This article records new
taxa
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
of
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
archosaur
Archosauria () or archosaurs () is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only extant taxon, extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistics ...
s of every kind that are
described during the year 2019, as well as other significant discoveries and events related to
paleontology
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure ge ...
of archosaurs that occurred in 2019.
Pseudosuchians
New pseudosuchian taxa
Pseudosuchian research
* A study on the bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of ''
Coahomasuchus chathamensis'', and on its implications for inferring
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
and growth strategy of this species, is published by Hoffman, Heckert & Zanno (2019).
* Tolchard ''et al.'' (2019) revise fragmentary archosaurian remains from the latest
Triassic
The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
lower
Elliot Formation
The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, so ...
(
South Africa
South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
), interpreting them as fossils of at least two distinct taxa of "
rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...
ns", thus representing the southernmost
palaeolatitude
Paleomagnetism (occasionally palaeomagnetism) is the study of prehistoric Earth's magnetic fields recorded in rocks, sediment, or archeological materials. Geophysicists who specialize in paleomagnetism are called ''paleomagnetists.''
Certain ...
s that these animals are known to have occurred, their first definitive remains from southern Africa, and some of the most recent records of members of this
grade
Grade most commonly refers to:
* Grading in education, a measurement of a student's performance by educational assessment (e.g. A, pass, etc.)
* A designation for students, classes and curricula indicating the number of the year a student has reach ...
.
* A study on the anatomy of the skeleton of ''
Poposaurus
''Poposaurus'' is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern and eastern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, bea ...
gracilis'' is published online by Schachner ''et al.'' (2019).
* A study on the age of sandstones of the
Badong Formation preserving fossils of ''
Lotosaurus
''Lotosaurus'' is an extinct genus of sail-backed poposauroid known from Hunan Province of central China.
Discovery
''Lotosaurus'' is known from the holotype IVPP V 4881 (or possibly V 4880), an articulated and well-preserved s ...
adentus'' is published by Wang ''et al.'' (2019).
* Description of the anatomy of the skull of a new specimen of ''Prestosuchus chiniquensis'' from the ''Dinodontosaurus'' Assemblage Zone of the Pinheiros-Chiniquá Sequence, Santa Maria Super sequence (
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
) is published by Mastrantonio ''et al.'' (2019), who also present the first description of a
rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...
n cranial
endocast
An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
.
* A study on habitat shifts during the evolutionary history of
Crocodylomorpha
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. Extinct crocodylomorphs were considerably mor ...
is published by Wilberg, Turner & Brochu (2019).
* A study on patterns of body size evolution of crocodylomorphs is published by Godoy ''et al.'' (2019).
* A study on the quality of the fossil record of non-marine crocodylomorphs is published by Mannion ''et al.'' (2019).
* A study on the evolution of skull shape in crocodylomorphs is published online by Godoy (2019).
* A study on the diversity of feeding ecologies of Mesozoic
crocodyliforms
Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
is published by Melstrom & Irmis (2019).
* A study on patterns of crocodyliform snout shape, on their inferred diet and on the relationship between form and function of crocodyliform skull shape throughout the evolutionary history of this group is published online by Drumheller & Wilberg (2019).
* New fossil material (an isolated left
dentary
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone ...
) of ''
Orthosuchus stormbergi'' is described from the Upper
Elliot Formation
The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, so ...
(
South Africa
South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
) by Dollman, Viglietti & Choiniere (2019), who also examine the
stratigraphic
Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks.
Stratigraphy has three related subfields: lithost ...
positions of all valid
crocodylomorph
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. Extinct crocodylomorphs were considerably mor ...
specimens from the main
Karoo Basin
The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a per ...
.
*
Teleosaurid and
metriorhynchid
Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period (Bajocian to early Aptian) of Europe, North America and South America. The name Metriorhynchidae ...
teeth are described from, respectively, the
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
(
Aalenian
The Aalenian () is a subdivision of the Middle Jurassic Epoch/Series of the geologic timescale that extends from about 174.7 ±0.8 Ma to about 170.9 ±0.8 Ma (million years ago). It was preceded by the Toarcian and succeeded by the Bajocian.
St ...
) and
Upper Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the name ...
(
Tithonian
In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
) of
Slovakia
Slovakia, officially the Slovak Republic, is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the west, and the Czech Republic to the northwest. Slovakia's m ...
by Čerňanský ''et al.'' (2019), representing the first record of members of both families from the country.
* Partial skeleton of a
teleosauroid
Teleosauroidea is an extinct superfamily of thalattosuchian crocodyliforms living from the Early Jurassic to the Early Cretaceous. It is phylogenetically defined by Mark T. Young and colleagues in 2024 in the ''PhyloCode'' as "the largest clade w ...
crocodylomorph, representing the most recent record of a definitive non-
machimosaurin teleosauroid in Africa reported so far, is described from the
Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
of
Tunisia
Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
by Dridi & Johnson (2019).
* Fossils of a member of Teleosauroidea with an estimated body length of 9.6 m, representing the most recent definitive record of Teleosauroidea reported so far, are described from the Lower Cretaceous (upper
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
)
Paja Formation
The Paja Formation (, K1p, Kip, Kimp, b3b6p) is an Early Cretaceous formation (geology), geologic formation of central Colombia. The formation extends across the northern part of the Altiplano Cundiboyacense, the Colombian emeralds#Western belt, W ...
(
Colombia
Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
) by Cortes ''et al.'' (2019).
* Redescription of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimens of ''
Mystriosaurus laurillardi'' and ''"
Steneosaurus
''Steneosaurus'' (from , 'narrow' and , 'lizard') is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic (Callovian or early Oxfordian) of France. The genus has been used as a wastebasket taxon for thalattosuchian ...
" brevior'' and a study on the taxonomic validity and phylogenetic relationships of these species is published by Sachs ''et al.'' (2019).
* A study on the anatomy and phylogenetic relationships of
metriorhynchoids
Metriorhynchoidea is an extinct superfamily (biology), superfamily of thalattosuchian crocodyliforms from the Early Jurassic to the Early Cretaceous (Toarcian - Valanginian, possibly as late as early Aptian) of Europe, North America and South Ame ...
from the Jurassic
Rosso Ammonitico Veronese Formation (
Italy
Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
) is published by Cau (2019), who provides a revised diagnosis of ''
Neptunidraco ammoniticus''.
* A three-dimensionally preserved
occiput
The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone lies over the occipital lobes of the ...
of a member of the genus ''
Torvoneustes'', indicating that members of this genus reached larger body sizes than previously supposed, is described from the Upper Jurassic
Kimmeridge Clay
The Kimmeridge Clay is a sedimentary rock, sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation (geology), form ...
Formation (
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
) by Young ''et al.'' (2019).
* A study on teeth morphology and
tooth enamel
Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The other ...
microstructure in ''
Mariliasuchus
''Mariliasuchus'' ("Marilia crocodile") is an extinct genus of Late Cretaceous notosuchian crocodyliforms found near Marilia, Brazil. The first bone remains were found and collected in 1995 by Brazilian paleontologist William Nava, in red rock ...
amarali'' is published by Augusta & Zaher (2019).
* A study on the arrangement and morphology of the
osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s of
baurusuchids
Baurusuchidae is a Gondwanan Family (biology), family of mesoeucrocodylians that lived during the Late Cretaceous. It is a group of terrestrial hypercarnivorous crocodilians from South America (Argentina and Brazil) and possibly Pakistan. Baurusu ...
is published by Montefeltro (2019).
* A study on the anatomy of the
pterygoid Pterygoid, from the Greek for 'winglike', may refer to:
* Pterygoid bone, a bone of the palate of many vertebrates
* Pterygoid processes of the sphenoid bone
** Lateral pterygoid plate
** Medial pterygoid plate
* Lateral pterygoid muscle
* Medial ...
region and skull airways of ''
Caipirasuchus
''Caipirasuchus'' is an extinct genus of sphagesaurid notosuchians known from the Late Cretaceous of northern São Paulo State and western Minas Gerais State, southeastern Brazil. The type species, ''C. paulistanus'', was named in 2011. A seco ...
paulistanus'' and ''C. montealtensis'' is published online by Dias ''et al.'' (2019), who report possible anatomical evidence of vocal capacity of ''C. montealtensis''.
* Description of fossils and possible
gastrolith
A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In ...
s of a large-bodied
sphagesaurid from the Upper Cretaceous Adamantina Formation (Brazil) is published online by Cunha ''et al.'' (2019).
* Description of new fossil material of ''
Pepesuchus'' from the
Upper Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
Adamantina Formation
The Adamantina Formation is a geological Formation (geology), formation in the Bauru Basin of western São Paulo (state), São Paulo state in southeastern Brazil.
Its strata date back to the Late Cretaceous epoch of the Cretaceous, Cretaceous Per ...
(
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
) and a study on the phylogenetic relationships of this taxon is published by Geroto & Bertini (2019).
* A study on the
diagenesis
Diagenesis () is the process of physical and chemical changes in sediments first caused by water-rock interactions, microbial activity, and compaction after their deposition. Increased pressure and temperature only start to play a role as sedi ...
of fossils of ''
Montealtosuchus
''Montealtosuchus'' is an extinct genus of terrestrial crocodyliform. It was discovered in 2004 in the Bauru Basin of Brazil, from Campanian-Maastrichtian deposits of the Adamantina Formation. The species was described in 2007, and assigned to t ...
arrudacamposi'' from the Upper Cretaceous Adamantina Formation is published by Marchetti ''et al.'' (2019).
* A study on the phylogenetic relationships of members of
Neosuchia
Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all Crocodylomorpha, crocodylomorphs more closely related to ''Croc ...
and on the evolution of longirostry in this group is published online by Groh ''et al.'' (2019).
* A study on the taxonomic status and phylogenetic relationships of ''
Sarcosuchus
''Sarcosuchus'' (), from Ancient Greek σάρξ (''sárx''), meaning "flesh", and σούχος (''soúkhos''), meaning "crocodile", is an extinct genus of crocodyliform a that lived during the Early Cretaceous, from the late Hauterivian to the ...
hartti'' is published online by Souza ''et al.'' (2019).
* Partial
dyrosaurid
Dyrosauridae is a family (biology), family of extinct neosuchian crocodyliforms that lived from the Campanian to the Eocene. Dyrosaurid fossils are globally distributed, having been found in Africa, Asia, Europe, North America and South America. ...
skeleton discovered in the 1930s in
Paleocene
The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
(
Danian
The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretac ...
) strata along the Atlantic coast of
Senegal
Senegal, officially the Republic of Senegal, is the westernmost country in West Africa, situated on the Atlantic Ocean coastline. It borders Mauritania to Mauritania–Senegal border, the north, Mali to Mali–Senegal border, the east, Guinea t ...
is described by Martin, Sarr & Hautier (2019).
* Description of new dyrosaurid specimens from the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
–early
Paleogene
The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
of
New Jersey
New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
(
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
), and a study on their implications for the validity of the species ''
Hyposaurus
''Hyposaurus'' is a genus of extinct marine Dyrosauridae, dyrosaurid crocodyliform. Fossils have been found in Paleocene aged rocks of the Iullemmeden Basin in West Africa, Campanian–Maastrichtian (Late Cretaceous, Late Cretaceous) Shendi Forma ...
rogersii'', is published online by Souza ''et al.'' (2019).
* Revision of the large-sized
neosuchia
Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all Crocodylomorpha, crocodylomorphs more closely related to ''Croc ...
ns ''
Kansajsuchus'' and "''
Turanosuchus''" from the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
of Central Asia is published by Kuzmin ''et al.'' (2019), who interpret ''Kansajsuchus'' as a member of
Paralligatoridae
Paralligatoridae is an extinct family of neosuchian crocodyliforms that existed during the Jurassic and Cretaceous periods. It includes the genera '' Paralligator'', '' Brillanceausuchus'', '' Kansajsuchus'', '' Shamosuchus'', '' Scolomastax'', ...
, and consider ''Turanosuchus aralensis'' to be a member of the genus ''Kansajsuchus'' belonging or related to the species ''K. extensus''.
* A study on the inner cavities of the skull of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''
Lohuecosuchus megadontos'' is published by Serrano-Martínez ''et al.'' (2019).
* Revision of the fossil material of ''
Allodaposuchus
''Allodaposuchus'' is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages, and possibly the Santonian stage, of the Late Cretaceous. Although generally classified as a non-cro ...
precedens'' from Vălioara (
Romania
Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
) is published online by Narváez ''et al.'' (2019), who emend the diagnosis for this species.
* A study on palaeodiversity of
eusuchia
Eusuchia is a clade of neosuchian Crocodylomorpha, crocodylomorphs that first appeared in the Early Cretaceous, which includes modern Crocodilia, crocodilians. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survi ...
ns over time is published online by De Celis, Narváez & Ortega (2019).
* A tooth of a juvenile specimen of ''
Deinosuchus
''Deinosuchus'' is an extinct genus of eusuchian, either an Alligatoroidea, alligatoroid Crocodilia, crocodilian or a stem-group crocodilian, which lived during the Late Cretaceous around . The first remains were discovered in North Carolina ...
'', providing new information on the
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
of this reptile, is described by Brownstein (2019).
* A well-preserved braincase of ''
Diplocynodon
''Diplocynodon'' is an extinct genus of eusuchian, either an alligatoroid crocodilian or a stem-group crocodilian, that lived during the Paleocene to Middle Miocene in Europe. Some species may have reached lengths of , while others probably did ...
tormis'' is described from the middle
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
site of 'Teso de la Flecha' (
Salamanca
Salamanca () is a Municipality of Spain, municipality and city in Spain, capital of the Province of Salamanca, province of the same name, located in the autonomous community of Castile and León. It is located in the Campo Charro comarca, in the ...
,
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) by Serrano-Martínez ''et al.'' (2019).
* A study on the anatomy and phylogenetic relationships of ''Diplocynodon hantoniensis'' is published online by Rio ''et al.'' (2020).
* Chroust, Mazuch & Luján (2019) describe new fossil material of ''Diplocynodon'' from four sites in the
Czech Republic
The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
dating to Eocene–Oligocene transition, and evaluate the implications of these fossils for the knowledge of the course of the Eocene–Oligocene cooling event in Central Europe.
* New crocodylian fossils, documenting the presence of four previously unrecognised alligatoroids, are described from the Lower
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
Castillo Formation (
Venezuela
Venezuela, officially the Bolivarian Republic of Venezuela, is a country on the northern coast of South America, consisting of a continental landmass and many Federal Dependencies of Venezuela, islands and islets in the Caribbean Sea. It com ...
) by Solórzano ''et al.'' (2019).
* A taxonomic and phylogenetic revision of ''
Necrosuchus
''Necrosuchus'' is an extinct genus of caiman from modern day Argentina that lived during the Paleocene epoch (Danian or possibly Selandian age, about 60 million years ago). It inhabited the marine or fluvio-lacustrine environment of the Patagoni ...
ionensis'' is published online by Cidade, Fortier & Hsiou (2019).
* Ten late Miocene specimens of ''
Mourasuchus
''Mourasuchus'' is an extinct genus of giant, aberrant caiman from the Miocene of South America. Its skull has been described as duck-like, being broad, flat, and very elongate, superficially resembling '' Stomatosuchus'' from the Late Cretace ...
