottletail squid *Order Myopsida: coastal squid *Order Oegopsida Oegopsida is one of the two orders of squid in the superorder Decapodiformes, in the class Cephalopoda. Together with the Myopsina, it was formerly considered to be a suborder of the order Teuthida, in which case it was known as Oegopsina. This ...: neritic squid References Further reading *abstract External links The Taxonomicon: Superorder Decapodiformes Co ...
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Cuttlefish
Cuttlefish or cuttles are marine molluscs of the order Sepiida. They belong to the class Cephalopoda which also includes squid, octopuses, and nautiluses. Cuttlefish have a unique internal shell, the cuttlebone, which is used for control of buoyancy. Cuttlefish have large, W-shaped pupils, eight arms, and two tentacles furnished with denticulated suckers, with which they secure their prey. They generally range in size from , with the largest species, the giant cuttlefish (''Sepia apama''), reaching in mantle length and over in mass. Cuttlefish eat small molluscs, crabs, shrimp, fish, octopus, worms, and other cuttlefish. Their predators include dolphins, sharks, fish, seals, seabirds, and other cuttlefish. The typical life expectancy of a cuttlefish is about 1–2 years. Studies are said to indicate cuttlefish to be among the most intelligent invertebrates.
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Palaeoteuthomorpha
Palaeoteuthomorpha is a superorder of primitive coleoid cephalopods, containing the Boletzkyida, characterized by an orthoceroid-like phragmocone early in ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ... and a teuthid living chamber later in ontogeny. This contrasts with the later Teuthida which have a reduced living chamber (gladius) throughout. References *Klaus Bandel, J Reitner, and W Sturmer, 1983. Coleoids from the Lower Devonian Black Slate (Hunstruck-Shiefer) of the Hunsruck (West Germany) N Jb Geol Paliont Abh 165(3) 397-414 Stuttgart, Apr 1983. Coleoidea Protostome superorders {{Cephalopod-stub ...
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Cephalopod Limb
All cephalopods possess flexible limbs extending from their heads and surrounding their cephalopod beak, beaks. These appendages, which function as muscular hydrostats, have been variously termed arms, legs or tentacles. Description In the scientific literature, a cephalopod ''arm'' is often treated as distinct from a ''Tentacle#Tentacles in invertebrates, tentacle'', though the terms are sometimes used interchangeably, often with the latter acting as an umbrella term for cephalopod limbs. Generally, arms have suckers along most of their length, as opposed to tentacles, which have suckers only near their ends.Young, R.E., M. Vecchione & K.M. Mangold 1999Cephalopoda Glossary Tree of Life web project. Barring a few exceptions, octopuses have eight arms and no tentacles, while squid and cuttlefish have eight arms (or two "legs" and six "arms") and two tentacles.Norman, M. 2000. ''Cephalopods: A World Guide''. ConchBooks, Hackenheim. p. 15. "There is some confusion around the ...
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Gladius (cephalopod)
The gladius (plural: ''gladii''), or pen, is a hard internal bodypart found in many cephalopods of the superorder Decapodiformes (particularly squids) and in a single extant member of the Octopodiformes, the vampire squid (''Vampyroteuthis infernalis''). It is so named for its superficial resemblance to the Roman short sword of the same name, and is a vestige of the ancestral mollusc shell, which was external. The gladius is located dorsally within the mantle and usually extends for its entire length. Composed primarily of chitin, it lies within the shell sac, which is responsible for its secretion. Gladii are known from a number of extinct cephalopod groups, including teudopseids (''e.g.'' '' Actinosepia'', '' Glyphiteuthis'', '' Muensterella'', '' Palaeololigo'', '' Teudopsinia'', '' Teudopsis'', and '' Trachyteuthis''), loligosepiids (''e.g.'' '' Geopeltis'', '' Jeletzkyteuthis'', and '' Loligosepia''), and prototeuthids (''e.g.'' '' Dorateuthis'', '' Paraplesioteuthis'' ...
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