Istiodactyliform
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Istiodactyliform
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs have been discovered worldwide except Antarctica, though most genera have been recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by pteranodontians and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not ''Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also regarded to have some of the largest pterosaur wingspans, such as the one estimated for t ...
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Mimodactylidae
Mimodactylidae is a group or family of pterosaurs known from Cretaceous deposits in East Asia and the Middle East. They are phylogenetically more closely related to the Istiodactylidae than to the Ornithocheiridae or the Anhangueridae, forming the clade Istiodactyliformes together with the Istiodactylidae and ''Hongshanopterus''.Kellner, Alexander W. A.; Caldwell, Michael W.; Holgado, Borja; Vecchia, Fabio M. Dalla; Nohra, Roy; Sayão, Juliana M.; Currie, Philip J. (2019). "First complete pterosaur from the Afro-Arabian continent: insight into pterodactyloid diversity". ''Scientific Reports''. 9(1). . In 2019, Alexander Wilhelm Armin Kellner, Michael Wayne Caldwell, Borja Holgado, Fabio Marco Dalla Vecchia, Roy Nohra, Juliana Manso Sayão and Philip John Currie defined the clade Mimodactylidae as the group consisting of ''Mimodactylus libanensis'' and all species more closely related to ''Mimodactylus'' than to ''Istiodactylus latidens'', '' Ikrandraco avatar'' or '' Anhang ...
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Istiodactylidae
Istiodactylidae is a small Family (biology), family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus ''Ornithodesmus''. Systematics and distribution Remains of taxa that can be confidently assigned to Istiodactylidae have been found in the UK and China, in rocks dating from the Early Cretaceous period (Barremian to Aptian stage). Victoria Arbour, Arbour and Currie (2011) described Canadian ''Gwawinapterus beardi'' as a member of Istiodactylidae living in the late Cretaceous (upper Campanian stage); however, Witton (2012) suggested the tooth replacement pattern in this animal does not match that of pterosaurs, suggesting that the species might be non-pterosaurian. Additional research suggested that the species was in fact a fish. The earliest known species might be ''Archaeoistiodactylus linglongtaensis'', from the Middle Jurassic of China; however, it also has been suggested that the holotype speci ...
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Hongshanopterus
''Hongshanopterus'' is a genus of pterodactyloid pterosaur from the Lower Cretaceous Jiufotang Formation of Liaoning, China. The type species ''Hongshanopterus lacustris'' was in 2008 named and described by Wang Xiaolin, Diogenes de Almeida Campos, Zhou Zhonghe and Alexander Wilhelm Armin Kellner. The generic name combines a reference to the Hongshan culture in Lioaning with a Latinised Greek ''pteron'', "wing". The specific name means "of the lake" in Latin, referring to the lake deposits the fossil was found in, at Dapinfang. ''Hongshanopterus'' is based on the holotype IVPP V14582, found in a layer of the Jiufotang Formation dating from the Aptian, It consists of a skull and five neck vertebrae of a single subadult individual. The wingspan of ''Hongshanopterus'' was estimated at 1.8 to 1.9 meters. It possessed a relatively high number of teeth, about thirty-six for both upper jaws combined. The teeth were robust and had triangular crowns which were flattened from mouth ...
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Linlongopterus
''Linlongopterus'' is a genus of pteranodontoidea, pteranodontoid pterodactyloidea, pterodactyloid pterosaur from the Early Cretaceous of China. It is known from a partial skull and mandible first named and described in 2015 by Rodrigues ''et al.''. The only known specimen was found in the Jiufotang Formation of the Liaoning Province or China, and lived around 120 million years ago. The full binomial of the taxon is ''Linlongopterus jennyae''. Discovery and naming The genus ''Linlongopterus'' was described and named in a 2015 paper by Taissa Rodrigues, Shunxing Jiang, Xin Cheng, Xiaolin Wang, and Alexander Kellner in ''Historical Biology'', containing the single species ''L. jennyae''. The species is based on a singular specimen, IVPP V15549, stored at the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing, China. It was discovered in the rocks of the Jiufotang Formation in Jianchang County, Liaoning province, and is split into two blocks of material which neede ...
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Valanginian
In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 137.05 ± 0.2 Ma and 132.6 ± 0.2 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretaceous and precedes the Hauterivian Stage of the Lower Cretaceous. Stratigraphic definitions The Valanginian was first described and named by Édouard Desor in 1853. It is named after Valangin, a small town north of Neuchâtel in the Jura Mountains of Switzerland. The base of the Valanginian is at the first appearance of calpionellid species '' Calpionellites darderi'' in the stratigraphic column. A global reference section (a GSSP) had in 2009 not yet been appointed. The top of the Valanginian (the base of the Hauterivian) is at the first appearance of the ammonite genus '' Acanthodiscus''. Subdivision The Valanginian is often subdivided in Lower and Upper substages. The Upper substage begins at the first appearance of ammonite speci ...
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Pterodactyloidea
Pterodactyloidea ( ; derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like '' Nyctosaurus'' and '' Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. " Pterodactyl" is also a common term for pterodactyloid pte ...
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Pterosaurs
Pterosaurs are an extinct clade of flying reptiles in the Order (biology), order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved flying and gliding animals, powered flight. Their wings were formed by a membrane of skin, muscle, and other tissue (biology), tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were ...
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Suborder
Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow consistent ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of '' Tiktaalik'' in the arctic of Canada. Paleontology includes the study of fossils: their age, method of formation, and evolutionary significance. Specimens are sometimes considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years to 4.1 billion years old. Early edition, published online before prin ...
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Targaryendraconia
Targaryendraconia is an extinct clade of lanceodontian pterosaurs that lived from the Early to Late Cretaceous period in Europe, North America, South America, and Australia. Classification Below is a cladogram following a topology by Pêgas and colleagues in 2019. In their analysis, they recovered Targaryendraconia as the sister taxon of the clade Anhangueria, both of which are within the more inclusive group Ornithocheirae. Targaryendraconia is split into two families: the Targaryendraconidae, which contains '' Aussiedraco'', '' Barbosania'', and '' Targaryendraco'', and the Cimoliopteridae, which contains '' Aetodactylus'', '' Camposipterus'', and '' Cimoliopterus''.Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, Contrastingly, a study by Alexander Averianov ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian–Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Onto ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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