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Mesosuchia
"Mesosuchia" is an obsolete name for a group of terrestrial, semi-aquatic, or fully aquatic crocodylomorph reptiles. Characteristics The marine crocodile '' Metriorhynchus'' had paddle-like forelimbs, '' Dakosaurus andiniensis'' had a skull that was adapted to eat large sea reptiles, and '' Shamosuchus'' was adapted for eating molluscs and gastropods. ''Shamosuchus'' also looked like modern crocodiles and was very closely related to their direct ancestor. History The "Mesosuchia" were formerly placed at Suborder rank as within Crocodylia. The "first" crocodiles were placed within their own suborder, Protosuchia; whilst extant species where placed within Suborder Eusuchia (meaning 'true crocodiles'). Mesosuchia were the crocodylians "in between".But it is no longer regarded as genuine because it belongs to a paraphyletic group. It is replaced by its phylogenetic equivalent Mesoeucrocodylia, which contains the taxa herein, the Crocodylia, and some allied forms more recently discover ...
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Mesoeucrocodylia
Mesoeucrocodylia is the clade that includes Eusuchia and crocodyliforms formerly placed in the paraphyletic group Mesosuchia. The group appeared during the Early Jurassic, and continues to the present day. Diagnosis It was long known that Mesosuchia was an evolutionary grade, a hypothesis confirmed by the phylogenetic analysis of Benton and Clark (1988) which demonstrated that Eusuchia (which includes all living crocodylian species) was nested within Mesosuchia. Due to the paraphyly of Mesosuchia, Mesoeucrocodylia was erected to replace Mesosuchia. Several anatomical characteristics differentiate Mesoeucrocodylia from the other crocodylomorph clades. The frontal bones of the skull are fused together into a single compound element, for example. Mesoeucrocodylians possess something of a secondary palate, formed by the posterior extension of sutured palatine bones. The otic aperture of the members of this clade is blocked posteriorly by the squamosal bone. Classification Phy ...
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Crocodylomorpha
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. Extinct crocodylomorphs were considerably more ecologically diverse than modern crocodillians. The earliest and most primitive crocodylomorphs are represented by " sphenosuchians", a paraphyletic assemblage containing small-bodied, slender forms with elongated limbs that walked upright, which represents the ancestral morphology of Crocodylomorpha. These forms persisted until the end of the Jurassic. During the Jurassic, crocodylomorphs morphologically diversified into numerous niches, with the subgroups Neosuchia (which includes modern crocodilians) and the extinct Thalattosuchia adapting to aquatic life, while some terrestrial groups adopted herbivorous and omnivorous lifestyles. Terrestrial crocodylomorphs would continue to co-exist alongside aquatic forms until becoming extinct d ...
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Thalattosuchia
Thalattosuchia is a clade of mostly marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia and records from Thailand and China suggest that some members lived in freshwater. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Teleosauroids are not greatly specialised for oceanic life, with back osteoderms similar to other crocodyliformes. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin, and probably gave Viviparity, live birth, seemingly uniquely among archosaurs. Taxonomy The term Thalattosuchia was coined by Fraas in 1901.Fraas E. 1901. Die Meerkrokodile (Thalattosuchia n. g.) eine neue Sauriergruppe der Juraform ...
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Notosuchia
Notosuchia is a clade of primarily Gondwanan mesoeucrocodylian Crocodylomorpha, crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogeny, phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group (see #Phylogeny, below); if Sebecosuchia is included within Notosuchia its existence is pushed into the Late Miocene or Zanclean, Early Pliocene, about 7 to 4.5 million years ago. Fossils have been found from South America, Africa, Asia, and Europe. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory (''Chimaerasuchus''), omnivory (''Simosuchus''), and terrestrial hypercarnivory (''Baurusuchus''). It included many members with highly derived traits unusual for crocodylomorphs, including Mammal, mammal-like teeth, flexible bands of shield-like body armor similar to those of armadillos (''Armadillosuchus''), and possibly fleshy cheeks and Pig, pig-like snouts (''Notosuchus''). Th ...
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Dyrosauridae
Dyrosauridae is a family of extinct neosuchian crocodyliforms that lived from the Campanian to the Eocene. Dyrosaurid fossils are globally distributed, having been found in Africa, Asia, Europe, North America and South America. Over a dozen species are currently known, varying greatly in overall size and cranial shape. A majority were aquatic, some terrestrial and others fully marine (see locomotion below), with species inhabiting both freshwater and marine environments. Ocean-dwelling dyrosaurids were among the few marine reptiles to survive the Cretaceous–Paleogene extinction event. The dyrosaurids were a group of mostly marine, long jawed, crocodile-like quadrupeds up to long. The largest dyrosaurid was probably '' Phosphatosaurus'' estimated at in length. Based on bone tissue evidence, it has been hypothesized that they were slow-growing near-shore marine animals with interlocking closed jaws, able to swim as well as walk on land. External nostrils at the posterior end ...
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Metriorhynchus
''Metriorhynchus'' is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, ''M. brevirostris'' was named in 1829 as a species of ''Steneosaurus'' before being named as a separate genus by the German palaeontologist Christian von Meyer in 1832. The name ''Metriorhynchus'' means "moderate snout", and is derived from the Greek ''Metrio''- ("moderate") and -''rhynchos'' ("snout"). Discovery and species Fossil specimens referrable to ''Metriorhynchus'' are known from Kimmeridgian (Late Jurassic) deposits of France. Valid species Only one valid species is recognized today, the type species ''M. geoffroyii'' (now called ''M. brevirostris''). ''"Metriorhynchus" hastifer'' and ''"M." palpebrosus'' are generically distinct from the ''Metriorhynchus'' type species, with ''hastifer'' being recovered as a geosaurine. Species in this genus were traditionally classed into two skull groups: longirostrine (long, narrow jaws) and brevirost ...
