Catatumbo Moist Forests
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The Catatumbo moist forests (NT0108) is an ecoregion in Venezuela and Colombia to the west and east of
Lake Maracaibo Lake Maracaibo () is located in northwestern Venezuela, between the states of Zulia, Trujillo, and Mérida. While Maracaibo is commonly referred to as a lake, its current hydrological characteristics may better classify it as estuary and/or ...
. It includes four regions of
moist forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in large ...
on slightly higher ground than the
dry forest The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
and
mangrove A mangrove is a shrub or tree that grows mainly in coastal saline water, saline or brackish water. Mangroves grow in an equatorial climate, typically along coastlines and tidal rivers. They have particular adaptations to take in extra oxygen a ...
s that surround the lake. The forest has unique flora related to the flora of the Amazon basin. It is badly degraded due to farming, livestock grazing and oil exploration.


Geography


Location

The Catatumbo moist forests ecoregion is in western Venezuela and northeastern Colombia found on both sides of Lake Maracaibo. To the west it extends to the foothills of the Cordillera Oriental of the Colombian Andes, and in the south reaches the foothills of the
Venezuelan Andes The Venezuelan Andes (Spanish: ''Andes Venezolanos'') also simply known as the Andes (Spanish: ''Los Andes'') in Venezuela, are a mountain system that form the northernmost extension of the Andes. They are fully identified, both by their geologi ...
. It has an area of . In Venezuela the forest is in the states of
Zulia Zulia State (, ; Wayuu: ''Mma’ipakat Suuria'') is one of the 23 states of Venezuela. The state capital is Maracaibo. As of the 2011 census, it has a population of 3,704,404, the largest population among Venezuela's states. It is also one of t ...
and
Lara Lara may refer to: People * Lara (name), can be a given name or a surname in several languages * Lara (mythology), a naiad nymph, daughter of the river Almo in Ovid's ''Fasti'' Places *Lara (state), a state in Venezuela * Electoral district ...
. In Colombia it is in the
Norte de Santander Department Norte de Santander (Spanish for Northern Santander) () is a departments of Colombia, department of northeastern Colombia. It is in the north of the country, bordering Venezuela. Its capital is Cúcuta, one of the country's major cities. Norte ...
. The moist forests are mainly found to the southwest of Lake Maracaibo, but include two patches to the east of the lake surrounded by
Maracaibo dry forests The Maracaibo dry forests (NT0222) is an ecoregion in Venezuela around Lake Maracaibo. It contains the country's main oil fields. The habitat is criss-crossed by roads and is severely degraded by farming and livestock grazing. Geography Location ...
. A strip of these dry forests runs through the western section of the Catatumbo moist forests. To the west the forests give way to the
Cordillera Oriental montane forests The Cordillera Oriental montane forests (NT0118) is an ecoregion in Venezuela and Colombia along the east slopes of the eastern cordillera of the Andes. The extensive region of submontane and montane forests includes distinctive flora and fauna in ...
and to the south to the
Venezuelan Andes montane forests The Venezuelan Andes montane forests (NT0175) is an ecoregion in the northern arm of the Andes in Venezuela. It contains montane and cloud forests, reaching up to the high-level Cordillera de Merida páramo high moor ecoregion. The forests are hom ...
. Along the west shore of Lake Maracaibo the forests adjoin a belt of Amazon-Orinoco-Southern Caribbean mangroves.