'', tentatively assigned to the species ''M. arendsi'', are described from
Bolivia
Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
and from the
Solimões Formation
The Pebas Formation is a lithostratigraphic unit of Miocene age, found in western Amazonia. The formation extends over , including parts of Brazil, Peru, Ecuador and Colombia.Wesselingh et al., 2006 It is interpreted as representing the deposit ...
of
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
by Cidade ''et al.'' (2019), who also discuss the morphology of ''Mourasuchus'' and
paleogeographic
Palaeogeography (or paleogeography) is the study of historical geography, generally physical landscapes. Palaeogeography can also include the study of human or cultural environments. When the focus is specifically on landforms, the term paleo ...
distribution of this genus in the Miocene of South America.
* A study on the anatomy of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
of ''Mourasuchus amazonensis'' and on the taxonomic status of species belonging to the genus ''Mourasuchus'' is published by Cidade ''et al.'' (2019).
* A study on the feeding habits of ''Mourasuchus'' is published by Cidade, Riff & Hsiou (2019).
* A study on the structure of the
vertebral column
The spinal column, also known as the vertebral column, spine or backbone, is the core part of the axial skeleton in vertebrates. The vertebral column is the defining and eponymous characteristic of the vertebrate. The spinal column is a segmente ...
of ''
Purussaurus
''Purussaurus'' is an extinct genus of giant caiman that lived in South America during the Miocene epoch, from the Friasian to the Huayquerian in the SALMA classification. It is known from skull material found in the Brazilian and Peruvian Am ...
mirandai'', providing evidence of a deviation from the vertebral count present in all extant crocodilians, is published by Scheyer ''et al.'' (2019).
* A study on the taxonomic status of ''
Balanerodus logimus'' and ''
Caiman venezuelensis'' is published by Cidade ''et al.'' (2019).
* Fossils of a specimen of ''
Asiatosuchus
''Asiatosuchus'' is an extinct genus of crocodyloid crocodilians that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to ''Asiatosuchus'' since the genus was named in 1940. These speci ...
depressifrons'' from the late
Paleocene
The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
of Mont de
Berru
Berru () is a commune in the Marne department in northeastern France.
Berru, along with the neighboring commune of Cernay-lès-Reims, is the site of a geologic formation (part of the Paris Basin) that has yielded a significant number of P ...
(
France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
), representing the oldest European
crocodyloid
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to th ...
remains reported so far, are described by Delfino ''et al.'' (2019).
* A study on geographical origin, historical
biogeography
Biogeography is the study of the species distribution, distribution of species and ecosystems in geography, geographic space and through evolutionary history of life, geological time. Organisms and biological community (ecology), communities o ...
and evolution of traits aiding dispersal of members of the genus ''
Crocodylus
''Crocodylus'' is a genus of true crocodiles in the family Crocodylidae.
Taxonomy
The Genus, generic name, ''Crocodylus'', was proposed by Josephus Nicolaus Laurenti in 1768. ''Crocodylus'' contains 13–14 extant taxon, extant (living) species ...
'' is published by Nicolaï & Matzke (2019).
* Evidence of
gavialine-specific
atavistic
In biology, an atavism is a modification of a biological traits structure or behavior whereby an ancestral genetic trait reappears after having been lost through evolutionary change in previous generations. Atavisms can occur in several ways, ...
characters in the skeletons of fossil
tomistomines ''
Penghusuchus
''Penghusuchus'' is an extinct genus of gavialid crocodylian. It is known from a skeleton found in Middle to Upper Miocene rocks of Penghu Island, off Taiwan. The taxon was described in 2009 by Shan and colleagues; the type species is ''P. pani ...
pani'' and ''
Toyotamaphimeia
''Toyotamaphimeia'' (named after Toyotama-hime) is a genus of extinct gavialid crocodylian which lived in Japan and Taiwan during the Middle Pleistocene. A specimen recovered in 1964 at Osaka University during the construction of a new science bu ...
machikanensis'' is presented by Iijima & Kobayashi (2019).
* Skull and
mandibular
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone i ...
elements of a tomistomine (probably belonging to the genus ''
Maomingosuchus'') are described from the late
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
lignite
Lignite (derived from Latin ''lignum'' meaning 'wood'), often referred to as brown coal, is a soft, brown, combustible sedimentary rock formed from naturally compressed peat. It has a carbon content around 25–35% and is considered the lowest ...
seams of Krabi (
Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
) by Martin ''et al.'' (2019), providing evidence of tomistomines living in the tropics in the late Eocene.
* A revision of members of the genus ''
Gavialis
''Gavialis'' is a genus of crocodylians that includes the living gharial ''Gavialis gangeticus'' and one known extinct species, '' Gavialis bengawanicus.'' ''G. gangeticus'' comes from the Indian Subcontinent, while ''G. bengawanicus'' is know ...
'' described on the basis of fossils from the
Sivalik Hills
The Sivalik Hills, also known as Churia Hills, are a mountain range of the outer Himalayas.
The literal translation of "Sivalik" is 'tresses of Shiva'. The hills are known for their numerous fossils, and are also home to the Soanian Middle Pale ...
of
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
and
Pakistan
Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
is published by Martin (2019).
* A study on the systematics of crocodilians known from the
Oligocene
The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
fossil locality of Monteviale (
Italy
Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
) is published by Macaluso ''et al.'' (2019).
* A revision of fossil record of
Cenozoic
The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
crocodilians from
Sardinia
Sardinia ( ; ; ) is the Mediterranean islands#By area, second-largest island in the Mediterranean Sea, after Sicily, and one of the Regions of Italy, twenty regions of Italy. It is located west of the Italian Peninsula, north of Tunisia an ...
(Italy) is published by Zoboli ''et al.'' (2019).
* A review of the fossil crocodylomorph fauna of the Cenozoic of South America is published by Cidade, Fortier & Hsiou (2019).
* A method for the quantification of size- and shape-
heterodont
In anatomy, a heterodont (from Greek, meaning 'different teeth') is an animal which possesses more than a single tooth morphology.
Human dentition is heterodont and diphyodont as an example.
In vertebrates, heterodont pertains to animals wher ...
y in members of Crocodylia is presented by D'Amore ''et al.'' (2019), who apply their method to extant and fossil crocodylomorphs.
* A study on the global diversification dynamics of crocodylians since the Cretaceous is published online by Solórzano ''et al.'' (2019).
* A study testing whether the bone ornamentation may play a role in terms of load-bearing capacity and mechanical strength of pseudosuchian
osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s, based on data from five osteoderms of crocodylomorphs (representing four species: ''
Caiman crocodilus
The spectacled caiman (''Caiman crocodilus''), also known as the white caiman, common caiman, and speckled caiman, is a crocodilian in the family Alligatoridae. It is brownish-, greenish-, or yellowish-gray colored and has a spectacle-like ridge ...
'', ''
Osteolaemus tetraspis
The dwarf crocodile (''Osteolaemus tetraspis''), also known as the African dwarf crocodile, broad-snouted crocodile (a name more often used for the Asian mugger crocodile) or bony crocodile, is an African crocodile that is also the smallest extan ...
'', ''
Hyposaurus
''Hyposaurus'' is a genus of extinct marine Dyrosauridae, dyrosaurid crocodyliform. Fossils have been found in Paleocene aged rocks of the Iullemmeden Basin in West Africa, Campanian–Maastrichtian (Late Cretaceous, Late Cretaceous) Shendi Forma ...
rogersii'', ''
Sarcosuchus
''Sarcosuchus'' (), from Ancient Greek σάρξ (''sárx''), meaning "flesh", and σούχος (''soúkhos''), meaning "crocodile", is an extinct genus of crocodyliform a that lived during the Early Cretaceous, from the late Hauterivian to the ...
imperator'') and one aetosaur osteoderm (''
Aetosaurus
''Aetosaurus'' is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: ''A. ferratus'', the type species from Germany a ...
'' sp.), is published by Clarac ''et al.'' (2019).
* A study on the utility of head width as a body size proxy in extant crocodylians, and on its implications for estimates of body size of extinct
crocodyliforms
Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
, is published by O'Brien ''et al.'' (2019).
* A study comparing skull anatomy and inferred head musculature, stress distribution in skulls and feeding mechanisms in members of the genera ''
Pelagosaurus
''Pelagosaurus'' (meaning "lizard of the open sea") is an extinct genus of thalattosuchian crocodyliform that lived during the Toarcian stage of the Lower Jurassic, around 183 Ma to 176 Ma (million years ago), in shallow epicontinental seas t ...
'' and ''
Gavialis
''Gavialis'' is a genus of crocodylians that includes the living gharial ''Gavialis gangeticus'' and one known extinct species, '' Gavialis bengawanicus.'' ''G. gangeticus'' comes from the Indian Subcontinent, while ''G. bengawanicus'' is know ...
'', and evaluating changes in
mandibular
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone i ...
function and feeding through time in the macroevolution of Crocodylomorpha, is published by Ballell ''et al.'' (2019).
* Description of fossils of longirostrine crocodylians from the
Bartonian
The Bartonian is, in the International Commission on Stratigraphy's (ICS) geologic time scale, a stage or age in the middle of the Eocene Epoch or Series. The Bartonian Age spans the time between . It is preceded by the Lutetian and is follow ...
of southern
Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
is published by Jouve, Khalloufi & Zouhri (2019), who also discuss the implications of these fossils for the knowledge of the evolution of crocodylians through the Eocene–Oligocene transition.
* A study on the diversity of Late Jurassic crocodylomorph teeth from Valmitão (
Lourinhã Formation
The Lourinhã Formation () is a fossil-rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the ...
,
Portugal
Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
), and on the ecological niches and feeding behaviours of crocodylomorphs from this assemblage, is published online by Guillaume ''et al.'' (2019).
* Rivera-Sylva ''et al.'' (2019) report the first crocodyliform remains from La Parrita locality (
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
Cerro del Pueblo Formation
The Cerro del Pueblo Formation is a geological Formation (geology), formation in Saltillo, capital city of the state of Coahuila, Mexico, whose strata date back to the latest Campanian of the Late Cretaceous, just before the Campanian-Maastric ...
,
Coahuila
Coahuila, formally Coahuila de Zaragoza, officially the Free and Sovereign State of Coahuila de Zaragoza, is one of the 31 states of Mexico. The largest city and State Capital is the city of Saltillo; the second largest is Torreón and the thi ...
,
Mexico
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
).
* Description of an isolated crocodyliform tooth from the upper Eocene Ergilin Dzo Formation (
Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
) and a study on the implications of this fossil for the knowledge of the regional paleoclimate of the area of Mongolia during the late Eocene is published by Iijima ''et al.'' (2019).
* A study on the morphological diversity and phylogenetic affinities of crocodylomorph teeth from the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Tremp Formation
The Tremp Formation (, ), alternatively described as Tremp Group (), is a geological Formation (geology), formation in the comarca Pallars Jussà, Lleida Province, Lleida, Spain. The formation is restricted to the Tremp-Graus Basin, Tremp or T ...
(north-eastern
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) is published online by Blanco ''et al.'' (2019).
Non-avian dinosaurs
New dinosaur taxa
General non-avian dinosaur research
* A study aiming to identify the most likely area for the geographic origin of dinosaurs is published by Lee ''et al.'' (2019).
* A study evaluating the impact of new fossil discoveries and changing phylogenetic hypotheses on
biogeographical scenarios for dinosaur origins is published by Marsola ''et al.'' (2019).
* A study aiming to determine the degree of differences of dinosaur phylogenies inferred from skull and postcranial characters is published online by Li, Ruta & Wills (2019).
* A study on the
chronostratigraphic
Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time.
The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geological ...
position of the uppermost Cretaceous dinosaur localities from south-western Europe, and on their implications for inferring the course of the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
dinosaur turnover, is published by Fondevilla ''et al.'' (2019).
* A study aiming to quantify the habitat of latest Cretaceous North American dinosaurs, based on data from fossil occurrences and climatic and environmental modelling, and evaluating its implications for inferring whether dinosaur diversity was in decline prior to the
Cretaceous–Paleogene extinction event
The Cretaceous–Paleogene (K–Pg) extinction event, also known as the K–T extinction, was the extinction event, mass extinction of three-quarters of the plant and animal species on Earth approximately 66 million years ago. The event cau ...
, is published by Chiarenza ''et al.'' (2019).
* A study on factors determining the community richness of large herbivorous dinosaurs from the
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) is published by Mallon (2019).
* A review of the fossil record of Late Cretaceous non-avian dinosaurs from the James Ross Basin (
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
) is published by
Lamanna ''et al.'' (2019), who also describe fragmentary new
ankylosaur
Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful l ...
and
ornithopod
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...
material from the Cape Lamb Member of the
Snow Hill Island Formation
The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
and the Sandwich Bluff Member of the
Lopez de Bertodano Formation.
* A review and evaluation of studies on molecular data from Mesozoic dinosaur fossils is published by
Schweitzer Schweitzer is a surname. Notable people with the surname include:
* Albert Schweitzer (1875–1965), German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize
* Anton Schweitzer (1735–1787), German op ...
''et al.'' (2019).
* A study on the nature of putative remains of ancient proteins, blood vessels, and cells preserved with dinosaur fossils, based on data from fossils of ''
Centrosaurus
''Centrosaurus'' ( ; ) is a genus of centrosaurine ceratopsian dinosaur from Campanian age of Late Cretaceous Canada. Their remains have been found in the Dinosaur Park Formation, dating from 76.5 to 75.5 million years ago.
Discovery and nami ...
apertus'' from the
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
), is published by Saitta ''et al.'' (2019).
* A study on the
olfactory bulb
The olfactory bulb (Latin: ''bulbus olfactorius'') is a neural structure of the vertebrate forebrain involved in olfaction, the sense of smell. It sends olfactory information to be further processed in the amygdala, the orbitofrontal cortex (OF ...
ratio (the size of the olfactory bulb relative to the cerebral hemisphere) in dinosaurs, and on its implication for inferring olfactory acuity of dinosaurs, is published by Hughes & Finarelli (2019).
* A study on vascular correlates in dinosaur skulls, evaluating their implications for the knowledge of thermoregulatory strategies of dinosaurs of different sizes, is published online by Porter &
Witmer (2019).
* A review of the diversity of the musculature of the skulls of herbivorous dinosaurs is published online by Nabavizadeh (2019).
* A study on the evolution of different modes of herbivory in non-avian dinosaurs is published online by Button &
Zanno (2019).
* A study on the structure of eggshells of eggs produced by ''
Lufengosaurus
''Lufengosaurus'' (, meaning "Lufeng lizard") is a genus of massospondylid dinosaur which lived during the Early Jurassic period in what is now southwestern China."Lufengosaurus." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, ...
'', ''
Massospondylus
''Massospondylus'' ( ) is a genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been ...
'' and ''
Mussaurus
''Mussaurus'' (meaning "mouse lizard") is a genus of herbivorous sauropodomorph dinosaur that lived in southern Argentina during the Sinemurian stage of the Early Jurassic. It receives its name from the small size of the skeletons of juvenile and ...
'', representing the oldest confirmed
amniote
Amniotes are tetrapod vertebrate animals belonging to the clade Amniota, a large group that comprises the vast majority of living terrestrial animal, terrestrial and semiaquatic vertebrates. Amniotes evolution, evolved from amphibious Stem tet ...
eggshells reported so far, is published by Stein ''et al.'' (2019).
