Pilot whales are

cetacean Cetacea (; , ) is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively c ...

s belonging to the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Globicephala''. The two

extant Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Exta ...

species are the

long-finned pilot whale The long-finned pilot whale, or pothead whale (''Globicephala melas'') is a large species of oceanic dolphin. It shares the genus ''Pilot whale, Globicephala'' with the short-finned pilot whale (''Globicephala macrorhynchus''). Long-finned pilo ...

(''G. melas'') and the

short-finned pilot whale The short-finned pilot whale (''Globicephala macrorhynchus'') is one of the two species of cetaceans in the genus ''Pilot whale, Globicephala'', which it shares with the long-finned pilot whale (''G. melas''). It is part of the oceanic dolphin ...

(''G. macrorhynchus''). The two are not readily distinguishable at sea, and analysis of the skulls is the best way to distinguish between the species. Between the two species, they range nearly worldwide, with long-finned pilot whales living in colder waters and short-finned pilot whales living in tropical and subtropical waters. Pilot whales are among the largest of the

oceanic dolphin Oceanic dolphins or Delphinidae are a widely distributed family of dolphins that live in the sea. Close to forty extant species are recognised. They include several big species whose common names contain "whale" rather than "dolphin", such as the ...

s, exceeded in size only by the

orca The orca (''Orcinus orca''), or killer whale, is a toothed whale and the largest member of the oceanic dolphin family. The only extant species in the genus '' Orcinus'', it is recognizable by its black-and-white-patterned body. A cosmopol ...

. They and other large members of the dolphin family are also known as blackfish. Pilot whales feed primarily on squid, but will also hunt large

demersal fish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...

such as

cod Cod (: cod) is the common name for the demersal fish genus ''Gadus'', belonging to the family (biology), family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gad ...

and

turbot The turbot ( ) ''Scophthalmus maximus'' is a relatively large species of flatfish in the family Scophthalmidae. It is a demersal fish native to marine or brackish waters of the Northeast Atlantic, Baltic Sea and the Mediterranean Sea. It is a ...

. They are highly social and may remain with their birth pod throughout their lifetime. Short-finned pilot whales are one of the few non-primate mammal species in which females go through

menopause Menopause, also known as the climacteric, is the time when Menstruation, menstrual periods permanently stop, marking the end of the Human reproduction, reproductive stage for the female human. It typically occurs between the ages of 45 and 5 ...

, and postreproductive females continue to contribute to their pod. Pilot whales are notorious for stranding themselves on beaches, but the reason behind this is not fully understood. Marine biologists have shed some light on the matter, suggesting that it is due to the mammals inner ear (their principal navigational sonar) being damaged from noise pollution in the ocean, such as from cargo ships or military exercises. The

conservation status The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservation ...

of short-finned and long-finned pilot whales has been determined to be

least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...

Naming

The animals were named "pilot whales" because pods were believed to be "piloted" by a leader. They are also called "pothead whales" and "blackfish". The genus name is a combination of the Latin word ''globus'' ("round ball" or "globe") and the Greek word ''Kephale'' ("head").

Taxonomy and evolution

Pilot whales are classified into two species; the

(''Globicephala melas'') and the

(''G. macrorhynchus''). The short-finned pilot whale was described, from skeletal materials only, by

John Edward Gray John Edward Gray (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for a z ...

in 1846. He presumed from the skeleton that the whale had a large beak. The long-finned pilot whale was first classified by

Thomas Stewart Traill Thomas Stewart Traill (29 October 1781 – 30 July 1862) was a British physician, chemist, meteorologist, zoologist and scholar of medical jurisprudence. He was the grandfather of the physicist, meteorologist and geologist Robert Traill Omon ...

in 1809 as ''Delphinus melas''. Its scientific name was eventually changed to ''Globicephala melaena''. Since 1986, the specific name of the long-finned pilot whale was changed to its original form ''melas''. Other species classifications have been proposed but only two have been accepted. There exist geographic forms of short-finned pilot whales off the east coast of Japan, which comprise genetically isolated stocks. Fossils of an extinct relative, ''Globicephala baereckeii'', have been found in

Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...

deposits in Florida. Another ''Globicephala'' dolphin was discovered in

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58G. etruriae''. Evolution of ''Tappanaga'', the endemic, larger form of short-finned pilots found in northern Japan, with similar characteristics to the whales found along

Vancouver Island Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian province of British Columbia. The island is in length, in width at its widest point, and in total area, while are of land. The island is the largest ...

and northern USA coasts, has indicated that the geniture of this form could be caused by the extinction of long-finned pilots in north Pacific in the 12th century, where ''Magondou'', the smaller, southern type possibly filled the former niches of long-finned pilots, adapting and colonizing into colder waters.

Description

Pilot whales are mostly dark grey, brown, or black, but have some light areas such as a grey saddle patch behind the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

. Other light areas are an anchor-shaped patch under the chin, a faint blaze marking behind the eye, a large marking on the belly, and a genital patch. The dorsal fin is set forward on the back and sweeps backwards. A pilot whale is more robust than most dolphins and has a distinctive large, bulbous

melon A melon is any of various plants of the family Cucurbitaceae with sweet, edible, and fleshy fruit. It can also specifically refer to ''Cucumis melo'', commonly known as the "true melon" or simply "melon". The term "melon" can apply to both the p ...

. Pilot whales' long, sickle-shaped flippers and tail stocks are flattened from side to side. Male long-finned pilot whales develop more circular melons than females, although this does not seem to be the case for short-finned pilot whales off the Pacific coast of Japan. Pilot whale spyhop

Long-finned and short-finned pilot whales are so similar, it is difficult to tell the two species apart. They were traditionally differentiated by the length of the pectoral flippers relative to total body length and the number of teeth. The long-finned pilot whale was thought to have 9–12 teeth in each row and flippers one-fifth of total body length, compared to the short-finned pilot whale with its 7–9 teeth in each row and flippers one-sixth of total body length. Studies of whales in the Atlantic showed much overlap in these characteristics between the species, making them

cline Cline may refer to: Science * Cline (biology), a measurable gradient in a single trait in a species across its geographical range * Cline (hydrology), a fluid layer with a property that varies * Cline (mathematics) or generalised circle, a ci ...

s instead of distinctive features. Thus, biologists have since used skull differences to distinguish the two species. The size and weight depend on the species, as long-finned pilot whales are generally larger than short-finned pilot whales. Their lifespans are about 45 years in males and 60 years in females for both species. Both species exhibit

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

. Adult long-finned pilot whales reach a body length of approximately 6.5 m, with males being 1 m longer than females. Their body mass reaches up to 1,300 kg in females and up to 2,300 kg in males.Jefferson, T. A, Webber, M. A., Pitman, R. L., (2008)
Marine mammals of the world
'. Elsevier, Amsterdam. For short-finned pilot whales, adult females reach a body length of about 5.5 m, while males reach 7.2 m and may weigh up to 3,200 kg.

Distribution and habitat

Pilot whales can be found in oceans nearly worldwide, but data about current population sizes is deficient. The long-finned pilot whale prefers slightly cooler waters than the short-finned and is divided into two populations. The smaller group is found in a circumpolar band in the

Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the world ocean, generally taken to be south of 60th parallel south, 60° S latitude and encircling Antarctica. With a size of , it is the seco ...

from about 20 to 65°S. It may be sighted off the coasts of

Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...

, and

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

. An estimated more than 200,000 individuals were in this population in 2006. The second, much larger, population inhabits the North

Atlantic Ocean The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...

, in a band from

South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...

in the United States across to the

Azores The Azores ( , , ; , ), officially the Autonomous Region of the Azores (), is one of the two autonomous regions of Portugal (along with Madeira). It is an archipelago composed of nine volcanic islands in the Macaronesia region of the North Atl ...

and

Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...

at its southern edge and from

Newfoundland Newfoundland and Labrador is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region of Labrador, having a total size of . As of 2025 the population ...

Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...