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Teleosauridae
Teleosauridae is a family of extinct typically marine crocodylomorphs similar to the modern gharial that lived during the Jurassic period. Teleosaurids were thalattosuchians closely related to the fully aquatic metriorhynchoids, but were less adapted to an open-ocean, pelagic lifestyle. The family was originally coined to include all the semi-aquatic (i.e. non-metriorhynchoid) thalattosuchians and was equivalent to the modern superfamily Teleosauroidea. However, as teleosauroid relationships and diversity was better studied in the 21st century, the division of teleosauroids into two distinct evolutionary lineages led to the establishment of Teleosauridae as a more restrictive family within the group, together with its sister family Machimosauridae. Amongst teleosauroids, teleosaurids were generally smaller and less common than machimosaurids, suggesting the two families occupied different niches, similar to modern species of crocodilians. However, teleosaurids were more div ...
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Sarcosuchus
''Sarcosuchus'' (), from Ancient Greek σάρξ (''sárx''), meaning "flesh", and σούχος (''soúkhos''), meaning "crocodile", is an extinct genus of crocodyliform a that lived during the Early Cretaceous, from the late Hauterivian to the early Albian stages, 130 to 112 million years ago of what is now Africa and South America. It was one of the largest pseudosuchians, with the largest specimen of ''S. imperator'' reaching approximately long and weighing up to . It is known from two species; ''S. imperator'' from the early Albian Elrhaz Formation of Niger, and ''S. hartti'' from the Late Hauterivian of northeastern Brazil. Other material is known from Morocco and Tunisia and possibly Libya and Mali. The first remains were discovered during several expeditions led by the French paleontologist Albert-Félix de Lapparent, spanning from 1946 to 1959, in the Sahara. These remains were fragments of the skull, vertebrae, teeth, and scutes. In 1964, an almost complete skull was fo ...
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Pholidosauridae
Pholidosauridae is an extinct family of aquatic neosuchian mesoeucrocodylian crocodylomorphs. Fossils have been found in Europe (Denmark, England, France, Germany, Spain and Sweden), Africa (Algeria, Niger, Mali, Morocco and Tunisia), North America (Canada and the United States) and South America (Brazil and Uruguay). The pholidosaurids first appeared in the fossil record during the Bathonian stage of the Middle Jurassic. Jouve & Jalil (2020) described postcranial material of a pholidosaurid from the Paleocene (Danian) of Ouled Abdoun Basin (Morocco), representing the most recent record of the family. The authors also reinterpreted putative Maastrichtian dyrosaurid '' Sabinosuchus'' as a pholidosaurid, and argued that at least two independent pholidosaurid lineages reached the Maastrichtian, among which one survived the Cretaceous–Paleogene extinction event. Before the publication of this study it was thought that the family became extinct during the Late Turonian stage of the L ...
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Trematochampsidae
Trematochampsidae is an extinct family of mesoeucrocodylian crocodyliforms. Fossils are present from Madagascar, Morocco, Niger, Argentina, and Brazil (in the case of '' Caririsuchus'', where some specimens have been found in the Romualdo Member of the Santana Formation). Possible trematochampsids have been found from Spain and France, but classification past the family level is indeterminant. The trematochampsids first appeared during the Barremian stage of the Early Cretaceous and became extinct during the late Maastrichtian stage of the Late Cretaceous. Description Trematochampsids are deep-snouted and have a ziphodont tooth structure. The dentition differs from most other crocodilians in that the teeth are recurved, serrated, and lateromedially compressed. This may be an adaptation to a terrestrial or at least semiterrestrial lifestyle as such teeth would be better suited for cutting and tearing into prey as opposed to capturing them and holding them underwater. Despite this ...
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Atoposauridae
Atoposauridae is an extinct family of neosuchian crocodyliforms, known from the Jurassic and Cretaceous of Eurasia. Characterised by their small size and their heterodont (having multiple tooth types) dentition, they are thought to have lived in terrestrial and semi-aquatic environments, and to have consumed small prey such as fish, insects and mammals. The oldest records of the group are known from the Middle Jurassic of Britain. The majority of the family are known from Late Jurassic to Early Cretaceous deposits in France, Portugal, and Bavaria in southern Germany. The discovery of the genus '' Aprosuchus'', however, extends the duration of the lineage to the end of the Cretaceous in Romania. Classification Atoposaurids are universally considered members of Neosuchia. Tennant et al. (2016) in their revision of the group, found atoposaurids to be basal neosuchians, and the group as traditionally defined to be paraphyletic with respect to Paralligatoridae, restricting Atoposaurid ...
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Elosuchidae
''Elosuchus'' is an extinct genus of neosuchian crocodyliform that lived during the Middle Cretaceous of what is now Africa (Niger, Morocco and Algeria). Description and taxonomy ''Elosuchus'' had an elongated snout like a gharial and was probably a fully aquatic animal. The type species, ''E. cherifiensis'' from Algeria and Morocco, was originally described as a species of ''Thoracosaurus'' by Lavocat, but was recognized as a genus separate from ''Thoracosaurus'' by de Broin in 2002. ''Elosuchus felixi'', described from the Echkar Formation of Niger, was renamed '' Fortignathus'' in 2016 and is either a dyrosaurid relative or a non-hyposaurine dyrosaurid.de Broin, F. de L., 2002, ''Elosuchus'', a new genus of crocodile from the Lower Cretaceous of the North of Africa: C. R. Palevol, v. 1, p. 275-285.Mark T. Young; Alexander K. Hastings; Ronan Allain; Thomas J. Smith (2016). "Revision of the enigmatic crocodyliform Elosuchus felixi de Lapparent de Broin, 2002 from the Lower–Upp ...
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