* Description of
dinosaur egg
Dinosaur eggs are the organic vessels in which a dinosaurs, dinosaur embryo develops. When the first scientifically documented remains of non-avian dinosaurs were being described in England during the 1820s, it was presumed that dinosaurs had la ...
fossils from the late
Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Chaochuan Formation (
Zhejiang
)
, translit_lang1_type2 =
, translit_lang1_info2 = ( Hangzhounese) ( Ningbonese) (Wenzhounese)
, image_skyline = 玉甑峰全貌 - panoramio.jpg
, image_caption = View of the Yandang Mountains
, image_map = Zhejiang i ...
,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) is published by Zhang ''et al.'' (2019), who name a new
ootaxon ''
Multifissoolithus chianensis''.
* Dinosaurs eggs assigned to the oofamily
Dendroolithidae
''Dendroolithus'' is an oogenus of Dendroolithid dinosaur egg found in the late Cenomanian Chichengshan Formation ( Tiantai Group), in the Gong-An-Zhai and Santonian Majiacun Formations of China and the Maastrichtian Nemegt and Campanian B ...
are described from the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
Zhaoying Formation (
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) by He ''et al.'' (2019), who name a new ootaxon ''
Pionoolithus quyuangangensis''.
* Dinosaurs eggs assigned to the oofamily
Faveoloolithidae
Faveoloolithidae is an oofamily of dinosaur eggs. It contains '' Faveoloolithus'', '' Hemifaveoloolithus'', '' Parafaveoloolithus'', and probably '' Sphaerovum''. However, unlike the other Faveoloolthids, ''Sphaerovum'' has compactituberculate ...
are described from the Upper Cretaceous (
Coniacian
The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by ...
–
Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
) siltstones within the Daeri Andesite of the Wido Volcanics (
South Korea
South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...
) by Kim ''et al.'' (2019), who name a new ootaxon ''
Propagoolithus widoensis''.
* Possible
dromaeosaurid
Dromaeosauridae () is a family (biology), family of feathered coelurosaurian Theropoda, theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous period (geology), Period. The name Drom ...
eggshells are described from the Upper Cretaceous Wido Volcanics (South Korea) by Choi & Lee (2019), who name a new ootaxon ''
Reticuloolithus acicularis''.
* Description of an intact dinosaur egg from the Cretaceous
Wayan Formation
The Wayan Formation is a geological Formation (geology), formation in Idaho whose strata date back to the latest Early Cretaceous and the earliest Late Cretaceous. Dinosaur, other reptile, mammal, and micro and macro-floral remains are among the f ...
(
Idaho
Idaho ( ) is a landlocked U.S. state, state in the Pacific Northwest and Mountain states, Mountain West subregions of the Western United States. It borders Montana and Wyoming to the east, Nevada and Utah to the south, and Washington (state), ...
,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) assigned to the oogenus ''
Macroelongatoolithus
''Macroelongatoolithus'' is an oogenus of large theropod dinosaur eggs, representing the eggs of giant caenagnathid oviraptorosaurs. They are known from Asia and from North America. Historically, several oospecies have been assigned to ''Macroelo ...
'' is published by Simon ''et al.'' (2019), who interpret this specimen as evidence of presence of a ''
Gigantoraptor
''Gigantoraptor'' () is a genus of large oviraptorosaur dinosaur that lived in Asia during the Late Cretaceous period. It is known from the Iren Dabasu Formation of Inner Mongolia, where the first remains were found in 2005.
''Gigantoraptor'' w ...
''-sized
oviraptorosaur
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or wi ...
in western North America.
* A study on the embryonic metabolism of ''
Troodon
''Troodon'' ( ; ''Troödon'' in older sources) is a controversial genus of relatively small, bird-like theropod dinosaurs definitively known from the Campanian age of the Late Cretaceous period (about 77 million years ago). It includes at l ...
formosus'', ''
Protoceratops
''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
andrewsi'' and ''
Hypacrosaurus
''Hypacrosaurus'' (meaning "near the highest lizard" reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high
Reek may refer to:
Places
* Reek, Netherlands, a village in the Dutch province of North Brabant
* Croagh Patrick, a mountain in the west of Ireland nicknamed "The Reek"
People
* Nikolai Reek (1890–1942), Estonian military commander
* Salme Ree ...
because it was almost but not quite as large as ''Tyrannosaurus'') is an extinct genus of hadrosaurid, duckbill dinosaur simila ...
stebingeri'', and on its implications for the knowledge of the incubation times for dinosaur eggs, is published by Lee (2019).
* A new dinosaur nesting site, preserving at least 15 egg clutches probably laid by a non-avian theropod dinosaur, is described from the Upper Cretaceous Javkhlant Formation (
Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
) by Tanaka ''et al.'' (2019), who interpret the finding as evidence of colonial nesting in a non-avian dinosaur.
* A study aiming to determine possible shifts from quadrupedality to bipedality during
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
in dinosaurs is published online by Chapelle ''et al.'' (2019).
* A review of evidence of probable responses of dinosaurs to serious injuries is presented by Hearn & Williams (2019).
* A study on the phylogenetic placement of ''
Chilesaurus
''Chilesaurus'' is an extinct genus of herbivorous dinosaur. While its exact classification is uncertain, many researchers believe it is a theropod, with a minority of academics suggesting that it may be an ornithischian. The type and only know ...
diegosuarezi'' and its implications for the phylogenetic relationships of major dinosaur groups is published by Müller & Dias-da-Silva (2019).
Saurischian research
Theropod research
* A study on specimen completeness in the fossil record of non-avian theropod dinosaurs is published by Cashmore & Butler (2019).
* A study on the distribution of discrete dental characters in theropod dinosaurs, and on the taxonomic value of theropod teeth, is published by Hendrickx ''et al.'' (2019).
* A study aiming to evaluate whether the maximum body size of theropod dinosaurs increased across the
Triassic
The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
-
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
boundary is published online by Griffin &
Nesbitt (2019).
* A revision of theropod dinosaur fossils from the Late Jurassic to mid-Cretaceous of Southeast Asia is published by Samathi, Chanthasit & Sander (2019).
* Description of theropod dinosaur teeth from the
Lower Cretaceous
Lower may refer to:
* ''Lower'' (album), 2025 album by Benjamin Booker
* Lower (surname)
* Lower Township, New Jersey
*Lower Receiver (firearms)
* Lower Wick Gloucestershire, England
See also
* Nizhny
{{Disambiguation ...
(
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
-
Aptian
The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
)
Ilek Formation
The Ilek Formation is a Lower Cretaceous geologic Formation (geology), formation in Western Siberia. Many different fossils have been recovered from the formation. It overlies the Late Jurassic Tyazhin Formation and underlies the Albian Kiya Form ...
(West Siberia,
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
) is published by Averianov, Ivantsov & Skutschas (2019).
* A study re-assessing the evidence for evolutionary
allometric
Allometry (Ancient Greek "other", "measurement") is the study of the relationship of body size to shape, anatomy, physiology and behaviour, first outlined by Otto Snell in 1892, by D'Arcy Thompson in 1917 in ''On Growth and Form'' and by Juli ...
trends in the forelimbs of non-avian theropod dinosaurs is published by Palma Liberona ''et al.'' (2019).
* Redescription of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''
Chindesaurus
''Chindesaurus'' ( ) is an extinct genus of basal saurischian dinosaur from the Late Triassic (213-210 million years ago) of the southwestern United States. It is known from a single species, ''C. bryansmalli'', based on a partial skeleton recov ...
bryansmalli'' and a study on the phylogenetic relationships of this species is published by Marsh ''et al.'' (2019).
* Description of two fragmentary
neotheropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodomor ...
specimens from the
Upper Triassic
The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...
Bull Canyon Formation
The Bull Canyon Formation is a geological Formation (geology), formation of Late Triassic (Norian) age in eastern New Mexico and the Texas Panhandle. It is one of several formations encompassed by the Dockum Group.
The Bull Canyon Formation prese ...
(
New Mexico
New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
), and a study on their implications for the knowledge of body size evolution among early theropods, is published by Griffin (2019).
* A study on the anatomy of the braincase, the skull
endocast
An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
and the inner ear of ''
Zupaysaurus
''Zupaysaurus'' (; "ZOO-pay-SAWR-us") is an extinct genus of early theropod dinosaur living during the Norian stage of the Late Triassic in what is now Argentina. Fossils of the dinosaur were found in the Los Colorados Formation of the Ischigual ...
rougieri'' is published by Paulina-Carabajal, Ezcurra &
Novas
A nova ( novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", Latin for "new") that slowly fades over weeks or months. All observed novae involve white d ...
(2019).
* A study on range of motion and functions of the forelimbs of ''
Dilophosaurus
''Dilophosaurus'' ( ) is a genus of theropod dinosaurs that lived in what is now North America during the Early Jurassic, about 186 million years ago. Three skeletons were discovered in northern Arizona in 1940, and the two best preserv ...
wetherilli'' is published by Senter & Sullivan (2019).
* A study on the paleobiology of ''
Cryolophosaurus
''Cryolophosaurus'' ( or ; ) is a genus of large theropod dinosaur known from only a single species, ''Cryolophosaurus ellioti'', from the Early Jurassic of Antarctica. It was one of the largest theropods of the Early Jurassic, with the subadult ...
'' is published by Yun (2019).
* A study on tooth formation and replacement rates in ''
Majungasaurus
''Majungasaurus'' (; ) is a genus of abelisaurid theropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period, making it one of the last-known non-avian dinosaurs that went extinct during th ...
'', ''
Ceratosaurus
''Ceratosaurus'' (from Greek 'horn' and 'lizard') is a genus of carnivorous theropod dinosaur that lived in the Late Jurassic period (Kimmeridgian to Tithonian ages). The genus was first described in 1884 by American paleontologist Othni ...
'' and ''
Allosaurus
''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
'' is published by D'Emic ''et al.'' (2019).
* A study on the ecology of ''
Ceratosaurus
''Ceratosaurus'' (from Greek 'horn' and 'lizard') is a genus of carnivorous theropod dinosaur that lived in the Late Jurassic period (Kimmeridgian to Tithonian ages). The genus was first described in 1884 by American paleontologist Othni ...
'' is published by Yun (2019).
* A study on the phylogenetic relationships of ''
Afromimus
''Afromimus'' (meaning "Africa mimic") is a genus of ceratosaurian theropod dinosaur from the Early Cretaceous Elrhaz Formation of Niger. It contains a single species, ''A. tenerensis'', named in 2017 by Paul Sereno from parts of the right leg, ...
tenerensis'' is published by Cerroni ''et al.'' (2019), who consider this taxon to be more likely an
abelisauroid rather than an
ornithomimosaur.
* Description of isolated neck vertebrae of abelisauroid theropods from the Cretaceous
Kem Kem Beds
The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations ...
(
Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
) and a study on the phylogenetic affinities of these fossils is published online by Smyth ''et al.'' (2019).
* Partially preserved
ilium of an indeterminate
abelisaur
Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with '' Ceratosaurus'' than with birds. The oldest known ceratosaur, '' Saltriovenator'', dates to the earlie ...
theropod is reported from the
Upper Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
Kem Kem Beds
The Kem Kem Group (commonly known as the Kem Kem beds) is a geological group in the Kem Kem region of eastern Morocco, whose strata date back to the Cenomanian stage of the Late Cretaceous. Its strata are subdivided into two geological formations ...
(
Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
) by Zitouni ''et al.'' (2019); however Smyth ''et al.'' (2019) reinterpret this specimen as a fossil of ''
Spinosaurus
''Spinosaurus'' (; ) is a genus of large spinosaurid theropod dinosaurs that lived in what now is North Africa during the Cenomanian faunal stage, stage of the Late Cretaceous Period (geology), period, about 100 to 94 annum, million year ...
aegyptiacus'',
while Samathi (2024) reinterprets this bone as a fossil of a spinosaurine spinosaurid of uncertain generic placement, possibly distinct from ''S. aegyptiacus''.
* A study on the anatomy of the brain, braincase and inner ear of ''
Carnotaurus
''Carnotaurus'' (; ) is a genus of Theropoda, theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 72 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a si ...
sastrei'' is published by Cerroni & Paulina-Carabajal (2019).
* A study on the phylogenetic affinities of a tooth previously considered to be part of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
of ''
Aerosteon
''Aerosteon'' is a genus of megaraptoran dinosaur from the Late Cretaceous period of Argentina. Its remains were discovered in 1996 in the Anacleto Formation, which is from the late Campanian. The type and only known species is ''A. riocolorad ...
riocoloradensis'' is published online by Hendrickx, Tschopp & Ezcurra (2019), who consider this fossil to be an
abelisaurid
Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
tooth.
*
Megalosaurid
Megalosauridae is a monophyletic family of carnivorous theropod dinosaurs within the group Megalosauroidea. Appearing in the Middle Jurassic, megalosaurids were among the first major radiation of large theropod dinosaurs. They were a relatively ...
teeth resembling teeth of ''
Torvosaurus
''Torvosaurus'' () is a genus of large Megalosaurinae, megalosaurine Theropoda, theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle Jurassic, Middle and Late Jurassi ...
'' are described from the Upper Jurassic of
Uruguay
Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
and
Tanzania
Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
by Soto, Toriño & Perea (2019).
* Isolated
spinosaurid
Spinosauridae (or spinosaurids) is a clade or Family (taxonomy), family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America, and Asia. ...
teeth are described from the Lower Cretaceous of Kut Island (
Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
) by Buffetaut ''et al.'' (2019).
* New spinosaurid specimens are described from the Kem Kem Beds (Morocco) by Arden ''et al.'' (2019), who interpret these specimens as providing evidence of aquatic adaptations in the skulls of spinosaurids, and name a new clade
Spinosaurini
Spinosauridae (or spinosaurids) is a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America, and Asia. Their remains have ...
; the study is subsequently criticized by Hone & Holtz (2019).
* New fossil material of juvenile spinosaurids is described from the Kem Kem Beds by Lakin & Longrich (2019).
* New theropod fossils, including partial tail vertebra of a member of
Megaraptora
Megaraptora is a clade of carnivorous theropod dinosaurs. Its derived members, the Megaraptoridae are noted for their large hand claws and powerfully-built forelimbs, which are usually reduced in size in other large theropods. Although undoubt ...
and an association of tail vertebrae and pelvic elements displaying a combination of characteristics that are present in megaraptorid and
carcharodontosaurid
Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae ...
theropods, are described from the early Late Cretaceous
Griman Creek Formation
The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian stages of the mid-Cretaceous. It is most notable being a major source of opal, ...
at
Lightning Ridge, New South Wales
Lightning Ridge is a small outback town in north-western New South Wales, Australia. Part of Walgett Shire, Lightning Ridge is situated near the southern border of Queensland, about east of the Castlereagh Highway. The Lightning Ridge area ...
(
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
) by Brougham, Smith & Bell (2019).
* Partial postcranial skeleton of a probable
carcharodontosauria
Carnosauria is an extinct group of carnivorous theropod dinosaurs that lived during the Jurassic and Cretaceous periods.
While Carnosauria was historically considered largely synonymous with Allosauroidea, some recent studies have revived Carn ...
n theropod is described from the
Upper Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the name ...
(
Tithonian
In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
)
Freixial Formation (
Portugal
Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
) by Malafaia ''et al.'' (2019).