, Iceland, and northern

Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of the Kingdom of ...

at its northern limit. This population was estimated at 778,000 individuals in 1989. It is also present in the western half of the

Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...

. The short-finned pilot whale is less populous. It is found in temperate and tropical waters of the Indian, Atlantic and

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the cont ...

Oceans. Its population overlaps slightly with the long-finned pilot whale in the temperate waters of the North Atlantic and Southern Oceans.Olson, P.A. (2008) "Pilot whale '' Globicephala melas and G. muerorhynchus''" pp. 847–52 in
Encyclopedia of Marine Mammals
', Perrin, W. F., Wursig, B., and Thewissen, J. G. M. (eds.), Academic Press; 2nd edition, About 150,000 individuals are found in the eastern tropical Pacific Ocean. More than 30,000 animals are estimated in the western Pacific, off the coast of Japan. Pilot whales are generally nomadic, but some populations stay year-round in places such as

Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...

and parts of California. They prefer the waters of the shelf break and

slope In mathematics, the slope or gradient of a Line (mathematics), line is a number that describes the direction (geometry), direction of the line on a plane (geometry), plane. Often denoted by the letter ''m'', slope is calculated as the ratio of t ...

. Once commonly seen off of Southern California, short-finned pilot whales disappeared from the area after a strong

El Niño EL, El or el may refer to: Arts and entertainment Fictional entities * El, a character from the manga series ''Shugo Chara!'' by Peach-Pit * Eleven (''Stranger Things'') (El), a fictional character in the TV series ''Stranger Things'' * El, fami ...

year in the early 1980s, according to the

National Oceanic and Atmospheric Administration The National Oceanic and Atmospheric Administration (NOAA ) is an American scientific and regulatory agency charged with Weather forecasting, forecasting weather, monitoring oceanic and atmospheric conditions, Hydrography, charting the seas, ...

. In October 2014, crew and passengers on several boats spotted a pod of 50–200 off

Dana Point, California Dana Point () is a city located in southern Orange County, California, United States. The population was 33,107 at the 2020 census. It has one of the few harbors along the Orange County coast; with ready access via State Route 1, it is a popu ...

Behavior and life history

Foraging and parasites

Although pilot whales are not known to have many predators, possible threats come from humans and killer whales. Both species eat primarily

squid A squid (: squid) is a mollusc with an elongated soft body, large eyes, eight cephalopod limb, arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida (though many other molluscs within the broader Neocoleoidea are also ...

. The whales make seasonal inshore and offshore movements in response to the dispersal of their prey. Fish that are consumed include

Atlantic cod The Atlantic cod (: cod; ''Gadus morhua'') is a fish of the family Gadidae, widely consumed by humans. It is also commercially known as '' cod'' or ''codling''.Atlantic mackerel,

Atlantic herring Atlantic herring (''Clupea harengus'') is a herring in the family Clupeidae. It is one of the most abundant fish species in the world. Atlantic herrings can be found on both sides of the northern Atlantic Ocean, congregating in large schools. ...

hake Hake is the common name for fish in the Merlucciidae family of the northern and southern oceans and the Phycidae family of the northern oceans. Hake is a commercially important fish in the same taxonomic order, Gadiformes, as cod and haddo ...

, and

spiny dogfish The spiny dogfish (''Squalus acanthias''), spurdog, mud shark, or piked dogfish is one of the best known species of the Squalidae (dogfish) family of sharks, which is part of the Squaliformes order. While these common names may apply to several ...

in the northwest Atlantic. In the

Faroe Islands The Faroe Islands ( ) (alt. the Faroes) are an archipelago in the North Atlantic Ocean and an autonomous territory of the Danish Realm, Kingdom of Denmark. Located between Iceland, Norway, and the United Kingdom, the islands have a populat ...

, whales mostly eat squid, but will also eat fish species such as greater argentine and blue whiting. However, Faroe whales do not seem to feed on

herring Herring are various species of forage fish, belonging to the Order (biology), order Clupeiformes. Herring often move in large Shoaling and schooling, schools around fishing banks and near the coast, found particularly in shallow, temperate wate ...