Terrain

The moist forest is found on higher land among the dry forests of the southern Maracaibo basin, and on the foothills of the mountains to the west and south. The largest block has a V shape, covering the foothills and lower valleys of the Eastern Ranges and the
Mérida Andes Mérida or Merida may refer to: Places *Mérida (state), one of the 23 states which make up Venezuela *Mérida, Mérida, the capital city of the state of Mérida, Venezuela *Merida, Leyte, Philippines, a municipality in the province of Leyte *Mà ...
, which extend to the north and east respectively. Between this block and Lake Maracaibo there is another block on land with elevations of . On the east side of the lake there are blocks of moist forest on the
Cerro Cerrón Cerro is Spanish for "hill" or "mountain". People * Francisco Cerro (born 1988), Argentine footballer * Francisco Cerro Chaves (born 1957), Spanish prelate, theologian, and philosopher of the Catholic Church * Ian Cerro (born 1996), American ...
and on a hill reaching in the northern foothills of the
Cordillera de Mérida The Cordillera de Mérida is a mountain range, or massif, in northwestern Venezuela. The Cordillera de Mérida is a northeastern extension of the Andes Mountains and the most important branch of the Venezuelan Andes. The range runs approximately 4 ...
. Rivers crossing the south and west of the ecoregion include the
Catatumbo The Catatumbo River () is a river rising in northern Colombia, flowing into Lake Maracaibo in Venezuela. The Catatumbo River is approximately long. It forms a part of the international boundary between the two countries. The river flows throug ...
,
Bravo Bravo(s) or The Bravo(s) may refer to: Arts and entertainment Music Groups and labels * Bravo (band), a Russian rock band * Bravo (Spanish group), represented Spain at Eurovision 1984 * Bravo Music, an American concert band music publishing compa ...
and
Onia Onia may refer to: * Plural of onium, a bound state of a particle and its antiparticle * Onia, Arkansas, an unincorporated community in Stone County, Arkansas * Onia (clothing), a New York City based clothing company See also * Onias (disambigu ...
. The Palmar, Apón and Santa Ana rivers have their sources in the
Serranía del Perijá The Serranía del Perijá, Cordillera de Perijá or Sierra de Perijá is a mountain range, an extension of the eastern Andean branch ( Cordillera Oriental), in northern South America, between Colombia and Venezuela, ending further north in the ...
.


Climate

The
Köppen climate classification The Köppen climate classification divides Earth climates into five main climate groups, with each group being divided based on patterns of seasonal precipitation and temperature. The five main groups are ''A'' (tropical), ''B'' (arid), ''C'' (te ...
is "Aw": equatorial, winter dry. At a sample location at coordinates mean temperatures are fairly constant throughout the year at . Annual rainfall at this location averages about . It rains throughout the year, with peaks in March and July. The highest rainfall is in the southwest of the Maracaibo basin, where the prevailing winds meet the cordilleras and release their moisture. Annual rainfall in this area is up to .


Ecology

The ecoregion is in the
neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeogra ...
realm, in the
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in la ...
biome. The portion of the ecoregion to the south and west of Lake Maracaibo is viewed as a
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
refugium for woody plant families, and is the only area north of the Andes still holding remnants of Brazil and Colombia's Amazonian flora. The forested lower slopes of the Onia River basin south of the lake have a number of relict plant species.