* Description of the anatomy of the
axial skeleton
The axial skeleton is the core part of the endoskeleton made of the bones of the head and trunk of vertebrates. In the human skeleton, it consists of 80 bones and is composed of the skull (28 bones, including the cranium, mandible and the midd ...
of ''
Concavenator
''Concavenator'' (meaning Cuenca hunter) is a genus of carcharodontosaurian dinosaur that lived in Spain during the Early Cretaceous epoch, about 125 million years ago. The genus contains a single species, ''Concavenator corcovatus'', named and ...
corcovatus'' is published by Cuesta, Ortega & Sanz (2019).
* A study on the anatomy of the brain and inner ear of ''
Giganotosaurus
''Giganotosaurus'' ( ) is a genus of large theropod dinosaur that lived in what is now Argentina, during the early Cenomanian age of the Late Cretaceous period (geology), period, approximately 99.6 to 95 million years ago. The holotype specim ...
carolinii'' is published online by Paulina-Carabajal & Nieto (2019).
* A study on the anatomy of ''
Murusraptor
''Murusraptor'' ("wall thief") is a genus of carnivorous megaraptoran theropod dinosaur from the Sierra Barrosa Formation, part of the Neuquén Group of Patagonia, in Argentina, South America. It is known from a single specimen that consists of ...
barrosaensis'', and on its implications for inferring the phylogenetic placement of
megaraptora
Megaraptora is a clade of carnivorous theropod dinosaurs. Its derived members, the Megaraptoridae are noted for their large hand claws and powerfully-built forelimbs, which are usually reduced in size in other large theropods. Although undoubt ...
ns within Theropoda, is published by Rolando,
Novas
A nova ( novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", Latin for "new") that slowly fades over weeks or months. All observed novae involve white d ...
& Agnolín (2019).
* New fossil material of a megaraptorid belonging or related to the species ''
Australovenator
''Australovenator'' (meaning "southern hunter") is a genus of megaraptoran theropod dinosaur from Cenomanian (Late Cretaceous)-age Winton Formation (dated to 95 million years ago) of Australia. Some specimens from the Albian-aged Eumeralla Forma ...
wintonensis'' is described from the Lower Cretaceous (
Albian
The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) Eric the Red West site (
Eumeralla Formation
The Eumeralla Formation is a geological formation in Victoria, Australia whose strata date back to the Early Cretaceous. It is Aptian to Albian in age. Dinosaur remains are among the fossils that have been recovered from the formation, particul ...
;
Victoria
Victoria most commonly refers to:
* Queen Victoria (1819–1901), Queen of the United Kingdom and Empress of India
* Victoria (state), a state of Australia
* Victoria, British Columbia, Canada, a provincial capital
* Victoria, Seychelles, the capi ...
,
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
) by Poropat ''et al.'' (2019).
* A study comparing different methods of assessing morphological diversity of
coelurosauria
Coelurosauria (; from Greek, meaning "hollow-tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
n
mandible
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone i ...
s is published online by Schaeffer ''et al.'' (2019).
* A study on the anatomy of the skull of ''
Bicentenaria argentina'' is published online by Aranciaga-Rolando, Cerroni & Novas (2019).
* New postcranial bones of ''
Kileskus
''Kileskus'' (meaning ''lizard'' in the Khakas language) is a genus of tyrannosauroid dinosaur known from partial remains found in Middle Jurassic (Bathonian stage) Itat Formation of Sharypovsky District, Krasnoyarsk Krai (Russia). Fossils r ...
aristotocus'', providing new information on the anatomy of this species, are described from the Middle Jurassic (
Itat Formation
The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from ...
)
Itat Formation
The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from ...
(
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
) by Averianov ''et al.'' (2019).
* A study on the agility and turning capability of tyrannosaurids and other large theropods is published by Snively ''et al.'' (2019), who argue that tyrannosaurids could turn with greater agility, thus pivoting more quickly, than other large theropods, which enhanced their ability to pursue and subdue prey.
* A study on the taxonomic identity of the tyrannosaurid specimen
CMN 11315 from the lower
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Tolman Member of the
Horseshoe Canyon Formation
The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller.
The Horseshoe Canyon Formation is part of th ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) is published online by Mallon ''et al.'' (2019).
* A study on the taxonomic identity of the juvenile tyrannosaurid specimen
TMP TMP may refer to:
Arts and media
* Tickle Me Pink, a rock band from Colorado, US (2005–2011)
* Tiny Moving Parts, an emo band from Minnesota, US
* Tom Malone Prize, an Australian glass art prize
* Tsukuyomi -Moon Phase-, a 2000–2008 anime ...
1994.143.1, formerly assigned to the genus ''
Daspletosaurus
''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 77 and 74.4 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three named ...
'', is published by Voris ''et al.'', who reinterpret this specimen as belonging to the species ''
Gorgosaurus
''Gorgosaurus'' ( ; ) is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period (Campanian), between about 76.5 and 75 million years ago. Fossil remains have been found in the Ca ...
libratus'', and describe a new
postorbital
The ''postorbital'' is one of the bones in vertebrate skulls which forms a portion of the dermal skull roof and, sometimes, a ring about the orbit. Generally, it is located behind the postfrontal and posteriorly to the orbital fenestra. In some ve ...
from the
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) belonging to a small juvenile ''
Daspletosaurus
''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 77 and 74.4 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three named ...
''.
* A study on the tooth replacement patterns in tyrannosaurid theropods, as indicated by data from a juvenile specimen of ''
Tarbosaurus
''Tarbosaurus'' ( ; meaning "alarming lizard") is a genus of large tyrannosaurid dinosaur that lived in Asia during the Late Cretaceous epoch, about 70 million years ago (Maastrichtian age). It contains the single type species: ''Tarbosaurus ...
bataar'', is published by Hanai & Tsuihiji (2019).
* A study on teeth of ''Tarbosaurus bataar'' and its potential prey species from the
Nemegt Formation
The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
(
Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
), aiming to infer the diet of this dinosaur and seasonal climatic variations in the area of Mongolia in the early
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
on the basis of stable isotope data from
tooth enamel
Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The other ...
, is published online by Owocki ''et al.'' (2019).
* A study on the complexity and modularity of the skull of ''
Tyrannosaurus
''Tyrannosaurus'' () is a genus of large theropod dinosaur. The type species ''Tyrannosaurus rex'' ( meaning 'king' in Latin), often shortened to ''T. rex'' or colloquially t-rex, is one of the best represented theropods. It lived througho ...
rex'' is published by Werneburg ''et al.'' (2019).
* Traces preserved on a tail vertebra of a
hadrosaurid
Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
dinosaur from the
Upper Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
(
Montana
Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) are described by Peterson & Daus (2019), who interpret their finding as feeding traces produced by a late-stage juvenile ''Tyrannosaurus rex''.
* A large specimen of ''Tyrannosaurus rex'' (
RSM P2523.8) with an estimated body mass exceeding other known ''T. rex'' specimens and representatives of all other gigantic terrestrial theropods is described by Persons,
Currie
Currie is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edinburgh Council. It is situated be ...
& Erickson (2019).
* A study testing the biomechanical performance of the skull of ''Tyrannosaurus rex'' is published online by Cost ''et al.'' (2019).
* A study on tooth replacement in a well-preserved skull of ''Tyrannosaurus rex'' from the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
(
Montana
Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) is published online by Sattler & Schwarz (2019).
* A study aiming to determine the processes contributing to the preservation of soft tissue structures and proteins of ''Tyrannosaurus rex'' is published by Boatman ''et al.'' (2019).
* Teeth on a (probably non-tyrannosaurid) tyrannosauroid and a dromaeosaurid are described from the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Mount Laurel Formation (
New Jersey
New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) by Brownstein (2019).
* Description of an
ornithomimid
Ornithomimidae (meaning "bird-mimics") is an extinct family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period ...
specimen
UALVP 16182, putatively assigned to the genus ''
Dromiceiomimus
''Dromiceiomimus'' is a genus of ornithomimid theropod from the Late Cretaceous (early Maastrichtian) of Alberta, Canada. The type species, ''D. brevitertius'', is considered a synonym of ''Ornithomimus edmontonicus'' by some authors, while othe ...
'', and a study on the validity of this genus is published by Macdonald &
Currie
Currie is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edinburgh Council. It is situated be ...
(2019).
* A study on the
morphometrics
Morphometrics (from Greek μορΦή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
of teeth of ''
Richardoestesia
''Richardoestesia'' is a morphogenus of theropod dinosaur teeth, originally described from the Late Cretaceous of what is now Canada, the United States and Kazakhstan. It currently contains two species, ''R. gilmorei'' and ''R. isosceles'', a ...
asiatica'' from the Upper Cretaceous
Khodzhakul,
Bissekty and Aitym formations of
Uzbekistan
, image_flag = Flag of Uzbekistan.svg
, image_coat = Emblem of Uzbekistan.svg
, symbol_type = Emblem of Uzbekistan, Emblem
, national_anthem = "State Anthem of Uzbekistan, State Anthem of the Republ ...
is published by Averianov &
Sues (2019).
* A study on the bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of a
metatarsal bone
The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges ( toes). Lacking individual names, the metatarsal bones are ...
of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''
Xixianykus
''Xixianykus'' is a genus of alvarezsaurid theropod dinosaur from the late Cretaceous period of China. Known from a slab of rock which contains a partial postcranial skeleton unearthed in the Late Cretaceous ( Coniacian-Santonian) Majiacun For ...
zhangi'' is published by Qin, Zhao &
Xu (2019).
* A study on the anatomy of the skull of ''
Beipiaosaurus
''Beipiaosaurus'' is a genus of Therizinosauroidea, therizinosauroid theropod dinosaurs that lived in China during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the disc ...
inexpectus'' is published by Liao &
Xu (2019).
* A study on form, function and evolution of skulls of members of
Oviraptorosauria
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period (geology), Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like s ...
is published online by Ma ''et al.'' (2019).
* A study on the wing performance of ''
Caudipteryx
''Caudipteryx'' (meaning "tail feather") is a genus of small oviraptorosaur dinosaurs that lived in Asia during the Early Cretaceous, around 124.6 million years ago. They were feathered and extremely birdlike in their overall appearance, to the ...
'' is published by Talori ''et al.'' (2019).
* A study on the
aerodynamic
Aerodynamics () is the study of the motion of atmosphere of Earth, air, particularly when affected by a solid object, such as an airplane wing. It involves topics covered in the field of fluid dynamics and its subfield of gas dynamics, and is an ...
capacity of feathered forelimbs of ''Caudipteryx'' is published by Talori & Zhao (2019).
* Description of an
avimimid bonebed assemblage from the
Iren Dabasu Formation
The Iren Dabasu Formation (also known as Erlian Formation) is a Late Cretaceous geologic Formation (geology), formation in the Erenhot, Iren Nor region of Inner Mongolia. Dinosaur remains diagnostic to the genus level are among the fossils that h ...
of northern China, providing new information on the growth of avimimids, is published by Funston ''et al.'' (2019).
* Description of new
caenagnathid
Caenagnathidae is a family of derived caenagnathoid dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized ...
mandible
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone i ...
s from the
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) and a study on their
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
is published online by Funston ''et al.'' (2019).
* A reconstruction of the architecture of the
oviraptorid
Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one ...
egg clutch, based on data from five clutches from the Upper Cretaceous Nanxiong Group (
Jiangxi
; Gan: )
, translit_lang1_type2 =
, translit_lang1_info2 =
, translit_lang1_type3 =
, translit_lang1_info3 =
, image_map = Jiangxi in China (+all claims hatched).svg
, mapsize = 275px
, map_caption = Location ...
,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) is presented by Yang ''et al.'' (2019), who re-evaluate the hypothesis of thermoregulatory contact incubation of eggs as an explanation for the discoveries of associations of adult oviraptorosaurs with egg clutches.
* A study on the reproductive biology of oviraptorids, based on data from a partial clutch of eggs from the Upper Cretaceous Nanxiong Group, is published online by Yang ''et al.'' (2019).
* A study on the function of the enlarged "sickle claw" on the second toe of
dromaeosaurid
Dromaeosauridae () is a family (biology), family of feathered coelurosaurian Theropoda, theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous period (geology), Period. The name Drom ...
theropods is published by Bishop (2019).
* An
ungual
An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropod
Sauropoda (), whose members are known as sauropods (; ...
phalanx
The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
of a dromaeosaurid theropod is described from the
Blagoveshchensk
Blagoveshchensk ( rus, Благовещенск, p=bləɡɐˈvʲeɕːɪnsk, ) is a types of inhabited localities in Russia, city and the administrative center of Amur Oblast, Russia. It is located at the confluence of the Amur River, Amur and the ...
area (
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
) by Bolotskii, Bolotskii & Sorokin (2019).
* A study on the anatomy,
taphonomy
Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek language, Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientis ...
, environmental setting and phylogenetic position of ''
Halszkaraptor escuilliei'' is published by Brownstein (2019); the study is subsequently criticized by Cau (2020).
* A study on a fossil lizard found in the abdomen of a specimen of ''
Microraptor
''Microraptor'' (Greek language, Greek, μικρός, ''mīkros'': "small"; Latin language, Latin, ''raptor'': "one who seizes") is a genus of small, four-winged dromaeosaurid dinosaurs. Numerous well-preserved fossil specimens have been recovere ...
zhaoianus'' from the Lower Cretaceous
Jiufotang Formation
The Jiufotang Formation () is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms (see Jehol Biota). It is a member of the Jehol group. T ...
(
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
), evaluating its implications for the knowledge of dromaeosaurid digestion, is published by O'Connor ''et al.'' (2019).
* Description of the anatomy of the skull of ''
Saurornitholestes
''Saurornitholestes'' ("lizard-bird thief") is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta and Saskatchewan) and the United States (Montana, New Mexico, Alabama, and South Carolina).
Two spe ...
langstoni'', based on data from an almost complete skeleton from the
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
), is published online by
Currie
Currie is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edinburgh Council. It is situated be ...
& Evans (2019).
*
Histological
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
analysis of the forelimb bones of ''
Daliansaurus
''Daliansaurus'' (meaning "Dalian reptile") is a genus of small troodontid theropod dinosaur, measuring approximately long, from the Early Cretaceous of China. It contains a single species, ''D. liaoningensis'', named in 2017 by Shen and colleag ...
liaoningensis'' is presented by Shen ''et al.'' (2019).
* Evidence indicating that the
pennaceous feather
The pennaceous feather is a type of feather present in most modern birds and in some other species of maniraptoriform dinosaurs.
Description
A pennaceous feather has a stalk or quill. Its basal part, called a ''calamus'', is embedded in the sk ...
s of ''
Anchiornis
''Anchiornis'' is a genus of small, four-winged Paraves, paravian dinosaurs, with only one known species, the type species ''Anchiornis huxleyi'', named for its similarity to modern birds. The Latin name ''Anchiornis'' derives from a Greek word m ...
'' were composed of both feather
β-keratins and
α-keratins is presented by Pan ''et al.'' (2019); the study is subsequently criticized by Saitta & Vinther (2019).
* Isolated theropod teeth, interpreted as most likely representing at least two species, are described from the
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
Valtos Sandstone and
Lealt Shale Formations of
Skye
The Isle of Skye, or simply Skye, is the largest and northernmost of the major islands in the Inner Hebrides of Scotland. The island's peninsulas radiate from a mountainous hub dominated by the Cuillin, the rocky slopes of which provide some o ...