, or

mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...

, even when they are abundant. Pilot whales generally take several breaths before diving for a few minutes. Feeding dives may last over ten minutes. They are capable of diving to depths of 600 meters, but most dives are to a depth of 30–60 m. Shallow dives tend to take place during the day, while deeper ones take place at night. When making deep dives, pilot whales often make fast sprints to catch fast-moving prey such as squid. Compared to

sperm whale The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the Genus (biology), genus ''Physeter'' and one of three extant species in the s ...

s and beaked whales, foraging short-finned pilot whales are more energetic at the same depth. When they reach the end of their dives, pilot whales will sprint, possibly to catch prey, and then make a few buzzes. This is unusual, considering that deep-diving, breath-holding animals would be expected to swim slowly to conserve oxygen. The animal's high metabolism possibly allows it to sprint at deep depths, which would also give it shorter diving periods than some other marine mammals. This may also be the case for long-finned pilot whales. In 2024, a gps-fitted long-finned pilot whale recorded a diving depth over 1,100 meters. Pilot whales are often infested with whale lice,

cestodes Cestoda is a class of parasitic worms in the flatworm phylum (Platyhelminthes). Most of the species—and the best-known—are those in the subclass Eucestoda; they are ribbon-like worms as adults, commonly known as tapeworms. Their bodies con ...

, and

nematodes The nematodes ( or ; ; ), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (he ...

. They also can be hosts to various pathogenic bacteria and viruses, such as ''

Streptococcus ''Streptococcus'' is a genus of gram-positive spherical bacteria that belongs to the family Streptococcaceae, within the order Lactobacillales (lactic acid bacteria), in the phylum Bacillota. Cell division in streptococci occurs along a sing ...

'', ''

Pseudomonas ''Pseudomonas'' is a genus of Gram-negative bacteria belonging to the family Pseudomonadaceae in the class Gammaproteobacteria. The 348 members of the genus demonstrate a great deal of metabolic diversity and consequently are able to colonize a ...

'', ''

Escherichia ''Escherichia'' ( ) is a genus of Gram-negative, non-Endospore, spore-forming, Facultative anaerobic organism, facultatively anaerobic, rod-shaped bacteria from the family Enterobacteriaceae. In those species which are inhabitants of the gastroin ...

'', ''

Staphylococcus ''Staphylococcus'', from Ancient Greek σταφυλή (''staphulḗ''), meaning "bunch of grapes", and (''kókkos''), meaning "kernel" or " Kermes", is a genus of Gram-positive bacteria in the family Staphylococcaceae from the order Bacillale ...

'', and

influenza Influenza, commonly known as the flu, is an infectious disease caused by influenza viruses. Symptoms range from mild to severe and often include fever, runny nose, sore throat, muscle pain, headache, coughing, and fatigue. These sympto ...

. One sample of Newfoundland pilot whales found that the most common illness was an

upper respiratory tract infection An upper respiratory tract infection (URTI) is an illness caused by an acute infection, which involves the upper respiratory tract, including the nose, sinuses, pharynx, larynx or trachea. This commonly includes nasal obstruction, sore throat ...

Social structure

Both species live in groups of 10–30, but some groups may number 100 or more. Data suggest the social structures of pilot whale pods are similar to those of " resident" killer whales. The pods are highly stable and the members have close

matrilineal Matrilineality, at times called matriliny, is the tracing of kinship through the female line. It may also correlate with a social system in which people identify with their matriline, their mother's lineage, and which can involve the inheritan ...

relationships. Pod members are of various age and sex classes, although adult females tend to outnumber adult males. They have been observed making various kin-directed behaviors, such as providing food.Pryor, K., Norris K. S, Marsh, H., Kasuya, T. (1991) "Changes in the role of a female pilot whale with age", pp. 281–86 in
Dolphin societies
', Pryor, K., Norris K. S. (eds), Berkeley, CA: University of California Press. Numerous pods will temporarily gather, perhaps to allow individuals from different pods to interact and mate, as well as provide protection. Both species are loosely

polygynous Polygyny () is a form of polygamy entailing the marriage of a man to several women. The term polygyny is from Neoclassical Greek πολυγυνία (); . Incidence Polygyny is more widespread in Africa than in any other continent. Some scholar ...

