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Teleosauridae is a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
typically marine crocodylomorphs similar to the modern
gharial The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family (biology), family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males ...
that lived during the
Jurassic period The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. The Jurassic constitutes the second and m ...
. Teleosaurids were thalattosuchians closely related to the fully aquatic metriorhynchoids, but were less adapted to an open-ocean,
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
lifestyle. The family was originally coined to include all the semi-aquatic (i.e. non-metriorhynchoid) thalattosuchians and was equivalent to the modern superfamily Teleosauroidea. However, as teleosauroid relationships and diversity was better studied in the 21st century, the division of teleosauroids into two distinct evolutionary lineages led to the establishment of Teleosauridae as a more restrictive family within the group, together with its sister family Machimosauridae. Amongst teleosauroids, teleosaurids were generally smaller and less common than machimosaurids, suggesting the two families occupied different niches, similar to modern species of
crocodilians Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchi ...
. However, teleosaurids were more diverse than machimosaurids, with generalist coastal predators ('' Mystriosaurus''), long-snouted marine
piscivores A piscivore () is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted repti ...
('' Bathysuchus''), and potentially even long-snouted, semi-terrestrial predators ('' Teleosaurus''). Additionally, teleosaurids occupied a wider range of habitats than machimosaurids, from semi-marine coasts and estuaries, the open-ocean, freshwater, and potentially even semi-terrestrial environments.


Classification

Teleosauridae is
phylogenetically In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
defined in the ''
PhyloCode The ''International Code of Phylogenetic Nomenclature'', known as the ''PhyloCode'' for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the ...
'' by Mark T. Young and colleagues as "the largest
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
within Teleosauroidea containing '' Teleosaurus cadomensis'' but not '' Machimosaurus hugii''. Teleosauridae is split into two subfamilies, the Teleosaurinae and the Aeolodontinae, the former defined in the ''PhyloCode'' as "the largest clade within Teleosauroidea containing ''Teleosaurus cadomensis'', but not '' Aeolodon priscus'' and the latter defined in the ''PhyloCode'' as "the largest clade within Teleosauroidea containing ''Aeolodon pricus'', but not ''Teleosaurus cadomensis''.


Palaeobiology

Teleosaurids were originally regarded as marine analogues to modern gharials, as they both typically share long, tubular snouts and narrow teeth. However, differences in the jaws, teeth, and skeleton of different teleosaurids suggest that they were more ecologically diverse than this. Earlier teleosaurids were coastal semi-aquatic generalists, while the two subfamilies were more specialised. Teleosaurines appear to have been semi-terrestrial, as they were more heavily armoured and had forward-facing nostrils. In contrast, aeolodontines have been found in deep marine waters and had reduced armour, implying that they were open water predators similar to metriorhynchoids (although the oldest aeolodontine, '' Mycterosuchus'', appears to have been semi-terrestrial, similar to teleosaurines).


Palaeoecology


Distribution

Definitive fossils of teleosaurids are restricted to Laurasia, with material found in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
(
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,
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,
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,
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
,
Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
,
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
and
Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
) and Asia (
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
and
Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
, and possibly
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
).


See also

* List of marine reptiles


References

{{Authority control Thalattosuchia Pseudosuchian families Jurassic crocodylomorphs Prehistoric marine crocodylomorphs Toarcian first appearances Hauterivian extinctions