(
Scotland
Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
) by Young ''et al.'' (2019).
Sauropodomorph research
* A study aiming to explain high diversity of early evolutionary branches of
sauropodomorph
Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...
dinosaurs is published online by Müller & Garcia (2019).
* A study on the anatomy and phylogenetic relationships of ''
Pampadromaeus
''Pampadromaeus'' is an extinct genus of basal sauropodomorph dinosaurs known from the Late Triassic (Carnian) Santa Maria Formation of the Paraná Basin in Rio Grande do Sul, southern Brazil.Cabreira et al., 2011
Discovery
''Pampadromaeus'' ...
barberenai'' is published by Langer ''et al.'' (2019).
* A
dinosauriform
Dinosauromorpha is a clade of avemetatarsalians (archosaurs closer to birds than to crocodilians) that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, ...
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
, possibly of a juvenile specimen of the species ''Pampadromaeus barberenai'', is described from the
Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
of southern
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
by Müller ''et al.'' (2019).
* A study on the anatomy of the braincase of ''
Saturnalia tupiniquim'' is published by Bronzati, Langer & Rauhut (2019).
* Description of all available skull bones of ''Saturnalia tupiniquim'' except the braincase, evaluating the implications of this taxon for the knowledge of the early evolution of the sauropodomorph feeding behaviour, is published by Bronzati, Müller & Langer (2019).
* A study on the phylogenetic relationships of ''
Unaysaurus
''Unaysaurus'' is a genus of unaysauridae, unaysaurid sauropodomorpha, sauropodomorph herbivore dinosaur. Discovered in southern Brazil, in the geopark of Paleorrota, in 1998, and announced in a press conference on Thursday, December 3, 2004, it ...
tolentinoi'' is published online by McPhee ''et al.'' (2019).
* A study on the anatomy of the skull of ''
Macrocollum
''Macrocollum'' is a genus of unaysauridae, unaysaurid sauropodomorph dinosaur that lived during the Late Triassic period (geology), period (early Norian) in what is now Brazil. It is one of the oldest dinosaurs known.
Discovery and naming
''Mac ...
itaquii'' and on the phylogenetic relationships of this species is published online by Müller (2019).
* A study on the
bony labyrinth
The bony labyrinth (also osseous labyrinth or otic capsule) is the rigid, bony outer wall of the inner ear in the temporal bone. It consists of three parts: the vestibule, semicircular canals, and cochlea. These are cavities hollowed out of the ...
scale and geometry through
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
in ''
Massospondylus
''Massospondylus'' ( ) is a genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been ...
carinatus'', evaluating whether the putative gait change from quadrupedal juvenile to bipedal adult is reflected in labyrinth morphology, will be published by Neenan ''et al.'' (2019).
* Description of the anatomy of the postcranial skeleton of the
neotype
In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...
specimen of ''Massospondylus carinatus'' is published by Barrett ''et al.'' (2019).
* Redescription of the anatomy of the skull of ''
Jingshanosaurus
''Jingshanosaurus'' (meaning "Jingshan lizard") is a genus of sauropodomorph dinosaurs from the early Jurassic period 201.3 million years ago that went extinct 199.3 million years ago in the Hettangian Age.
Its maximum weight was around 4.3 t ...
xinwaensis'' is published online by Zhang ''et al.'' (2019), who consider ''
Chuxiongosaurus
''Chuxiongosaurus'' (meaning "Chuxiong lizard") is a genus of basal sauropodomorph dinosaur which lived during the Early Jurassic Period. Fossils of this genus have been found in the Lower Lufeng Formation, Yunnan Province, southern China. Ident ...
lufengensis'' to be a
junior synonym
In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently.
...
of ''J. xinwaensis''.
* A study on the anatomy of the
axial skeleton
The axial skeleton is the core part of the endoskeleton made of the bones of the head and trunk of vertebrates. In the human skeleton, it consists of 80 bones and is composed of the skull (28 bones, including the cranium, mandible and the midd ...
of ''
Xingxiulong
''Xingxiulong'' (meaning "Xingxiu Bridge dragon") is a genus of bipedal massopodan sauropodomorph from the Early Jurassic of China. It contains two species; the type species is ''X. chengi'', described by Wang et al. in 2017 from three specimens, ...
chengi'' is published online by Wang ''et al.'' (2019).
* A study on changes of body mass and
center of mass
In physics, the center of mass of a distribution of mass in space (sometimes referred to as the barycenter or balance point) is the unique point at any given time where the weight function, weighted relative position (vector), position of the d ...
of ''
Mussaurus
''Mussaurus'' (meaning "mouse lizard") is a genus of herbivorous sauropodomorph dinosaur that lived in southern Argentina during the Sinemurian stage of the Early Jurassic. It receives its name from the small size of the skeletons of juvenile and ...
patagonicus'' during its
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
, and on their potential relationship with the locomotor stance of this dinosaur, is published by Otero ''et al.'' (2019).
* A study on the leverage of forelimb muscles in the transition from the narrow-gauge stance of
basal sauropods to a wide-gauge stance in
titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
s is published by Klinkhamer ''et al.'' (2019).
* A study on the hind foot posture and
biomechanical capabilities of ''
Rhoetosaurus
''Rhoetosaurus'' (meaning "Rhoetos lizard"), named after Rhoetus, a titan in Greek mythology, is a genus of sauropod dinosaur from the Jurassic ( Oxfordian) Walloon Coal Measures of what is now eastern Australia. ''Rhoetosaurus'' is estimated ...
brownei'' is published by Jannel ''et al.'' (2019).
* A study on the age of the fossils of ''Rhoetosaurus brownei'' is published by Todd ''et al.'' (2019).
* An isolated tooth-crown of a member of
Eusauropoda
Eusauropoda (meaning "True Lizard Foot") is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of '' Shunosaurus'', and possibly '' Barapas ...
, possibly a member of
Mamenchisauridae
Mamenchisauridae is a family of sauropod dinosaurs belonging to the Eusauropoda known from the Jurassic and Early Cretaceous of Asia and Africa. Mamenchisaurids are characterized by their proportionately extremely long necks. Some members of the ...
or
Euhelopodidae
Euhelopodidae is a family of sauropod dinosaurs of disputed membership and affinities, which contains '' Euhelopus'' and its close relatives. Most proposed euhelopodids are from East Asia.
Euhelopodidae was first recognized by Carl Wiman in 1929 ...
, is described from the
Upper Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the name ...
Qigu Formation
The Qigu Formation is a Late Jurassic ( Oxfordian) geologic formation in the Southern Junggar Basin in China. Indeterminate Dinosaur remains are among the fossils that have been recovered from the formation, including the stegosaur (" Angustungu ...
(
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) by Maisch & Matzke (2019), representing the first record of a eusauropod from this formation reported so far.
* A
cervical vertebra
In tetrapods, cervical vertebrae (: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In sauropsid s ...
of a member of the genus ''
Omeisaurus
''Omeisaurus'' (meaning "Omei lizard") is a genus of sauropod dinosaur from the Middle to Late Jurassic Period (Bathonian- Oxfordian stage) of what is now China. Its name comes from Mount Emei, where it was discovered in the lower Shaximiao ...
'' is described from the
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
Lower Member of the
Shaximiao Formation
The Shaximiao Formation () is a Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the ...
(
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) by Tan ''et al.'' (2019), providing new information on the skeletal
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of this genus, and representing the easternmost occurrence of ''Omeisaurus'' reported so far.
* Possible
mamenchisaurid teeth are described from the
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
Itat Formation
The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from ...
(
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
) by Averianov ''et al.'' (2019).
* A study on the age of the fossils of members of the genus ''
Mamenchisaurus
''Mamenchisaurus'' ( , or spelling pronunciation ) is an extinct genus of sauropod dinosaurs known for their remarkably long necks which made up nearly half the total body length. Numerous species have been assigned to the genus; however, the val ...
'' from the
Suining Formation in the Sichuan Basin (China) is published by Wang ''et al.'' (2019).
* A study on the anatomy and affinities of ''
Lapparentosaurus
''Lapparentosaurus'' is a genus of sauropod dinosaur from the Middle Jurassic. Its fossils were found in Madagascar (Isalo III Formation). The type species is ''L. madagascariensis''.
Discovery and naming
In 1895 Richard Lydekker named a new s ...
madagascariensis'' is published by Raveloson, Clark & Rasoamiaramana (2019).
* Partial vertebra of a sauropod dinosaur belonging to the group
Turiasauria
Turiasauria is an unranked clade of eusauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa.
Description
Turiasauria was originally erected by Royo-Torres et al. (2006) to include '' Tur ...
is described from the
Lower Cretaceous
Lower may refer to:
* ''Lower'' (album), 2025 album by Benjamin Booker
* Lower (surname)
* Lower Township, New Jersey
*Lower Receiver (firearms)
* Lower Wick Gloucestershire, England
See also
* Nizhny
{{Disambiguation ...
Wealden Supergroup
The Wealden Group, occasionally also referred to as the Wealden Supergroup, is a group (a sequence of rock strata) in the lithostratigraphy of southern England. The Wealden group consists of paralic to continental (freshwater) facies sedimentar ...
(
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
) by Mannion (2019).
* Description and a study on the affinities of sauropod teeth from the
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
(
Bathonian
In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
)
Sakahara Formation (
Madagascar
Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
) is published online by Bindellini & Dal Sasso (2019), who report evidence of presence of
Titanosauriformes
Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located (hence ...
in the Bathonian.
* Description of isolated sauropod vertebrae from the
Oxford Clay
The Oxford Clay (or Oxford Clay Formation) is a Jurassic marine sedimentary rock formation underlying much of southeast England, from as far west as Dorset and as far north as Yorkshire. The Oxford Clay Formation dates to the Jurassic, specific ...
Formation (
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
), indicative of a higher sauropod biodiversity in this formation than previously recognised, is published by Holwerda, Evans & Liston (2019).
* Revision of the taxonomic diversity of sauropod dinosaurs from a historic
Carnegie Museum locality (Red Fork of the Powder River Quarry B) in northern
Wyoming
Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
(
Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
) is published by Tschopp ''et al.'' (2019).
* A study on pneumatic structures in the vertebrae of ''
Pilmatueia faundezi'' is published online by Windholz,
Coria & Zurriaguz (2019).
* A study on the anatomy of the
appendicular skeleton
The appendicular skeleton is the portion of the vertebrate endoskeleton consisting of the bones, cartilages and ligaments that support the paired appendages ( fins, flippers or limbs). In most terrestrial vertebrates (except snakes, legless li ...
of ''
Europasaurus
''Europasaurus'' (meaning 'Europe lizard') is a basal macronarian sauropod, a form of quadrupedal herbivorous dinosaur. It lived during the Late Jurassic (middle Kimmeridgian, from about 154 to 151 million years ago) of northern Germany, and has ...
holgeri'' and on the phylogenetic relationships of this species is published online by Carballido ''et al.'' (2019).
* Redescription of
brachiosaurid
The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the l ...
fossil material from the Upper Jurassic Morrison Formation (
Colorado
Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
), including a mostly complete skull discovered in
1883
Events
January
* January 4 – ''Life'' magazine is founded in Los Angeles, California, United States.
* January 10 – A fire at the Newhall Hotel in Milwaukee, Wisconsin, United States, kills 73 people.
* January 16 – ...
, is published online by D'Emic & Carrano (2019).
* A study on the phylogenetic relationships of ''
Galvesaurus
''Galvesaurus'', or ''Galveosaurus'', (meaning " Galve lizard") is a genus of brachiosaurid sauropod dinosaur from the Late Jurassic period. Fossils of the only known species, ''G. herreroi'', were found in Galve, Spain, hence its generic name, ...
herreroi'' is published by Pérez-Pueyo ''et al.'' (2019).
* A study on the phylogenetic relationships of the
Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
sauropod dinosaurs from the
Tendaguru Formation
The Tendaguru Formation, or Tendaguru Beds are a highly List of fossiliferous stratigraphic units in Tanzania, fossiliferous Lithostratigraphy, formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represe ...
of
Tanzania
Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
(''
Australodocus
''Australodocus'' (meaning "southern beam" from the Latin language, Latin ''australis'' "southern" and the Ancient Greek, Greek ''dokos''/δοκоς "beam") is a genus of sauropod dinosaur that lived during the Late Jurassic, Jurassic period, arou ...
bohetii'', ''
Janenschia
''Janenschia'' (named after Werner Janensch) is a large herbivorous sauropod dinosaur from the Late Jurassic Tendaguru Formation of Lindi Region, Tanzania around 155 million years ago.
Discovery and naming
''Janenschia'' has had a convoluted nom ...
robusta'' and ''
Tendaguria
''Tendaguria'' ( ; meaning "the Tendaguru one") is a genus of herbivorous sauropod dinosaur from the Late Jurassic of Lindi Region, Tanzania.
Discovery and naming
In 1911, German geologist Wilhelm Bornhardt at Nambango in German East Africa disc ...
tanzaniensis'') is published by Mannion ''et al.'' (2019).
* The first confirmed fossil of a sauropod dinosaur from
Ethiopia
Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
(an isolated tooth) is reported from the Upper Jurassic Mugher Mudstone by Goodwin ''et al.'' (2019).
* A study on the affinities of the sauropod dinosaur known from an isolated Metacarpal bones, metacarpal from the Upper Jurassic (Oxfordian (stage), Oxfordian) Jagua Formation (Cuba) is published online by Apesteguía, Izquierdo & Iturralde-Vinent (2019).
* A study on isolated sauropod teeth from the Early Cretaceous Teete locality (Batylykh Formation) (Yakutia,
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
), representing the northernmost sauropod record in the Northern Hemisphere reported so far, is published online by Averianov ''et al.'' (2019).
* Redescription of ''Jiangshanosaurus lixianensis'', a study on the anatomy of ''Dongyangosaurus sinensis'' and a study on the phylogenetic relationships of these species is published by Mannion ''et al.'' (2019).
* New fossil material of Titanosauriformes, titanosauriform sauropods is described from the Upper Cretaceous Daijiaping Formation (Hunan,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) by Han ''et al.'' (2019).
* A study on the long bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
in early juvenile titanosaur sauropods, evaluating its implications for the knowledge of early stages of development of these dinosaurs, is published online by González ''et al.'' (2019).
* A study on the neurology and phylogenetic affinities of a titanosaurian braincase from the
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
locality of Fox-Amphoux-Métisson (southeastern
France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
) is published by Knoll ''et al.'' (2019).
* Tail vertebrae of lithostrotian titanosaurs are described from the Lower Cretaceous
Ilek Formation
The Ilek Formation is a Lower Cretaceous geologic Formation (geology), formation in Western Siberia. Many different fossils have been recovered from the formation. It overlies the Late Jurassic Tyazhin Formation and underlies the Albian Kiya Form ...
(Krasnoyarsk Krai,
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
) by Averianov, Ivantsov & Skutschas (2019).
* A study on the anatomy of the
appendicular skeleton
The appendicular skeleton is the portion of the vertebrate endoskeleton consisting of the bones, cartilages and ligaments that support the paired appendages ( fins, flippers or limbs). In most terrestrial vertebrates (except snakes, legless li ...
of South American titanosaur sauropods and on its implications for the knowledge of the phylogenetic relationships of these sauropods is published by Bernardo Javier González Riga, González Riga ''et al.'' (2019), who name a new clade Colossosauria.