. Data suggest both males and females remain in their mother's pod for life; despite this, inbreeding within a pod does not seem to occur. During aggregations, males will temporarily leave their pods to mate with females from other pods. Male reproductive dominance or competition for mates does not seem to exist. After mating, a male pilot whale usually spends only a few months with a female, and an individual may sire several offspring in the same pod. Males return to their own pods when the aggregations disband, and their presence may contribute to the survival of the other pod members. No evidence of "bachelor" groups has been found. Pilot whale pods off southern California have been observed in three different groups: traveling/hunting groups, feeding groups and loafing groups. In traveling/hunting groups, individuals position themselves in chorus lines stretching two miles long, with only a few whales underneath. Sexual and age-class segregation apparently occurs in these groups. In feeding groups, individuals are very loosely associated, but may move in the same direction. In loafing groups, whales number between 12 and 30 individuals resting. Mating and other behaviors may take place.

Reproduction and lifecycle

Pilot whales have one of the longest birth intervals of the cetaceans, calving once every three to five years. Most matings and calvings occur during the summer for long-finned pilot whales.Jefferson T. A., Leatherwood, S., Webber, M. A. (1993) "FAO Species identification guide. Marine mammals of the world". ''UNEP/FAO'', Rome. tp://ftp.fao.org/docrep/fao/009/t0725e/t0725e00.pdf Preview For short-finned pilot whales of the Southern Hemisphere, births are at their highest in spring and autumn, while in Northern Hemisphere, the time in which calving peaks can vary by population. For long-finned pilot whales, gestation lasts 12–16 months, and short-finned pilot whales have a 15-month gestation period. The calf nurses for 36–42 months, allowing for extensive mother-calf bonds. Young pilot whales will take milk until as old as 13–15 years of age. Short-finned pilot whale females will go through

, but this is not as common in females of long-finned pilot whales. Postreproductive females possibly play important roles in the survival of the young. Postreproductive females will continue to lactate and nurse young. Since they can no longer bear young of their own, these females invest in the current young, allowing them to feed even though they are not their own. Short-finned pilot whales grow more slowly than long-finned pilot whales. For the short-finned pilot whale, females become sexually mature at 9 years old and males at about 13–16 years. For the long-finned pilot whale, females reach maturity at around eight years and males at around 12 years.

Vocalizations

Pilot whales emit echolocation clicks for foraging and whistles and burst pulses as social signals (e.g. to keep contact with members of their pod). With active behavior, vocalizations are more complex, while less-active behavior is accompanied by simple vocalizations. Differences have been found in the calls of the two species. Compared with short-finned pilot whales, long-finned pilot whales have relatively low-frequency calls with narrow frequency ranges. In one study of North Atlantic long-finned pilot whales, certain vocalizations were heard to accompany certain behaviors. When resting or "milling", simple whistles are emitted. Surfacing behavior is accompanied by more complex whistles and pulsed sounds. The number of whistles made increases with the number of subgroups and the distance in which the whales are spread apart. A study of short-finned pilot whales off the southwest coast of

Tenerife Tenerife ( ; ; formerly spelled ''Teneriffe'') is the largest and most populous island of the Canary Islands, an Autonomous communities of Spain, autonomous community of Spain. With a land area of and a population of 965,575 inhabitants as of A ...

in the

Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...

has found the members of a pod maintained contact with each other through call repertoires unique to their pod. A later study found, when foraging at around 800 m deep, short-finned pilot whales make tonal calls. The number and length of the calls seem to decrease with depth despite being farther away from conspecifics at the surface. As such, the surrounding water pressure affects the energy of the calls, but it does not appear to affect the frequency levels. When in stressful situations, pilot whales produce "shrills" or "plaintive cries", which are variations of their whistles. To elude predators, long-finned pilot whales off the southern coast of Australia have been observed to mimic the calls of orcas while scavenging for food. This behaviour is thought to deter orca pods from approaching the pilot whales.

