''Gongbusaurus'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
ornithischia
Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
n, perhaps
ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous worl ...
,
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived between about 160 and 157 million years ago, in the
Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the ...
period. A small
herbivore
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
, it is very poorly known. Two species have been assigned to it, but as the original name is based on teeth, there is no concrete evidence to connect the two species. Its fossils have been found in
China.
Description
''Gongbusaurus'', by extrapolation from the remains of possible species "G." ''wucaiwanensis'' and other
basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
ornithopods, was a herbivorous
bipedal
Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
animal around long. The
tibia
The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
of "G." ''wucaiwanensis'' is 19.5 centimetres long. It would have been a strong runner.
Classification
Dong Zhiming
Dong Zhiming ( Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at th ...
, Zhou Shiwu, and Zhang Zicheng, who originally described the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
''G. shiyii'', thought it was most similar to ''
Fabrosaurus'' and assigned it to the nebulous
Fabrosauridae.
Upon description of the second species "G." ''wucaiwanensis'' several years later, Dong elected to assign it to
Hypsilophodontidae, an equally nebulous (
paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
) family of somewhat more
derived small bipedal
ornithischia
Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
ns,
[ while at about the same time, ]David B. Weishampel
Professor David Bruce Weishampel (born November 16, 1952) is an American palaeontologist in the Center for Functional Anatomy and Evolution at Johns Hopkins University School of Medicine. Weishampel received his Ph.D. in Geology from the Univers ...
and Larry Witmer found ''Gongbusaurus'' to be an indeterminate basal ornithischian. The most recent reviews also found the genus to be a dubious ornithischian, and recommended renaming the better-known second species. Peter Galton has noted that the teeth on which ''Gongbusaurus'' is based resemble those of ''Sarcolestes
''Sarcolestes'' (meaning "flesh robber") is an extinct genus of ankylosaurian ornithischian dinosaur from the Oxford Clay of England. The current type and only species is ''S. leedsi'', and the holotype is a single partial left mandible. The ...
'' and '' Gastonia'', so the genus may actually be an ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs ...
n.
History
Dong and his coauthors established ''Gongbusaurus'' on two small teeth, holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
IVPP V9069: one, V9069-1, from the beak (premaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...
), and the other, V9069-2, from the cheek (maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
). These remains came from the Oxfordian-age Upper Shaximiao Formation in Sichuan
Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of th ...
, China and were found by a unit of the Chinese aerial survey behind an elementary school in the village of Huangtong, part of the Duxin commune, in Rongxian or Rong County. The type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
''Gongbusaurus shiyii'' was named and described by Dong, Zhou, and Zhang in 1983. The generic name is derived from the imperial ministry, ''bu'', of public works, ''gong'', in honour of the great poet Du Fu who in Sichuan worked for that ministry after 760, in the rank of ''shi yi'', "junior consultant", which explains the specific name. As ''shi yi'' also can mean "register of lost objects" the specific name is at the same time a pun on the fact that the teeth were later separately found among the assorted bones collected during the dig.[ Dong added a second species, "G." ''wucaiwanensis'', in 1989 for a fragmentary skeleton (holotype IVPP 8302) including a partial lower jaw, three tail ]vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characterist ...
e, and a partial forelimb, and added another specimen (paratype
In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...
IVPP 8303) consisting of two hip vertebrae, eight tail vertebrae, and two complete hind limbs. A partial foot (IVPP 8304) and four separate dorsal vertebrae and a caudal vertebra were referred. These remains came from the roughly contemporaneous Shishugou Formation
The Shishugou Formation () is a geological formation in Xinjiang, China.
Its strata date back to the Late Jurassic period. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Di ...
of Wucaiwan, Xinjiang
Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwest ...
.[
Tooth species are not well regarded in dinosaur paleontology, because dinosaur teeth are generally not distinctive enough to hold a name. Therefore, it is unsurprising that other paleontologists have suggested removing "G." ''wucaiwanensis''.][ A possible replacement name, " Eugongbusaurus",] has accidentally gotten to the public, but remains informal
Formal, formality, informal or informality imply the complying with, or not complying with, some set of requirements (forms, in Ancient Greek). They may refer to:
Dress code and events
* Formal wear, attire for formal events
* Semi-formal atti ...
.
References
{{Taxonbar, from=Q1656761
Ornithischian genera
Late Jurassic dinosaurs of Asia
Paleontology in Sichuan
Nomina dubia
Taxa named by Dong Zhiming
Fossil taxa described in 1983