* Description of titanosaur sauropod vertebrae from the Upper Cretaceous Lameta Formation (
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
) is published by Jeffrey A. Wilson, Wilson ''et al.'' (2019).
* Description of the anatomy of the braincase of ''Malawisaurus dixeyi'' is published by Andrzejewski ''et al.'' (2019), who present digital reconstructions of the
endocast
An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
and inner ear of this species based on CT scanning.
* A study on the anatomy and phylogenetic relationships of ''Uberabatitan ribeiroi'' is published by Silva ''et al.'' (2019).
* A study on vertebral pneumaticity in ''Uberabatitan ribeiroi'', indicating that
diagenesis
Diagenesis () is the process of physical and chemical changes in sediments first caused by water-rock interactions, microbial activity, and compaction after their deposition. Increased pressure and temperature only start to play a role as sedi ...
can obliterate traces of bone pneumaticity, is published online by Aureliano ''et al.'' (2019).
* Fossils of a titanosaur sauropod related to ''Rapetosaurus'' and the indeterminate Italian titanosaur specimen Museo Civico di Storia Naturale di Milano, MSNM V7157 are described from the Algora vertebrate fossil site located in the Cenomanian strata of the Arenas de Utrillas Formation (
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) by Mocho ''et al.'' (2019).
* Description of five articulated sauropod dorsal vertebrae from the Upper Cretaceous
Nemegt Formation
The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians ...
, possibly belonging to the species ''Nemegtosaurus mongoliensis'', is published by Averianov & Lopatin (2019), who also study the anatomy of sauropod Femur, femora from the Nemegt Formation, and argue that ''N. mongoliensis'' is likely to be distinct from ''Opisthocoelicaudia skarzynskii''.
Ornithischian research
* A study on the evolution of the Femur, femoral fourth trochanter in ornithischian dinosaurs is published online by Persons &
Currie
Currie is a village and suburb on the outskirts of Edinburgh, Scotland, situated south west of the city centre. Formerly within the County of Midlothian, it now falls within the jurisdiction of the City of Edinburgh Council. It is situated be ...
(2019).
* A study on the
taphonomy
Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek language, Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientis ...
and
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of the ornithischian (ankylosaurian and ornithopod) fossils from the La Cantalera-1 site (
Lower Cretaceous
Lower may refer to:
* ''Lower'' (album), 2025 album by Benjamin Booker
* Lower (surname)
* Lower Township, New Jersey
*Lower Receiver (firearms)
* Lower Wick Gloucestershire, England
See also
* Nizhny
{{Disambiguation ...
Blesa Formation,
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) is published by Perales-Gogenola ''et al.'' (2019).
Thyreophoran research
* Studies on the anatomy of the skull and postcranial skeleton of ''Scelidosaurus harrisonii'' are published online by David B. Norman, Norman (2019).
* A study on the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Bienosaurus lufengensis'', and on the taxonomic validity and phylogenetic relationships of this dinosaur, is published by Raven ''et al.'' (2019).
* A study on the Morphology (biology), morphological diversity of Stegosauria, stegosaurs through the evolutionary history of the group is published by Romano (2019).
* Two new stegosaurian specimens from the northernmost outcrops of the
Morrison Formation
The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
in
Montana
Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
, one of which is the northernmost occurrence of a dinosaur from the Morrison Formation reported so far, are described by Woodruff, Trexler & Susannah Maidment, Maidment (2019).
* Description of a new specimen of ''Miragaia longicollum'' and a study on the taxonomic validity and phylogenetic relationships of this species is published by Costa & Octávio Mateus, Mateus (2019), who transfer the species ''Alcovasaurus longispinus'' to the genus ''Miragaia''.
* Plates of an armored dinosaur from the Lower Jurassic (Sinemurian-Pliensbachian) Lower Kota Formation (
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
) are redescribed by Peter Galton, Galton (2019), who considers these fossils to be more similar to plates of ankylosaurians than
basal thyreophorans, and interprets them as the earliest ankylosaurian fossils reported so far.
* Description of an assemblage of 12 partial, articulated or associated ankylosaurian skeletons and thousands of isolated bones and teeth from the Cretaceous (
Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
) Iharkút vertebrate locality (Hungary) will be published by Ősi ''et al.'' (2019).
* A study on the evolution of morphological traits associated with tail weaponry in ankylosaurs and glyptodonts, aiming to quantitatively test the hypothesis that tail weaponry of these groups is an example of convergent evolution, is published online by Victoria Arbour, Arbour &
Zanno (2019).
* Murray, Riguetti & Rozadilla (2019) describe ankylosaur
osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s from the Upper Cretaceous (
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
-
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
) Allen Formation (Argentina).
* Description of three new skull specimens of ''Talarurus plicatospineus'' from the Upper Cretaceous (Cenomanian–
Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
) Bayan Shireh Formation (
Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
), and a study on the phylogenetic relationships of this species, is published online by Park ''et al.'' (2019).
* Evidence for sexual dimorphism in ''Tianzhenosaurus youngi'' is presented by Ma (2019).
* A study on the brain morphology and topography of cranial nerves of ''Bissektipelta archibaldi'' is published by Alifanov & Saveliev (2019).
* A study on the bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Antarctopelta oliveroi'' and on its implication for the knowledge of paleobiology of this species is published by Cerda ''et al.'' (2019).
Cerapod research
* A study on the age of the Kulinda locality (south-eastern Siberia,
Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
) which yielded fossils of ''Kulindadromeus zabaikalicus'' is published by Cincotta ''et al.'' (2019).
* First Photogrammetry, photogrammetric models of the type locality burrow of ''Oryctodromeus cubicularis'' are presented by Wilson & Varricchio (2019).
* A study on the
taphonomy
Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek language, Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientis ...
of fossils of ''Oryctodromeus cubicularis'' is published by Krumenacker ''et al.'' (2019), who also report discovery of new burrows of this dinosaur.
* New fossil material of ornithopod dinosaurs is described from the Cretaceous Flat Rocks locality (Wonthaggi Formation,
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
) by Herne ''et al.'' (2019), who also revise ''Qantassaurus intrepidus'' and study the phylogenetic relationships of the Victoria (Australia), Victorian ornithopods.
* Two minuscule ornithopod Femur, femora, likely belonging to individuals around the point of hatching, are described from the Cenomanian
Griman Creek Formation
The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian stages of the mid-Cretaceous. It is most notable being a major source of opal, ...
(Australia) by Kitchener ''et al.'' (2019).
* Description of new fossil material of large ornithopod dinosaurs from the Lower Cretaceous localities in El Castellar (Maestrazgo Basin, Teruel,
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
), and a study on the implications of these fossils for the knowledge of ornithopod diversity in the Lower Cretaceous of the Iberian Peninsula, is published by Verdú ''et al.'' (2019).
* Description of the anatomy of the skeleton of ''Talenkauen santacrucensis'' is published by Rozadilla, Agnolín &
Novas
A nova ( novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", Latin for "new") that slowly fades over weeks or months. All observed novae involve white d ...
(2019).
* A study on the anatomy of the skeleton of ''Macrogryphosaurus gondwanicus'' is published online by Rozadilla, Cruzado-Caballero & Jorge O. Calvo, Calvo (2019).
* Skeletal pathologies affecting a subadult specimen of ''Tenontosaurus tilletti'' from the Antlers Formation of southeastern Oklahoma are described by Hunt ''et al.'' (2019).
* A study on the anatomy and phylogenetic relationships of the ornithopod dinosaurs from the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
of Crimea, including ''Riabininohadros weberae'', is published by Averianov & Lopatin (2019).
* Redescription of the fossil material of ''Orthomerus dolloi'' and a study on the phylogenetic affinities of this taxon is published online by Madzia, Jagt & Mulder (2019).
* A study on patterns and processes of morphological evolution of Hadrosauroidea, hadrosauroid dinosaurs is published by Stubbs ''et al.'' (2019).
* A study on the nature of the Fluvial processes, fluvial systems of Laramidia during the Late Cretaceous, as indicated by data from vertebrate and invertebrate fossils from the Kaiparowits Formation of southern Utah, and on the behavior of
hadrosaurid
Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...
dinosaurs over these landscapes, will be published by Crystal ''et al.'' (2019).
* Evidence of three-dimensional preservation of eumelanin-bearing bodies, dermal cells and blood vessel fragments in a hadrosaur specimen Peabody Museum of Natural History, YPMPU 016969 is presented by Fabbri ''et al.'' (2019).
* A study on the osteology and phylogenetic relationships of ''"Tanius laiyangensis"'' is published online by Zhang ''et al.'' (2019).
* A study on the bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of tibiae of ''Maiasaura peeblesorum'', focusing on the composition, frequency and cortical extent of localized vascular changes, is published by Woodward (2019).
* A study on hadrosaurine skulls from the
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
), aiming to assess the influence of
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
on skull morphology, and evaluating proposed synonymy between ''Gryposaurus incurvimanus'' and ''G. notabilis'', is published online by Lowi-Merri & Evans (2019).
* Three juvenile specimens of ''Prosaurolophus maximus'', providing new information on the
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
of this taxon, are described from the Bearpaw Formation (
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) by Drysdale ''et al.'' (2019).
* A study on the structure and contents of a large piece of amber attached to a jaw of a specimen of ''Prosaurolophus maximus'' from the Cretaceous
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
), evaluating the implications of this finding for the knowledge of the habitat and taphonomy of the dinosaur, is published by McKellar ''et al.'' (2019).
* A study on the impact of bone tissue structure, early Diagenesis, diagenetic regimes and other Taphonomy, taphonomic variables on the preservation potential of soft tissues in vertebrate fossils, as indicated by data from fossils of ''Edmontosaurus annectens'' from the Standing Rock Hadrosaur Site (
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Hell Creek Formation
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The Formation (stratigraphy), formation s ...
, South Dakota), is published by Ullmann, Pandya & Nellermoe (2019), who report the first recovery of osteocytes and vessels from a fossil vertebral centrum and ossified tendons.
* A
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
of an early juvenile hadrosaurid, probably belonging to the species ''Edmontosaurus annectens'', is described from the Hell Creek Formation (
Montana
Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
, United States) by Farke & Yip (2019), providing new information on Ontogeny, ontogenetic changes in the skeleton of this dinosaur.
* Skull remains of nestling-sized hadrosaurids, probably belonging to the species ''Edmontosaurus annectens'', are described from the Hell Creek Formation (Montana, United States) by Wosik, Goodwin & Evans (2019).
* A study of three-dimensionally preserved squamous skin of a member of the genus ''Edmontosaurus'' from the Upper Cretaceous (
Campanian
The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
) Wapiti Formation (
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) is published by Barbi ''et al.'' (2019).
* The first definitive Lambeosaurinae, lambeosaurine fossil (an isolated skull bone) is described from the Liscomb Bonebed of the Prince Creek Formation (Alaska,
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
) by Takasaki ''et al.'' (2019).
* Fossils of a lambeosaurine related to the Eurasian Tsintaosaurini are described from the lower
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
of the Els Nerets locality (eastern Tremp Syncline, northeastern
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) by Conti ''et al.'' (2019).
* A study on the microwear of hadrosaur teeth from the La Parrita locality (
Cerro del Pueblo Formation
The Cerro del Pueblo Formation is a geological Formation (geology), formation in Saltillo, capital city of the state of Coahuila, Mexico, whose strata date back to the latest Campanian of the Late Cretaceous, just before the Campanian-Maastric ...
,
Mexico
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
) and on its implications for the knowledge of jaw mechanics and feeding ecology of these hadrosaurs is published by Rivera-Sylva ''et al.'' (2019).
* A study on bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of ''Psittacosaurus lujiatunensis'' through its
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
is published by Zhao ''et al.'' (2019).
* A study on the morphological changes in the braincase of ''Psittacosaurus lujiatunensis'' through its ontogeny, based on data from three specimens from the Lower Cretaceous Yixian Formation (
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
), is published by Bullar ''et al.'' (2019).
* A three-dimensional virtual
endocast
An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
of a member of the genus ''Auroraceratops'' is reconstructed on the basis of a well-preserved skull by Zhang ''et al.'' (2019).
* Studies on the preservation of fossils of ''Auroraceratops rugosus'', on their stratigraphic provenance, and on the anatomy and phylogenetic relationships of this species are published by Suarez ''et al.'' (2019), Suarez ''et al.'' (2019), Morschhauser ''et al.'' (2019), Li ''et al.'' (2019), Morschhauser ''et al.'' (2019) and Morschhauser ''et al.'' (2019).
* A study on the nature of the observed variation in morphology and size of skulls of ''Bagaceratops rozhdestvenskyi'' is published online by Czepiński (2019), who considers the species ''Gobiceratops minutus'', ''Lamaceratops tereschenkoi'', ''Platyceratops tatarinovi'' and ''Magnirostris dodsoni'' to be Synonym (taxonomy), junior synonyms of ''B. rozhdestvenskyi''.
* The first postcranial skeleton of ''Bagaceratops'' reported so far is described from the Upper Cretaceous Barun Goyot Formation (
Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
) by Kim, Yun & Lee (2019).
* A study on the anatomy of the
appendicular skeleton
The appendicular skeleton is the portion of the vertebrate endoskeleton consisting of the bones, cartilages and ligaments that support the paired appendages ( fins, flippers or limbs). In most terrestrial vertebrates (except snakes, legless li ...
of ''
Protoceratops
''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
andrewsi'' and on its implications for the knowledge of the locomotor abilities of this species throughout its ontogeny is published by Słowiak, Tereshchenko & Fostowicz-Frelik (2019).
* A study on the bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
and skeletal growth of ''Avaceratops'' and ''Yehuecauhceratops'' is published online by Hedrick ''et al.'' (2019).
* New information on the anatomy of the skeleton of ''Pachyrhinosaurus perotorum'' is presented by Tykoski, Anthony Fiorillo, Fiorillo & Chiba (2019), who also provide a new diagnosis of this species.
* A study on the morphological variation of the skulls of specimens of ''Styracosaurus albertensis'' is published online by Holmes ''et al.'' (2019).
* A study on the affinities of two Chasmosaurinae, chasmosaurine skulls from the
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 7 ...
(
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
), previously referred to the species ''Chasmosaurus belli'', is published by Campbell ''et al.'' (2019), who transfer the species ''Vagaceratops irvinensis'' to the genus ''Chasmosaurus'', and consider the two studied skulls to be fossils of members of the genus ''Chasmosaurus'' of uncertain specific assignment, with morphology intermediate between ''C. belli'' and ''C. irvinensis''.
* A study on the
taphonomy
Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek language, Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientis ...
of hadrosaurid and ceratopsid fossils from the Scabby Butte locality (St. Mary River Formation;
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
) is published online by Campbell, Ryan & Anderson (2019).
*A study on the taxonomic status of ''Teyuwasu barberenai'', in which it was proposed as a second specimen of the Herrerasauridae, herrerasaurid ''Staurikosaurus pricei'' rather than a separate genus and species, is published by Garcia, Müller & Dias-da-Silva (2019).