Antagonistic interactions with other species

Pilot whales have been occasionally observed mobbing or chasing other species of cetaceans. In several parts of the world, including off Iceland, long-finned pilot whales have been frequently documented chasing killer whales. The reasons for these chases are unknown, but it has been proposed that they might be due to either competition for prey or an anti-predation strategy. In 2021 an adult female killer whale with a newborn pilot whale travelling alongside her was observed off Western Iceland, leading scientists to question whether the relationship between these species might be far more complex than previously suggested. It is not known whether the newborn was adopted or abducted, but this same female killer whale was seen a year later interacting with a larger group of pilot whales. Based on experimentation involving familiar sounds of Orcas that consume fish and unfamiliar vocalizations of mammal-hunting Killer Whales, one study suggests that long-finned pilot whales can distinguish between familiar and unaccustomed types of Orca, noting behavioral differences like the ceasing of feeding when mammal-hunting Orcas' sounds were played. The study suggests that antagonistic interactions against fish-eating Killer Whales could either be an anti-predatory behavior or an attempt to maintain territory, while actions done in response to mammal-hunting Killer Whales could be a response to a more dangerous threat.

Stranding

Of the cetaceans, pilot whales are among the most common stranders. Because of their strong social bonds, whole groups of pilot whales will strand. Single stranders have been recorded and these are usually diseased. Group stranding tends to be of mostly healthy individuals. Several hypotheses have been proposed to explain group strandings. When using magnetic fields for navigation, the whales have been suggested to get perplexed by geomagnetic anomalies or they may be following a sick member of their group that got stranded. The pod also may be following a member of high importance that got stranded and a secondary social response makes them keep returning.Ridgway, S. H. (1998).
Handbook of Marine Mammals: The second book of dolphins and the porpoises, Volume 6
', Elsevier. pp. 245–69. Researchers from New Zealand have successfully used secondary social responses to keep a stranding pod of long-finned pilot whales from returning to the beach. In addition, the young members of the pod were taken offshore to buoys, and their distress calls lured the older whales back out to sea. In September 2022, nearly 200 pilot whales died after becoming stranded on Ocean Beach, part of Tasmania's west coast. Authorities said only about 35 survived of the 230 that were stranded.

Human interaction

The

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

lists long-finned pilot whales as "least concern" in the

Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological spe ...

. Long-finned pilot whales in the North and Baltic Seas are listed in Appendix II of the

Convention on the Conservation of Migratory Species of Wild Animals The Convention on the Conservation of Migratory Species of Wild Animals, also known as the Convention on Migratory Species (CMS) or the Bonn Convention, is an international agreement that aims to conserve migratory species throughout their r ...

(CMS). Those from northwest and northeast Atlantic may also need to be included to Appendix II of CMS. The short-finned pilot whale is listed on Appendix II of

CITES CITES (shorter acronym for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of inte ...

Hunting

The long-finned pilot whale has traditionally been hunted by "

driving Driving is the controlled operation and movement of a land vehicle, including cars, motorcycles, trucks, and buses. A driver's permission to drive on public highways is granted based on a set of conditions being met, and drivers are required to ...

", which involves many hunters and boats gathering in a semicircle behind a pod of whales close to shore, and slowly driving them towards a bay, where they become stranded and are then slaughtered. This practice was common in both the 19th and 20th centuries. The whales were hunted for bone, meat, oil, and fertilizer. In the

, pilot whale hunting started at least in the 16th century, and continued into modern times, with thousands being killed during the 1970s and 1980s. In other parts of the North Atlantic, such as

, West Greenland, Ireland and

Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer months. The ...

, pilot whales have also been hunted, but to a lesser extent. One fishery at Cape Cod harvested 2,000–3,000 whales per year during the late 19th and early 20th centuries.

