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Varanidae
The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely related to ''Varanus'' than to the earless monitor lizard (''Lanthanotus''). ''Varanus'' includes the Komodo dragon (the largest living lizard), crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and Chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia. Taxonomy The Varanidae were defined (using morphological characteristics) by Estes, de Queiroz and Gauthier (1988) as the clade containing the most recent common ancestor of '' Lanthanotus'' and ''Varanus'' and all of its descendants. A similar definition was formulated by Conrad ' ...
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Varanoidea
Varanoidea is a superfamily of lizards, including the well-known family Varanidae (the monitors and goannas). Also included in the Varanoidea are the Lanthanotidae ( earless monitor lizards), and the extinct Palaeovaranidae. Throughout their long evolutionary history, varanoids have exhibited great diversity, both in habitat and form. This superfamily includes the largest-known terrestrial lizard, Megalania (~5 meters), and the largest extant lizard, the Komodo dragon (''Varanus komodoensis'', ~3 meters). Evolution Either synonymous with, or a subgroup of, the group Platynota, the varanoids first appear in the fossil record in the latter part of Early Cretaceous, but possible varanoid ancestors have been traced back as far as Early Jurassic times. Among the earliest known varanoids are the monitor-like necrosaurids '' Palaeosaniwa canadensis'' from the Campanian (roughly 71–82 mya) of North America and '' Estesia mongoliensis'' and '' Telmasaurus grangeri'', both from the ...
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Saniwa
''Saniwa'' is an extinct genus of varanid lizard that lived during the Eocene epoch. It is known from well-preserved fossils found in the Bridger and Green River Formations of Wyoming, United States. The type species ''S. ensidens'' was described in 1870 as the first fossil lizard known from North America. A second species, ''S.'' ''orsmaelensis'', is recognised from remains found in Europe. It is a close relative of '' Varanus'', the genus that includes monitor lizards. Description ''Saniwa'' measured . Like other varanid lizards, ''Saniwa'' had a long, pointed snout and nostrils placed farther back in the skull than most lizards and a tail that was almost twice as long as the body. Although similar in appearance to extant monitor lizards, ''Saniwa'' had many primitive traits, including teeth on its palate, a jugal bone beneath the eye that extended farther forward, and a suture between the frontal and parietal bones that was straight rather than curved. A study in 2018 by ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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Lizard
Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic islands, oceanic Archipelago, island chains. The grouping is Paraphyly, paraphyletic as some lizards are more closely related to snakes than they are to other lizards. Lizards range in size from chameleons and geckos a few centimeters long to the 3-meter-long Komodo dragon. Most lizards are quadrupedal, running with a strong side-to-side motion. Some lineages (known as "legless lizards") have secondarily lost their legs, and have long snake-like bodies. Some lizards, such as the forest-dwelling ''Draco (genus), Draco'', are able to glide. They are often Territory (animal), territorial, the males fighting off other males and signalling, often with bright colours, to attract mates and to intimidate rivals. Lizards are mainly carnivorous, often b ...
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Jacques Gauthier
Jacques Armand Gauthier (born June 7, 1948, in New York City) is an American vertebrate paleontologist, comparative morphologist, and systematist, and one of the founders of the use of cladistics in biology. Life and career Gauthier is the son of Edward Paul Gauthier and Patricia Marie Grogan. He received a B.S. degree in zoology at San Diego State University in 1973, a master's in biological science at the same institute in 1980, and a PhD in paleontology from the University of California, Berkeley, in 1984. Currently he is a professor of geology and geophysics and ecology and evolutionary biology and curator of vertebrate paleontology and vertebrate zoology at Yale University. His master's thesis, the content of which was published in 1982, is a classic work on the paleontology and phylogeny of the lizard clade Anguimorpha that remains a core reference for morphological research on Xenosauridae and Anguidae in particular. His PhD thesis constituted the first major cladi ...
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Kevin De Queiroz
Kevin de Queiroz is a vertebrate, Evolutionary biology, evolutionary, and Systematics, systematic biologist. He has worked in the phylogenetics and evolutionary biology of squamate reptiles, the development of a unified species concept and of a Phylogenetic nomenclature, phylogenetic approach to biological nomenclature, and the philosophy of systematic biology. Early life and career De Queiroz was born and raised in Los Angeles, California. He received a B.S. in Biology from the University of California, Los Angeles (1978), a M.S. in Zoology from San Diego State University (1985), and a Ph.D. in Zoology from the University of California, Berkeley (1989). He was a Tilton Postdoctoral Fellow at the California Academy of Sciences and is currently a Research Zoologist and a curator of the collection of Amphibians and Reptiles at the National Museum of Natural History, Smithsonian Institution. He is a former president of the Society of Systematic Biologists and was the first presid ...