*A study on a putative
sauropodomorph
Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...
ilium, from Carnian rocks of the Candelária Sequence/Santa Maria Formation (
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
), aims to discuss muscle attachment scars, in the context of
ontogeny
Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the ovum, egg to adult. The term can also be used to refer to t ...
and phylogeny of
basal Dinosauriformes, dinosauriforms, focusing in Saturnaliinae, saturnaliine sauropodomorphs is published by Garcia ''et al.'' (2019).
Birds
New bird taxa
Avialan research
* A study on early bird evolution, aiming to determine their divergence times and evolutionary rates, is published by Zhang & Wang (2019).
* A study on the impact of varying oxygen concentrations, global temperatures and air densities on the flight performance of extinct birds and on major diversification events which took place during the evolution of birds is published by Serrano ''et al.'' (2019).
* A study aiming to determine whether there is a relationship between the volume of Lacuna (histology), lacunae of osteocytes derived from static osteogenesis and biological parameters such as genome size, body mass, growth rate, metabolic rate or red blood cell size in extant birds is published by Grunmeier & D'Emic (2019), who evaluate the implications of their finding for inferring Physiology, physiological paraments in extinct birds, and potentially other vertebrates, on the basis of osteocyte lacunar volumes.
* A study on the expression patterns of the anterior genes ''GLI3, Gli
3'' and ''ALX4, Alx
4'' in limb buds of emu, chicken and zebra finch embryos, and on their implications for the knowledge of evolution of the Origin of birds#Digit homology, avian digital pattern, is published by Kawahata ''et al.'' (2019).
* A study on the diversity of regulatory gene expression profiles of amniote Digit (anatomy), digits, evaluating its implications for the knowledge of the origin of the avian digital pattern, is published by Stewart ''et al.'' (2019).
* A study on the total mass of the dentition of Mesozoic birds, and on the impact of the reduction and loss of teeth on total body mass of Mesozoic birds, is published by Zhou, Sullivan & Zhang (2019).
* A review of the available evidence of the diet of Mesozoic birds, especially those known from the Lower Cretaceous Jehol Lagerstätte (China), is published by O'Connor (2019).
* A study on the early evolution of the digestive system of birds, as indicated by data from Paraves, paravians from the Jurassic Yanliao Biota and the Cretaceous Jehol Biota (
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
), is published online by O'Connor & Zhou (2019).
* A study on the early evolution of the diel activity patterns in diapsid lineages, focusing on the common ancestor branch of living birds, is published by Yu & Wang (2019).
* A study on the diversity of melanosome
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
in Iridescence, iridescent feathers of extant birds, and on its implications for inferring iridescence in fossil feathers in general and in
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
birds cf. ''Primotrogon'' and ''Scaniacypselus'' in particular, is published by Nordén ''et al.'' (2019).
* A study on the morphology of melanosomes from feathers of extant birds that express non-iridescent structural colour, and on its implications for the possibility of detection of these melanosomes in the fossil record in general and in Crown group#Stem groups, stem group Coraciidae, roller ''Eocoracias'' in particular, is published by Babarović ''et al.'' (2019).
* Amino acids are detected in two specimens of fossil feathers from the Cretaceous amber from Myanmar and
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
Baltic amber by McCoy ''et al.'' (2019).
* Description of new amber specimens preserving feathers from the Cretaceous of Myanmar, evaluating the implications of these feathers for the knowledge of the development of the rachis-dominated feathers of Mesozoic birds, is published by Carroll, Luis M. Chiappe, Chiappe & Bottjer (2019).
* A study on ''Praeornis sharovi'' from the
Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
of Kazakhstan is published by Agnolin, Rozadilla & Carvalho (2019), who interpret the fossil as a tail feather of a
basal bird.
* A geochemical halo of the calamus of the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
feather of ''Archaeopteryx lithographica'', detected using Laser-induced fluorescence, Laser-Stimulated Fluorescence, is reported by Kaye ''et al.'' (2019), who also assess the implications of their findings for the identification of this feather; the study is subsequently criticized by Carney, Tischlinger & Shawkey (2020).
* A study on the postcranial skeleton of the Specimens of Archaeopteryx#The Berlin specimen, Berlin specimen of ''Archaeopteryx lithographica'', reporting pneumatic structures visible under ultraviolet light and confirming that numerous postcranial bones of ''Archaeopteryx'' were reduced in mass via hollow interiors, is published by Schwarz ''et al.'' (2019).
* A comparative study of all named taxa referred to Confuciusornithiformes, taxonomic revision of the group and a study on the phylogenetic relationships of members of the group is published by Wang, O'Connor & Zhou (2019).
* Evidence of beak preservation in a referred specimen of ''Confuciusornis sanctus'' (originally the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
of ''Jinzhouornis zhangjiyingia'') is presented by Falk ''et al.'' (2019).
* A study on bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of ''Confuciusornis sanctus'', and on its implications for the knowledge of the life history of this species, is published online by Anusuya Chinsamy-Turan, Chinsamy ''et al.'' (2019).
* Fully fledged feathering is reported in a hatchling Enantiornithes, enantiornithine specimen from the
Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
Las Hoyas locality in
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
(first described by Knoll ''et al.'', 2018) by Kate ''et al.'' (2019).
* A remarkably well-preserved foot of an enantiornithine bird, accompanied by part of the wing plumage, is described from the Cretaceous amber from Myanmar by Xing ''et al.'' (2019).
* A foot of a bird (likely a member of Enantiornithes; made the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
of the species ''Fortipesavis prehendens'' in a later publication), revealing a morphology previously unrecognized in Mesozoic birds, and a range of feathers representing multiple morphotypes are reported for the Cretaceous amber from Myanmar by Xing ''et al.'' (2019).
* O'Connor ''et al.'' (2019) describe the integument preserved in four juvenile enantiornithine specimens from the Early Cretaceous Jehol Biota, interpreted by the authors as mid to late immature feathers.
* Description of two new specimens of ''Protopteryx fengningensis'' from the Lower Cretaceous Huajiying Formation (China), preserving most of the skeleton and plumage, and providing new information on the anatomy and flight performance of members of this species, is published online by Luis M. Chiappe, Chiappe ''et al.'' (2019).
* A study on the bone microstructure of ''Yanornis'', and on its implications for the knowledge of the growth strategy of this bird, is published online by Wang ''et al.'' (2019).
* A study on the evolution and function of avian predentary found in Mesozoic Ornithuromorpha, ornithuromorphs, based on data from a specimen of ''Yanornis martini'', is published by Bailleul ''et al.'' (2019).
* A study comparing the hindlimb
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of Hesperornithiformes, hesperornithiforms and modern foot-propelled diving birds is published by Bell, Wu & Luis M. Chiappe, Chiappe (2019).
* A study on the paleobiogeography of hesperornithiforms, evaluating its implications for the knowledge of the paleoecology of the Late Cretaceous Western Interior Seaway, is published by Wilson (2019).
* A large bird
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
referred to the species ''Gargantuavis philoinos'', providing new information on the anatomy of this species, is described from the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
of southern
France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
by Buffetaut & Angst (2019), who name a new family Gargantuaviidae.
* Description of a well-preserved pelvis of ''Gargantuavis'' from the Maastrichtian Sânpetru Formation (
Romania
Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
), preserving characteristics previously unknown in ''Gargantuavis'' and constituting the first record of this genus outside the area of the Late Cretaceous Ibero-Armorican Island, is published online by Gerald Mayr, Mayr ''et al.'' (2019), who evaluate the implications of this finding for the knowledge of the phylogenetic relationships of ''Gargantuavis''; the study is subsequently criticized by Éric Buffetaut, Buffetaut & Angst (2020).
* A study on the evolution of body size of Palaeognathae, palaeognath birds is published by Crouch & Julia Clarke, Clarke (2019).
* A study on the hindlimb anatomy and phylogenetic relationships of ''Palaeotis weigelti'' is published by Gerald Mayr, Mayr (2019).
* A study on wing anatomy, body mass, wing surface area, wing span and probable flight parameters of ''Calciavis grandei'' is published online by Torres, Mark Norell, Norell & Clarke (2019).
* A study on changes of ostrich eggshell bead diameter throughout the Holocene, testing the proposed relationship between changes of ostrich eggshell bead diameter and the spread of herding in Africa, is published by Miller & Sawchuk (2019).
* A study on the taxonomic identification of Rhea (bird), rhea bones from four archaeological sites in the Mendoza Province (Argentina), based on genetic data, is published by Abbona ''et al.'' (2019).
* A study aiming to evaluate whether Invasive species in New Zealand, introduced deer and hares fill the same ecological niches as extinct moa birds in New Zealand, as indicated by data from pollen extracted from moa coprolites and mammal feces, is published by Wood & Wilmshurst (2019).
* A study on the anatomy of the Bone#Cancellous bone, cancellous bone in the
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
, tibiotarsus and fibula of three moa species is published by Bishop, Scofield & Hocknull (2019).
* A study on population densities and on the relationship between body mass and population densities in moa birds is published online by Latham ''et al.'' (2019).
* A
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
of a very large specimen of ''Struthio dmanisensis, "Struthio" dmanisensis'' is described from the Early Pleistocene of the Crimean Peninsula by Zelenkov ''et al.'' (2019), who transfer this species to the genus ''Pachystruthio'' and estimate body mass of this species.
* A study on the microstructure of the bones of ''Vegavis iaai'' is published by Garcia Marsà, Agnolín &
Novas
A nova ( novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", Latin for "new") that slowly fades over weeks or months. All observed novae involve white d ...
(2019).
* Eggshells and a small ovoid-shaped egg of Neognathae, neognathous birds, probably members of the family Presbyornithidae, as well as a carpometacarpus of a presbyornithid are described from the
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of the Glib Zegdou Formation (Algeria) by Garcia ''et al.'' (2019).
* A study on the putative Cariamiformes, cariamiform
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
from the
Maastrichtian
The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
Sandwich Bluff Member of the
Lopez de Bertodano Formation (Vega Island,
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
) reported by Case ''et al.'' (2006) is published by West ''et al.'' (2019), who reinterpret this specimen as a fossil of an unnamed large-bodied member of the genus ''Vegavis''.
* A study on the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen and other fossils attributed to the species ''Cayaoa bruneti'' is published by De Mendoza & Tambussi (2019), who present a revised diagnosis of this species.
* A study on the phylogenetic relationships of ''Cayaoa bruneti'' is published by De Mendoza (2019).
* A study on the Cenozoic fossil record of Anatidae, anatids from Eurasia is published by Zelenkov (2019).
* A study on the morphology of the postcranial skeleton of the
Oligocene
The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
-
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
Galliformes, galliform ''Palaeortyx'', and on the phylogenetic relationships of this taxon, is published by Zelenkov (2019).
* A study on the phylogenetic relationships of extant and fossil members of Strisores is published by Chen ''et al.'' (2019).
* Description of new fossil material of ''Pellornis mikkelseni'', providing new information on the anatomy of this species, and a study on the phylogenetic relationships of this species is published by Musser, Ksepka & Field (2019).
* A study on the phylogenetic relationships of the adzebills, as indicated by data from near-complete mitochondrial genome sequences, is published by Boast ''et al.'' (2019).
* A study on the phylogenetic relationships of the adzebills, as indicated by morphological and molecular data, is published by Musser & Joel Cracraft, Cracraft (2019).
* A study on two Humerus, humeri of Rail (bird), rails belonging to the genus ''Dryolimnas'' from the Pleistocene of the Picard Island (Seychelles) is published by Hume & Martill (2019), who interpret these humeri as bones of a flightless rail, and consider them to be evidence of repeated evolution flightlessness in members of the genus ''Dryolimnas'' inhabiting the Aldabra Atoll – before the atoll was completely submerged in the late Pleistocene, as well as after it emerged from the ocean again.
* A study on the phylogenetic relationships and evolutionary history of living and extinct flightless lineages of the white-throated rail from the Aldabra Group is published by van de Crommenacker ''et al.'' (2019).
* A revision of extinct endemic rails of the Mascarene Islands and a study on their ecology and extinction chronologies is published by Hume (2019).
* A study on the taxonomic status of the Canary Islands oystercatcher (''Haematopus meadewaldoi'') is published online by Senfeld ''et al.'' (2019).
* A study on the fossil material attributed to the species ''Becassius charadriioides'' is published online by De Pietri, Gerald Mayr, Mayr & Scofield (2019), who assign this species to the family Glareolidae.
* A nearly complete tarsometatarsus of the least seedsnipe (''Thinocorus rumicivorus'') is described from the Ensenadan of Argentina by Picasso, De Mendoza & Gelfo (2019).
* A study aiming to determine the drivers of the extinction of the great auk, based on data from mitochondrial genome sequences from across its geographic range, is published by Thomas ''et al.'' (2019).
* Pedal
phalanx
The phalanx (: phalanxes or phalanges) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar polearms tightly packed together. The term is particularly used t ...
of a penguin affected by osteomyelitis is described from the
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of West Antarctica by Jadwiszczak & Rothschild (2019).
* A set of skeletal elements of a penguin attributable to the species ''Delphinornis larseni'', providing new information on the anatomy of this species, is described from the Eocene Submeseta Formation (Seymour Island, Antarctica) by Jadwiszczak & Mörs (2019).
* The first skull reliably assigned to ''Anthropornis grandis'' is described from the Eocene (
Bartonian
The Bartonian is, in the International Commission on Stratigraphy's (ICS) geologic time scale, a stage or age in the middle of the Eocene Epoch or Series. The Bartonian Age spans the time between . It is preceded by the Lutetian and is follow ...
) Submeseta Formation (Seymour Island, Antarctica) by Acosta Hospitaleche ''et al.'' (2019).
* A study on the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Tereingaornis moisleyi'', evaluating the taxonomic validity of this species, is published online by Thomas ''et al.'' (2019).
* A fossil humerus of the Magellanic penguin or a relative of this species is described from
Uruguay
Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
by Acosta Hospitaleche ''et al.'' (2019), representing the first fossil of a penguin from Uruguay reported so far.
* A study on changes in the population size of the Adélie penguin colonies and relative krill abundance in the Prydz Bay (
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
) during the 2nd millennium, as indicated by data from ornithogenic sediment cores from the Vestfold Hills, will be published by Gao ''et al.'' (2019).
* A vertebra of a stork similar to the maguari stork is described from the late Pleistocene of the Santa Vitória Formation (Rio Grande do Sul,
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
) by Lopes, Pereira & Ferigolo (2019), who evaluate the implications of this finding for reconstructions of local paleoenvironment.
* Restudy of a putative bill of an ibis-like bird from the
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
La Meseta Formation (
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
) described by Jadwiszczak, Gaździcki & Tatur (2008) is published by Agnolin, Bogan & Rozadilla (2019), who consider this specimen to be more likely to be a dorsal spine of a Chimaera, chimaeroid Chondrichthyes, cartilaginous fish.
* A study on the body mass evolution in the clade Telluraves, incorporating data from 76 extinct species, is published by Crouch & Mason-Gamer (2019).
* A study on the demographic history of the Andean condors in southern South America and on the causes of their survival of late Quaternary megafauna extinctions is published by Perrig ''et al.'' (2019).
* Hindlimb bones of an extinct eagle of uncertain phylogenetic placement are described from the late Quaternary of Hispaniola by David Steadman, Steadman, Milan & Mychajliw (2019).
