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Flying Insects
Pterygota ( ) is a subclass of insects that includes all winged insects and groups who lost them secondarily. Pterygota group comprises 99.9% of all insects. The orders not included are the Archaeognatha (jumping bristletails) and the Zygentoma (silverfishes and firebrats), two primitively wingless insect orders. Unlike Archaeognatha and Zygentoma, the pterygotes do not have styli or vesicles on their abdomen (also absent in some zygentomans), and with the exception of the majority of mayflies, are also missing the median terminal filament which is present in the ancestrally wingless insects. The oldest known representatives of the group appeared during the mid-Carboniferous, around 328–324 million years ago, and the group subsequently underwent rapid diversification. Claims that they originated substantially earlier during the Silurian or Devonian based on molecular clock estimates are unlikely based on the fossil record, and are likely analytical artefacts. __TOC__ Syste ...
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Empis Livida
''Empis livida'' is a species of fly in the Empididae, Empididae family. It is included in the subgenus ''Kritempis'' of the genus ''Empis''. Males range from , females . The male's abdomen is brownish and its wings appear faintly brown and clouded. The female's abdomen is gray and its wings are clear. ''E. livida'' lives in hedgerows, feeding on the nectar of several species of Rosaceae, several species of Asteraceae, and ''Heracleum sphondylium'' nectar; they also feed on other insects. They live all across temperate and Northern Europe, the only species with such a wide distribution. ''E. livida'' larvae are also carnivorous and live in damp soil and leaf litter. Adults fly in between April and July. References

Empis Asilomorph flies of Europe Flies described in 1758 Taxa named by Carl Linnaeus {{Empidoidea-stub ...
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Paraneoptera
Paraneoptera or Acercaria is a Order (biology), superorder of insects which includes Psocodea, lice (Psocoptera, bark lice and Louse, true lice), thrips, and hemipterans, the true bugs. It also includes the extinct order Permopsocida, known from fossils dating from the Cisuralian, Early Permian to the mid-Cretaceous. All of the insects classified here exhibit various “reductions” or “simplifications” from the primitive body-plan found in typical polyneopterans. Cercus, Cerci, for example, are entirely absent in all living paraneopterans (Acercaria meaning without cerci). Other “reductions” occur in wing venation, in the number of tarsal segments (no more than three), only four Malpighian tubules, and only one complex of abdominal ganglia. The mouthparts of the Paraneoptera reflect diverse feeding habits. Basal groups are microbial surface feeders, whereas more advanced groups feed on plant or animal fluids. Phylogeny Paraneoptera consists of Psocodea (lice), along ...
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Paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor exc ...
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Paleoptera
The name Palaeoptera (from Greek ( 'old') + ( 'wing')) has been traditionally applied to those ancestral groups of winged insects (most of them extinct) that lacked the ability to fold the wings back over the abdomen as characterizes the Neoptera. The Diaphanopterodea, which are palaeopteran insects, had independently and uniquely evolved a different wing-folding mechanism. Both mayflies and dragonflies lack any of the smell centers in their brain found in Neoptera. Disputed status The complexities of the wing-folding mechanism, as well as the mechanical operation of the wings in flight ( indirect flight muscles), are such that it indicates that Neoptera is a monophyletic lineage. The plesiomorphic absence of wing-folding does not necessarily mean the Palaeoptera form a natural group – they may be an assemblage containing all insects, closely related or not, that "are not Neoptera", an example of a wastebasket taxon. If the extinct lineages are taken into account, it is ...
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Infraclass
In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of organisms (a ''taxon'') in a hierarchy that reflects evolutionary relationships. Thus, the most inclusive clades (such as Eukarya and Animalia) have the highest ranks, whereas the least inclusive ones (such as ''Homo sapiens'' or ''Bufo bufo'') have the lowest ranks. Ranks can be either relative and be denoted by an indented taxonomy in which the level of indentation reflects the rank, or absolute, in which various terms, such as species, genus, family, order, class, phylum, kingdom, and domain designate rank. This page emphasizes absolute ranks and the rank-based codes (the Zoological Code, the Botanical Code, the Code for Cultivated Plants, the Prokaryotic Code, and thCode for Viruses require them. However, absolute ranks are not req ...