's long-finned pilot whale fishery was at its highest in 1956, but declined shortly after and is now defunct. In the Southern Hemisphere, exploitation of long-finned pilot whales has been sporadic and low. Currently, long-finned pilot whales are only hunted at the Faroe Islands and Greenland. According to the IUCN the harvesting of this species for food in the Faroe Islands and Greenland has not resulted in any detectable declines in abundance. The short-finned pilot whale has also been hunted for many centuries, particularly by Japanese whalers. Between 1948 and 1980, hundreds of whales were exploited at

Hokkaido is the list of islands of Japan by area, second-largest island of Japan and comprises the largest and northernmost prefectures of Japan, prefecture, making up its own list of regions of Japan, region. The Tsugaru Strait separates Hokkaidō fr ...

and Sanriku in the north and Taiji, Izu, and

Okinawa most commonly refers to: * Okinawa Prefecture, Japan's southernmost prefecture * Okinawa Island, the largest island of Okinawa Prefecture * Okinawa Islands, an island group including Okinawa itself * Okinawa (city), the second largest city in th ...

in the south. These fisheries were at their highest in the late 1940s and early 1950s; 2,326 short-finned pilot whales were harvested in the mid- to late 1980s. This had decreased to about 400 per year by the 1990s. Pilot whales have also fallen victim to bycatches. In one year, around 30 short-finned pilot whales were caught by the squid round-haul fishery in southern California. Likewise, California's drift gill net fishery took around 20 whales a year in the mid-1990s. In 1988, 141 whales caught on the east coast of the U.S. were taken by the foreign Atlantic

fishery, which forced it to be shut down.

Pollution

As with other marine mammals, pilot whales are susceptible to certain pollutants. Off the Faroes, France, the UK, and the eastern US, pilot whales were found to have been contaminated with high amounts of DDT and PCB. Pollutants such as DDT and mercury can be passed from mothers to their babies during gestation and lactation. The Faroes whales have also been contaminated with

cadmium Cadmium is a chemical element; it has chemical symbol, symbol Cd and atomic number 48. This soft, silvery-white metal is chemically similar to the two other stable metals in group 12 element, group 12, zinc and mercury (element), mercury. Like z ...

and mercury. However, pilot whales from

and

Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...

were found to have had very low levels of DDT. Short-finned pilot whales off the west coast of the US have had high amounts of DDT and PCB in contrast to the low amounts found in whales from Japan and the Antilles.

Cuisine

Pilot

whale meat Whale meat, broadly speaking, may include all cetaceans (whales, dolphins, porpoises) and all parts of the animal: muscle (meat), organs (offal), skin (muktuk), and fat (blubber). There is relatively little demand for whale meat, compared to far ...

is available for consumption in very few areas of

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

, mainly along the central Pacific coast, and also in other areas of the world, such as the Faroe Islands. The meat is high in

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...

(higher than beef) and low in fat. Because a whale's fat is contained in the layer of

blubber Blubber is a thick layer of Blood vessel, vascularized adipose tissue under the skin of all cetaceans, pinnipeds, penguins, and sirenians. It was present in many marine reptiles, such as Ichthyosauria, ichthyosaurs and plesiosaurs. Description ...

beneath the skin, and the muscle is high in myoglobin, the meat is a dark shade of red. In Japan, where pilot whale meat can be found in certain restaurants and '' izakayas'', the meat is sometimes served raw, as ''

sashimi is a Japanese cuisine, Japanese delicacy consisting of fresh raw fish or Raw meat, meat sliced into thin pieces and often eaten with soy sauce. Origin The word ''sashimi'' means 'pierced body', i.e., "wikt:刺身, 刺身" = ''sashimi'', whe ...

'', but just as often pilot whale steaks are marinated, cut into small chunks, and grilled. When grilled, the meat is slightly flaky and quite flavorful, somewhat gamey, though similar to a quality cut of

beef Beef is the culinary name for meat from cattle (''Bos taurus''). Beef can be prepared in various ways; Cut of beef, cuts are often used for steak, which can be cooked to varying degrees of doneness, while trimmings are often Ground beef, grou ...