Flora

Common plant families are ''
Bombacaceae Bombacaceae were long recognised as a family of flowering plants or Angiospermae. The family name was based on the type genus '' Bombax''. As is true for many botanical names, circumscription and status of the taxon has varied with taxonomic poi ...
'', ''
Combretaceae The Combretaceae, often called the white mangrove family, are a family of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, G ...
'', ''
Lecythidaceae The Lecythidaceae ( ) comprise a family of about 20 genera and 250–300 species of woody plants native to tropical South America, Africa (including Madagascar), Asia and Australia. Well known members of the family include the cannonball tree ...
'', ''
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
'', ''
Sapotaceae 240px, '' Madhuca longifolia'' var. ''latifolia'' in Narsapur, Medak district, India The Sapotaceae are a family of flowering plants belonging to the order (biology)">order Ericales">family (biology)">family of flowering plants belonging to th ...
'', ''
Tiliaceae Tiliaceae () is a family of flowering plants. It is not a part of the APG, APG II and APG III classifications, being sunk in Malvaceae mostly as the subfamilies Tilioideae, Brownlowioideae and Grewioideae, but has an extensive historical re ...
'' and ''
Vochysiaceae Vochysiaceae is a plant family belonging to the order Myrtales. Description Trees or shrubs with opposite leaves; flowers are zygomorph 1-(3)-5 merous; ovary inferior or superior; one fertile stamen; fruits samara or capsules. Biogeography ...
''. The upper canopy at about includes ''
Anacardium excelsum ''Anacardium excelsum'', the wild cashew, ''espavél'' or ''espavé'', is a tree in the flowering plant family Anacardiaceae. The tree is common in the tropical and subtropical dry broadleaf forests of Pacific and Atlantic watersheds of Central ...
'', ''
Carapa guianensis ''Carapa guianensis'' is a species of tree in the family Meliaceae, also known by the common names andiroba or crabwood. Description Andiroba is native to the Amazon and is widely used by the indigenous populations of the northern region of ...
'', ''
Ceiba pentandra ''Ceiba pentandra'' is a tropical tree of the order (biology), order Malvales and the family (biology), family Malvaceae (previously emplaced in the family Bombacaceae), native to Mexico, Central America and the Caribbean, northern South America, ...
'', '' Coumarouna punctata'', '' Couroupita guianenesis'', ''
Eschweilera ''Eschweilera'' is a genus of woody plants in the family Lecythidaceae first described as a genus in 1828.Sterculia apetala ''Sterculia apetala'', the Panama tree, is a species of flowering plant in the family Malvaceae. It is found in Florida, southern Mexico, Central America, and northern South America, and has been introduced to the Caribbean islands. ''Sterculia a ...
''. The middle canopy at about contains '' Calophllum brasiliense'', '' Guarea thichioides'', ''
Parkia pendula ''Parkia pendula'' is a species of neotropical evergreen tree found throughout Central and South America.Hopkins, H.C., Parkia (Leguminosae: Mimosoideae). Flora Neotropica, 1986: p. 1-123. It is part of the ''Parkia'' genus, a group of flowering ...
'', '' Pentaclethra macrobola'' and ''
Swartzia ''Swartzia'' is a genus of flowering plants in the family Fabaceae. It was named in honor of Swedish botanist Olof Swartz and contains about 200 species. ''Swartzia'' is restricted in its geographical distribution to the New World Tropics, where ...
'' species. The lower canopy at about contains ''
Combretum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar ...
'' species, ''
Inga ''Inga'' is a genus of small tropical, tough-leaved, nitrogen-fixing treesElkan, Daniel. "Slash-and-burn farming has become a major threat to the world's rainforest" ''The Guardian'' 21 April 2004 and shrubs, subfamily Mimosoideae. ''Inga''s l ...
'' species, ''
Luehea ''Luehea'' is a genus of trees in the family Malvaceae. Its native range stretches from Mexico to southern tropical America and Cuba. It is native to the countries of; Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, El S ...
'' species, ''
Protium Hydrogen (H) has three naturally occurring isotopes: H, H, and H. H and H are stable, while H has a half-life of years. Heavier isotopes also exist; all are synthetic and have a half-life of less than 1 zeptosecond (10 s). Of these, H is ...
'' species, '' Trichilia pleeana'' and '' Trichilia maynasiana''. Other flora include '' Gustavia hexapetala'', '' Cariniana pyriformis'', ''
Faramea capillipes ''Faramea'' is a genus of plants in the family Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs tha ...
'', '' Ochoterenaea colombiana'', '' Miconia mocquerysii'' and '' Vochysia lehmannii''. In the western and southern parts of the ecoregion, which has been degraded by logging, agriculture and livestock grazing, there is secondary growth of the genera '' Casearia'', ''
Cecropia ''Cecropia'' is a Neotropical genus consisting of 61 recognized species with a highly distinctive lineage of dioecious trees. The genus consists of pioneer trees in the more or less humid parts of the Neotropics, with the majority of the speci ...
'', '' Croton'', ''
Inga ''Inga'' is a genus of small tropical, tough-leaved, nitrogen-fixing treesElkan, Daniel. "Slash-and-burn farming has become a major threat to the world's rainforest" ''The Guardian'' 21 April 2004 and shrubs, subfamily Mimosoideae. ''Inga''s l ...
'', '' Isertia'', ''
Jacaranda ''Jacaranda'' is a genus of 49 species of flowering plants in the family Bignoniaceae, native to tropical and subtropical regions of the Americas while cultivated around the world. The generic name is also used as the common name. The species ' ...
'', ''
Trema Trema may refer to: * a Greek and Latin root meaning ''hole'' * Trema, a term for the two dots (diacritic) ** Tréma, (French), a diaeresis * ''Trema'' (plant), a genus of about 15 species of small evergreen trees * Tréma (record label), a Fr ...
'' and ''
Vismia ''Vismia'' is a genus of flowering plants in the family Hypericaceae. Members of the genus are small trees and shrubs found in tropical and subtropical areas of Central America and South America. Including the countries of Belize, Bolivia, Brazil ...
''. Isolated species along the Onia River include ''
Faramea capillipes ''Faramea'' is a genus of plants in the family Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs tha ...
'' associated with other lowland tropical forest species such as ''
Tapirira guianensis ''Tapirira guianensis'' is a species of plant in the family Anacardiaceae. It is native to Cerrado and Atlantic Forest (ecoregions) vegetation in Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South ...
'', '' Diplasis karataefolia'', '' Maprounea guianensi'', '' Olyra micrantha'', '' Leandra solenifera'', '' Miconia barbinervis'', '' Miconia nervosa'', '' Mouriri myrtifolia'', '' Abuta pahni'' and '' Psychotria capitata inudata''. Relict plant species in Catatumbo and eastern Colombia include '' Ochoterineae colombiana'', '' Miconia mocquerysii'', '' Palicourea buntingii'' and '' Vochysia lehmannii''. Endemic flora include '' Anthurium praemontanum'', '' Philodendron mesae'', '' Rodospatha perezii'', '' Spanthiphyllum perezii'' and '' Besleria ornata''. ''Besleria ornata'' is related to '' Besleria immitis'', which grows in the Colombian and Peruvian Amazon.