* A study on the origin and evolution of the Haast's eagle and the Eyles's harrier, as indicated by complete mitochondrial genome data, is published by Knapp ''et al.'' (2019).
* Evidence from Neanderthal-associated sites in Europe indicating that Neanderthals practiced catching the golden eagles is presented by Finlayson ''et al.'' (2019).
* Evidence of Châtelperronian Neanderthals using Pes (anatomy), pedal Phalanx bone, phalanges of imperial eagles for symbolic practices is reported from the Cova Foradà site (
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) by Rodríguez-Hidalgo ''et al.'' (2019).
* A study on the impact of the climate changes of the last 35,000 years on the long-eared owls and burrowing owls, as indicated by data from fossils from the La Brea Tar Pits, is published by Madan, Donald Prothero, Prothero & Syverson (2019).
* A study on the geographical origin and evolutionary history of Coraciiformes, based on data from extant taxa and from fossils, is published by McCullough ''et al.'' (2019).
* New skull remains of ''Phorusrhacos longissimus'' are described from the Cerro de los fósiles site in the
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
Santa Cruz Formation (Argentina) by Degrange ''et al.'' (2019).
* A study on the phylogenetic relationships of the Bahaman caracara, based on data from a nearly complete mitochondrial genome recovered from a bone of a member of this species, is published by Oswald ''et al.'' (2019).
* A study on the
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Calcardea junnei'' is published by Gerald Mayr, Mayr, Philip D. Gingerich, Gingerich & Smith (2019), who reject the interpretation of this species as a heron, and claim that this bird resembled parrot-like taxon ''Vastanavis'' from the early
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
.
* A study on the identity of a parakeet specimen held at National Museums Scotland, interpreted as most likely originating from Mauritius by Cheke & Jansen (2016), is published by Jones ''et al.'' (2019), who consider this parakeet to be the only known skin specimen of extinct Echo parakeet, Réunion parakeet.
* Complete genomic sequence of a specimen of the Carolina parakeet is generated by Gelabert ''et al.'' (2019), who evaluate the implications of their findings for the knowledge of the phylogenetic relationships of this species, its demographic history and adaptation to a toxic diet.
* A study on the phylogenetic relationships,
biogeography
Biogeography is the study of the species distribution, distribution of species and ecosystems in geography, geographic space and through evolutionary history of life, geological time. Organisms and biological community (ecology), communities o ...
and diversification rates of passerine birds throughout their evolutionary history, aiming to evaluate the impact of major events in Earth history on the evolution of passerines, is published by Oliveros ''et al.'' (2019).
* Dussex ''et al.'' (2019) sequence whole genomes of the huia and the South Island kōkako, South Island kokako, and evaluate whether the loss of genomic diversity played a role in their extinction.
* A review of Cretaceous and
Paleogene
The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
bird fossils from the James Ross Basin (
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
) is published by Acosta Hospitaleche ''et al.'' (2019).
* A study on drivers of bird distribution shifts throughout the Cenozoic is published by Saupe ''et al.'' (2019).
* A review of the bird fossil assemblage from the
Paleocene
The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
locality of Menat, Puy-de-Dôme, Menat (Puy-de-Dôme,
France
France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
), including a new fossil specimen with exceptional soft tissue preservation, is published by Gerald Mayr, Mayr, Hervet & Buffetaut (2019).
* New bird fossils, including the oldest European record of the Gastornithidae which is temporally well-constrained, are described from the Paleocene localities from the North Sea Basin in Belgium (Maret) and France (Templeuve and Rivecourt-Petit Pâtis) by Mayr & Smith (2019).
* A revision of bird fossils from the
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
(Ypresian) fossil sites of the North American Okanagan Highlands, mainly in British Columbia (
Canada
Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
), is published by Mayr ''et al.'' (2019), who report, among other findings, a skeleton of a possible member of the family Songziidae, and fossil wings which might constitute the earliest known record of Gaviiformes.
* An assemblage of 54 bird bones from early Eocene marine sediments of the Ampe quarry near Egem in Belgium is described by Mayr & Smith (2019).
* New Eocene bird fossils, including remains of members of Crown group#Pan-group, Pan-Charadriiformes, a member of Pan-Mirandornithes and a member or a relative of the family Quercymegapodiidae, are described from the Bumban Member of the Naranbulag Formation (
Mongolia
Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
) by Hood ''et al.'' (2019).
* Flamingo-like and Anatidae, anatid-like fossil bird footprints are described from the Vinchina Formation (Argentina) by Farina ''et al.'' (2019), who name new ichnotaxa ''Phoenicopterichnum lucioi'' and ''P. vinchinaensis''.
* A study on the date of extinction of the Tristan moorhen, the Inaccessible Island finch and the Tristan albatross on the main island of the Tristan da Cunha archipelago, aiming to place these extinctions in the context of the changing island ecosystems of the nineteenth and early twentieth centuries, is published by Bond, Carlson & Burgio (2019).
* Description of a fossil bird assemblage from the early Pliocene of the Na Burguesa-1 site (Mallorca,
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) is published by Torres-Roig ''et al.'' (2019).
* A study on the impact of Plio-Pleistocene environmental changes on the bird fauna of New Zealand is published by Rawlence ''et al.'' (2019).
* Description of Late Pleistocene and Holocene bird remains from Jerimalai (cave), Jerimalai and Matja Kuru 1 sites in East Timor will be published by Meijer, Louys & O'Connor (2019).
* Description of bird remains from the Castelcivita Caves, Grotta di Castelcivita site (
Italy
Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
) and a study on their implications for the knowledge of local environment and human-bird interactions in the Paleolithic is published by Fiore ''et al.'' (2019).
* Description of bird remains from the Qesem cave (Israel) dated to between 420 and 200 Year#SI prefix multipliers, ka, and a study on their implications for the knowledge of interactions of birds and humans occupying the site, is published by Blasco ''et al.'' (2019).
* A study on the phylogenetic relationships of the dodo and the great auk, as indicated by data from proteins extracted from bone material, is published by Horn ''et al.'' (2019).
* A study on bone surface modifications of Pleistocene bird fossils from Mata Menge site (Flores, Indonesia) is published by Meijer ''et al.'' (2019), who report no unambiguous evidence for exploitation of birds from Mata Menge by early hominins.
* A study on the impact of human colonization of New Zealand on the diversity dynamics of New Zealand bird fauna is published by Valente, Etienne & Garcia-R (2019).
Pterosaurs
New pterosaur taxa
Pterosaur research
* A study on the evolution of vertebral Skeletal pneumaticity, pneumaticity in pterosaurs is published by Buchmann & Rodrigues (2019).
* A study on the development of pterosaur embryos is published by Unwin & Deeming (2019).
* A study on three pterosaur coprolites from the
Upper Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the name ...
of Poland, probably produced by Ctenochasmatidae, ctenochasmatids, and on the probable diet of their producers is published by Qvarnström ''et al.'' (2019).
* Pterosaur#Pycnofibers, Pycnofibers showing diagnostic features of feathers are reported in two specimens of Anurognathidae, anurognathid pterosaurs (probably belonging either to the genus ''Jeholopterus'' or ''Dendrorhynchoides'') from the
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
of
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
by Yang ''et al.'' (2019); the study is subsequently criticized by Unwin & Martill (2020).
* Redescription of the holotype specimen of ''Mythunga camara'' is published by Pentland & Poropat (2019).
* Partial left Metacarpal bones, metacarpal of a large Ornithocheiridae, ornithocheirid pterosaur is described from the Lower Cretaceous (
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) Wessex Formation (
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
) by Martill & Coram (2019).
* New specimen of ''Sinopterus atavismus'' (a nearly complete skeleton) is described from the Lower Cretaceous (
Aptian
The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
)
Jiufotang Formation
The Jiufotang Formation () is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms (see Jehol Biota). It is a member of the Jehol group. T ...
(
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) by Zhang ''et al.'' (2019).
* A study on intervertebral foramina in ''Vectidraco'', ''Anhanguera (pterosaur), Anhanguera'' and ''Coloborhynchus'', and on their implications for inferring palaeoecology and locomotion of these pterosaurs, is published by Martin-Silverstone, Sykes & Darren Naish, Naish (2019).
* The first pterosaur postcranial bone (a left ulna) from the
Albian
The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
Lohan Cura Formation (Argentina) is described by Bellardini & Codorniú (2019).
* A study on the melanin content of the soft tissue headcrest of a specimen of ''Tupandactylus imperator'' is published by Pinheiro ''et al.'' (2019).
* Alexander Kellner, Kellner ''et al.'' (2019) describe fossils of Pterodactyloidea, pterodactyloid pterosaurs from the Upper Cretaceous Santa Marta Formation and
Snow Hill Island Formation
The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
(
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
).
Other archosaurs
Other new archosaur taxa
Other archosaur research
* Fossils of an immature Lagerpetidae, lagerpetid, representing a new morphotype different from other known lagerpetids, and co-occurring with dinosaurs and Silesauridae, silesaurids, are described from the Candelária Sequence of the
Upper Triassic
The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. T ...
Santa Maria Formation (
Brazil
Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
) by Garcia ''et al.'' (2019).
* A study on the
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
and gross
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
of a growth series of Femur, femora of ''Dromomeron romeri'' is published by Griffin ''et al.'' (2019).
* A study reevaluating the taxonomic validity of ''Lagosuchus talampayensis'' is published by Agnolin & Ezcurra (2019).
* A study on the microstructure of the long bones (
femur
The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg.
The Femo ...
and tibiae) of ''Lewisuchus admixtus'' is published by Garcia Marsà, Agnolín &
Novas
A nova ( novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", Latin for "new") that slowly fades over weeks or months. All observed novae involve white d ...
(2019).
* New specimen of ''Lewisuchus admixtus'' is described from the Chañares Formation (Argentina) by Ezcurra ''et al.'' (2019).
* A study on the anatomy of the skeleton of ''Asilisaurus kongwe'' is published online by
Nesbitt, Langer & Ezcurra (2019).
* A study on the anatomy of the braincase of ''Silesaurus opolensis'' is published by Piechowski, Niedźwiedzki & Tałanda (2019).
* Coprolites containing beetle remains, most likely produced by ''Silesaurus opolensis'', are described from the Upper Triassic Krasiejów locality (Poland) by Qvarnström ''et al.'' (2019).
* A study on bone
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
of ''Sacisaurus agudoensis'' is published by Veiga ''et al.'' (2019).
* A study on the phylogenetic relationships of ''Pisanosaurus mertii'' is published by Baron (2019), who interprets this taxon as a likely Silesauridae, silesaurid.
General research
* A study on patterns of evolutionary integration among regions of the archosaur skull, based on data from extant and fossil taxa, is published by Felice ''et al.'' (2019).
* A review of the Biogeography, biogeographic history of
crocodyliforms
Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
, Sauropoda, sauropod dinosaurs, nonavian Theropoda, theropod dinosaurs and mammals from the Mesozoic of Gondwana is published by David W. Krause, Krause ''et al.'' (2019).
* A study on the biogeography of Cretaceous terrestrial tetrapods, including terrestrial crocodyliforms, non-avian dinosaurs, birds and pterosaurs, is published by Kubo (2019).
* A study on size and shape differences between brains and
endocast
An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
s of extant American alligator and Chicken, domestic chicken, and on its implications for inferring whether endocasts are a reliable proxy for brain
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
in archosaurs in general, is published by Watanabe ''et al.'' (2019).
* A study comparing the mechanical properties of teeth of ''Suchomimus tenerensis'' and ''
Sarcosuchus
''Sarcosuchus'' (), from Ancient Greek σάρξ (''sárx''), meaning "flesh", and σούχος (''soúkhos''), meaning "crocodile", is an extinct genus of crocodyliform a that lived during the Early Cretaceous, from the late Hauterivian to the ...
imperator'' is published by Kundanati ''et al.'' (2019).
* A study on the distribution of medullary bone in the skeletons of living birds, aiming to refine the set of criteria used to evaluate purported records of medullary bone tissue in fossil avemetatarsalians, is published by Canoville,
Schweitzer Schweitzer is a surname. Notable people with the surname include:
* Albert Schweitzer (1875–1965), German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize
* Anton Schweitzer (1735–1787), German op ...
&
Zanno (2019).
* A study comparing the anatomy of hindlimbs of cursorial birds, non-avian theropod dinosaurs and other cursorial animals, aiming to determine whether cursorial birds are good kinematic model for reconstructions of theropod dinosaur locomotion, is published by Grossi ''et al.'' (2019).
* A study on the microstructure of eggshells in birds and non-avian maniraptoran dinosaurs is published by Choi, Han & Lee (2019).
* Hu ''et al.'' (2019) reconstruct the vomer of ''Sapeornis'' and ''Sinovenator'', and evaluate their implications for the knowledge of the evolution of the skull of Paraves, paravians.
* A study on the anatomy of skull fenestrae in Sauropsida, sauropsids, and on its implications for reconstructions of dinosaur soft tissues, is published online by Holliday ''et al.'' (2019).
* A study aiming to determine the likely karyotype of the dinosaur and the early diapsid ancestor of birds is published by Griffin, Larkin & O'Connor (2019).
* A study on the evolution of bipedality in archosaurs is published by Grinham, VanBuren & David B. Norman, Norman (2019).
* A study on the evolution of the brain of bird-line archosaurs is published by Beyrand ''et al.'' (2019).
* A review of the progress in the field of archosaur paleo
histology
Histology,
also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at large ...
, focusing in particular on the study of the dinosaurs, is published by Bailleul, O'Connor &
Schweitzer Schweitzer is a surname. Notable people with the surname include:
* Albert Schweitzer (1875–1965), German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize
* Anton Schweitzer (1735–1787), German op ...
(2019).
* A study on the phylogenetic distribution of the hyposphene-hypantrum articulation in the vertebrae of archosaurs is published by Stefanic &
Nesbitt (2019).
* A study comparing growth patterns of the American alligator, chicken and ''Tenontosaurus tilletti'' is published by Brunner ''et al.'' (2019).
* A study comparing the position, size and number of Skeletal pneumaticity, pneumatic foramina in the vertebrae of pterosaurs and birds is published by Buchmann, Avilla & Rodrigues (2019).
* A study on non-avian dinosaur and bird tracks (representing some of the oldest known bird tracks) preserved in slabs used as building stones at the Chengde Mountain Resort, originating from the Tuchengzi Formation (
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
) and dating to the Jurassic-Cretaceous boundary, is published online by Xing ''et al.'' (2019).
* Description of non-avian dinosaur and bird tracks from the Upper Cretaceous Chignik Formation (southwestern Alaska), evaluating their implications for the knowledge of habitat preferences of northern high-latitude dinosaurs, is published by Fiorillo ''et al.'' (2019).
* An assemblage of non-avian dinosaur and bird feathers is described from the Lower Cretaceous Koonwarra, Victoria#Koonwarra fossil bed, Koonwarra Fossil Bed (
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
) by Kundrát ''et al.'' (2019).
* A study on Feather#Classification, barb angles in birds and non-avian dinosaurs, evaluating their implications for the knowledge of feather shape evolution and the utility of barb angles for determination of flight abilities of fossil taxa, is published by Wang, Tang & Julia Clarke, Clarke (2019).
References
{{portal bar, Dinosaurs, Evolutionary biology, Paleontology
2019 in paleontology
2010s in paleontology
2019 in science