, with distinct yet subtle undertones recalling its marine origin. In both Japan and the Faroe Islands, the meat is contaminated with mercury and cadmium, causing a health risk for those who frequently eat it, especially children and pregnant women. In November 2008, an article in ''

New Scientist ''New Scientist'' is a popular science magazine covering all aspects of science and technology. Based in London, it publishes weekly English-language editions in the United Kingdom, the United States and Australia. An editorially separate organ ...

'' reported that research done on the Faroe Islands resulted in two chief medical officers recommending against the consumption of pilot whale meat, considering it to be too toxic. In 2008, the local authorities recommended that pilot whale meat should no longer be eaten due to the contamination. This has resulted in reduced consumption, according to a senior Faroese health official.

Captivity

Pilot whales, mostly short-finned pilot whales, have been kept in captivity in various marine parks, arguably starting in the late 1940s. Since 1973, some long-finned pilot whales from New England waters were taken and temporarily kept in captivity. Short-finned pilot whales off southern California, Hawaii and Japan have been kept in

aquarium An aquarium (: aquariums or aquaria) is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. fishkeeping, Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aquati ...

s and

oceanarium An oceanarium can be either a marine mammal park, such as Marineland of Canada, or a large-scale Public aquarium, aquarium, such as the Lisbon Oceanarium, presenting an ocean habitat with marine (ocean), marine animals, especially large ocean dwe ...

s. Several pilot whales from southern California and Hawaii were taken into captivity during the 1960s and early 1970s, two of which were placed at

SeaWorld San Diego SeaWorld San Diego is a theme park in Mission Bay Park in San Diego, California. It is a marine mammal park, oceanarium, public aquarium, and marine animal rehabilitation center. SeaWorld, the theme park's proprietor, is owned and operated by Un ...

. During the 1970s and early 1980s, six pilot whales were captured alive by drive hunts and taken for public display. Pilot whales have historically had low survival rates in captivity, with the average annual survival being 0.51 years during the mid-1960s to early 1970s. There have been a few exceptions to the rule. Bubbles, a female short-finned pilot whale, who was displayed in Marineland of the Pacific and eventually at Sea World California, lived to be somewhere in her 50s when she eventually died on 12 June 2016. In 1968, a pilot whale was captured, given the name Morgan, and trained by the U.S. Navy's Deep Ops to retrieve deeper-attached objects from the ocean floor. He dove a record depth of 1654 feet and was used for training until 1971.

Films

There are two documentaries entirely dedicated to the pilot whales. * Full-length ''Cheetahs of the deep'' (49’, 2014, directed by Rafa Herrero Massieu) — tells about the way of life, features of social interaction, the subtleties of hunting, games and breeding on the example of a group of non-migrating short-finned pilot whales living between the Islands of Tenerife and

La Gomera La Gomera () is one of Spain's Canary Islands, in the Atlantic Ocean off the coast of Africa. With an area of , it is the third-smallest of the archipelago's eight main islands. It belongs to the province of Santa Cruz de Tenerife. La Gomer ...

of the Canary archipelago. A curious feature of the film is that: “all marine mammals filmed in

freediving Freediving, free-diving, free diving, breath-hold diving, or skin diving, is a mode of underwater diving that relies on breath-holding until resurfacing rather than the use of breathing apparatus such as scuba gear. Besides the limits of breat ...

”. * Short film '' My Pilot, Whale'' (28’, 2014, directed by Alexander and Nicole Gratovsky) demonstrates the possibility of interaction between humans and free-living pilot whales, offering the viewer a number of philosophical questions related to cetaceans: about their attitude to the world, what we have in common, what we — humans — can learn from them, and so on. The film has received a number awards of international film festivals.

References

External links

Pilot Whales.org

{{Authority control Globicephalinae Cosmopolitan mammals Animals that use echolocation Taxa named by René Lesson

Naming

Taxonomy and evolution

Description

Distribution and habitat

Behavior and life history

Foraging and parasites

Social structure

Reproduction and lifecycle

Vocalizations

Antagonistic interactions with other species

Stranding

Human interaction

Hunting

Pollution

Cuisine

Captivity

Films

See also

References

External links