Fauna

The fauna of the Catatumbo moist forest is poorly documented. The
brown hairy dwarf porcupine The brown hairy dwarf porcupine (''Coendou vestitus'') is a species of rodent in the family Erethizontidae. Found in the Andes in Colombia and Venezuela, its natural habitat is subtropical or tropical moist lowland forests. It is not easy to stu ...
(''Coendou vestitus''), a very rare mammal, lives in warm lowlands up to elevations of about . Endangered mammals include
Geoffroy's spider monkey Geoffroy's spider monkey (''Ateles geoffroyi''), also known as the black-handed spider monkey or the Central American spider monkey, is a species of spider monkey, a type of New World monkey, from Central America, parts of Mexico and possibly a ...
(''Ateles geoffroyi''),
red-crested tree-rat The red-crested tree-rat or Santa Marta toro (''Santamartamys rufodorsalis'') is a species of tree-rat found in the monotypic genus ''Santamartamys'' in the family Echimyidae. It is nocturnal and is believed to feed on plant matter, and is mainly ...
(''Santamartamys rufodorsalis'') and
woodland Oldfield mouse The woodland Oldfield mouse (''Thomasomys hylophilus'') is a species of rodent in the family Cricetidae. It is found in Colombia and Venezuela Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast ...
(''Thomasomys hylophilus''). Endangered birds include black-and-chestnut eagle (''Spizaetus isidori'').


Status

The
World Wildlife Fund The World Wide Fund for Nature (WWF) is a Swiss-based international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the ...
gives the ecoregion the status "Critical/Endangered". Large areas of moist and dry forests have been destroyed around Lake Maracaibo, and the area is fragmented by the many roads in the region. Grazing animals, farms and oil exploration have drastically changed large areas. As of 2002 of the total in the ecoregion was agricultural land, while (27.9%) had natural cover or extractive land uses. Colonization is increasing, and oil exploration continues. On the mountain slopes areas have been deforested for shifting cultivation. The southwestern part of the region, considered to be a forest refuge, is the most degraded. Protected areas include the
Ciénagas del Catatumbo National Park The Ciénagas del Catatumbo National Park () Also Catatumbo Marshes National Park or Ciénagas de Juan Manuel National Park Is a protected area of Venezuela. The park is the second natural park of the Zulia state. It includes a segment of the Cat ...
in Venezuela. The
Catatumbo Barí National Natural Park The Catatumbo River () is a river rising in northern Colombia, flowing into Lake Maracaibo in Venezuela. The Catatumbo River is approximately long. It forms a part of the international boundary between the two countries. The river flows through ...
in the western section in Colombia mostly protects Cordillera Oriental montane forests rather than moist forests.


References


Bibliography

* * * * {{Biodiversity of Colombia Neotropical tropical and subtropical moist broadleaf forests Ecoregions of Colombia Ecoregions of Venezuela