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Chinese Crocodile Lizard
The Chinese crocodile lizard (''Shinisaurus crocodilurus'') is a semiaquatic anguimorph lizard found only in cool forests in southeastern China and northeastern Vietnam. The Chinese crocodile lizard spends much of its time in shallow water or in overhanging branches and vegetation, where it hunts its prey of insects, snails, tadpoles, and worms. Individuals in captivity may be fed baby mice.Chinese Crocodile Lizard (''Shinisaurus crocodilurus'').
The Sacramento Zoological Society.
A rare and little-studied lizard, it is listed in Appendix II, which regulates international trade of specimens. This is the only species in the

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Goanna
A goanna is any one of several species of lizard of the genus ''Monitor lizard, Varanus'' found in Australia and Southeast Asia. Around 70 species of ''Varanus'' are known, 25 of which are found in Australia. This varied group of carnivorous reptiles ranges greatly in size and fills several ecological niches. The goanna features prominently in Dreamtime, Aboriginal mythology and Australian folklore. Being predatory lizards, goannas are often quite large with sharp teeth and claws. The largest is the perentie (''V. giganteus''), which can grow over in length. Not all goannas are so large; pygmy goannas may be smaller than the arm of an adult human. The smallest of these, the short-tailed monitor (''V. brevicauda''), reaches only in length. They survive on smaller prey, such as insects and mice. Goannas combine predatory and scavenging behaviours. They prey on any animal they can catch that is small enough to eat whole. They have been blamed by farmers for the death of sheep, ...
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Savannah Monitor
The savannah monitor (''Varanus exanthematicus'') is a medium-sized species of monitor lizard native to Africa. The species is known as Bosc's monitor in Europe, since French scientist Louis Bosc first described the species. It belongs to the subgenus '' Polydaedalus''. Etymology The specific name ''exanthematicus'' is derived from the Greek word ''exanthem'' , meaning an eruption or blister of the skin. French botanist and zoologist Louis Augustin Guillaume Bosc originally described this lizard as ''Lacerta exanthematica'' in reference to the large oval scales on the back of its neck. The species was formerly known as ''Lacerta exanthematicus''. Description Savannah monitors are stoutly built, with relatively short limbs and toes, and skulls and dentition adapted to feed on hard-shelled prey. They are robust creatures, with powerful limbs for digging, powerful jaws and blunt, peglike teeth. Maximum size is rarely more than 100 cm. The skin coloration pattern varies accor ...
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Crocodile Monitor
The crocodile monitor (''Varanus salvadorii''), also known as the Papuan monitor or Salvadori's monitor, is a species of monitor lizard Endemism, endemic to New Guinea. It is the largest monitor lizard in New Guinea and is one of the List of largest extant lizards, longest lizards, verified at up to . Its tail is exceptionally long, with some specimens having been claimed to exceed the length of the Komodo dragon, however less massive. Crocodile monitors are arboreal lizard with a dark green body marked with bands of yellowish spots. It has a characteristic blunt snout and a very long prehensile tail. It lives among the mangrove swamps and coastal rainforests of the southeastern part of New Guinea, feeding opportunistically on everything from birds and small mammals to eggs, other reptiles, amphibians and carrion. Its large, backwards-curving teeth are better adapted than those of most monitors for seizing fast-moving prey. Like all monitors, it has anatomical features that enabl ...
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Earless Monitor Lizard
The earless monitor lizard (''Lanthanotus borneensis'') is a semiaquatic, brown lizard native to the Southeast Asian island of Borneo. It is the only living species in the family Lanthanotidae and it is related to the true monitor lizards. Taxonomy The earless monitor lizard was described in 1878 by Franz Steindachner. The genus name ''Lanthanotus'' means "hidden ear" and the species name ''borneensis'' refers to its home island of Borneo. The uniqueness of the species was immediately recognized and Steindachner placed it in its own family, Lanthanotidae. In 1899, George Albert Boulenger relegated it to the family Helodermatidae, together with the beaded lizards and Gila monster, on the basis of morphological similarities. Further studies were conducted in the 1950s where it was found that although it is related to Helodermatidae, this relationship is relatively distant. The similarity is in part the result of convergent evolution and they should be recognized as separate famil ...
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Genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. Phylogeneti ...
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