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Molecular Clock
The molecular clock is a figurative term for a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged. The biomolecular data used for such calculations are usually nucleotide sequences for DNA, RNA, or amino acid sequences for proteins. Early discovery and genetic equidistance The notion of the existence of a so-called "molecular clock" was first attributed to Émile Zuckerkandl and Linus Pauling who, in 1962, noticed that the number of amino acid differences in hemoglobin between different lineages changes roughly linearly with time, as estimated from fossil evidence. They generalized this observation to assert that the rate of evolutionary change of any specified protein was approximately constant over time and over different lineages (known as the molecular clock hypothesis). The genetic equidistance phenomenon was first noted in 1963 by Emanuel Margoliash, who wrote: "It appears that the number ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Firebrat
The firebrat (''Thermobia domestica'') is a small insect (typically 1–1.5 cm) in the order Zygentoma. Habitat Firebrats prefer relatively warm temperatures (36–39 °C) and require some humidity. They are commonly found indoors near heat sources such as furnaces and boilers. They feed on a wide variety of carbohydrates and starches that are also protein sources such as dog food, flour and book bindings. They are distributed throughout most parts of the world and are normally found outdoors under rocks, plant litter, and in similar environments, but are also often found indoors where they are considered pests. They do not cause major damage, but they can contaminate food, damage paper goods, and stain clothing. Otherwise they are mostly harmless. Behavior Firebrats utilize pheromones to lead other firebrats to attract one another and congregate. To maintain a group, firebrats must remain in contact with one another. Breeding At 1.5 to 4.5 months of age t ...
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Silverfish
The silverfish (''Lepisma saccharinum'') is a species of small, primitive, wingless insect in the order Zygentoma (formerly Thysanura). Its common name derives from the insect's silvery light grey colour, combined with the fish-like appearance of its movements. The scientific name (''L. saccharinum'') indicates that the silverfish's diet consists of carbohydrates such as sugar or starches. While the common name ''silverfish'' is used throughout the global literature to refer to various species of Zygentoma, the Entomological Society of America restricts use of the term solely for ''Lepisma saccharinum''. Description The silverfish is a nocturnal insect typically long. Its abdomen tapers at the end, giving it a fish-like appearance. The newly hatched are whitish, but develop a greyish hue and metallic sheen as they get older. It has two long cerci and one terminal filament at the tip of the abdomen between the cerci. It also has two small compound eyes, although other members ...
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Zygentoma
Zygentoma are an order in the class Insecta, and consist of about 550 known species. The Zygentoma include the so-called silverfish or fishmoths, and the firebrats. A conspicuous feature of the order are the three long caudal filaments. The two lateral filaments are cerci, and the medial one is an epiproct or ''appendix dorsalis''. In this they resemble the Archaeognatha, although the cerci of Zygentoma, unlike in the latter order, are nearly as long as the epiproct. Until the late twentieth century the Zygentoma were regarded as a suborder of the Thysanura, until it was recognized that the order Thysanura was paraphyletic, thus raising the two suborders to the status of independent monophyletic orders, with Archaeognatha as sister group to the Dicondylia, including the Zygentoma. Etymology The name "Zygentoma" is derived from the Greek ('), in context meaning "yoke" or "bridge"; and ('), "insects" (literally meaning "cut into", in reference to the segmented anatomy o ...
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Archaeognatha
The Archaeognatha are an order of apterygotes, known by various common names such as jumping bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the same time as the arachnids. Specimens that closely resemble extant species have been found as both body and trace fossils (the latter including body imprints and trackways) in strata from the remainder of the Paleozoic Era and more recent periods. For historical reasons an alternative name for the order is Microcoryphia. Until the late 20th century the suborders Zygentoma and Archaeognatha comprised the order Thysanura; both orders possess three-pronged tails comprising two lateral cerci and a medial epiproct or ''appendix dorsalis''. Of the three organs, the appendix dorsalis is considerably longer than the two cerci; in this the Archaeognatha differ from the Zygentoma, in which the three organs are subequal in length. In the late 20th ...
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