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A goanna is any one of several
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of lizard of the genus '' Varanus'' found in Australia and Southeast Asia. Around 70 species of ''Varanus'' are known, 25 of which are found in Australia. This varied group of
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s ranges greatly in size and fills several
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of Resource (biology), resources an ...
s. The goanna features prominently in
Aboriginal mythology Australian Aboriginal religion and mythology is the sacred spirituality represented in the stories performed by Aboriginal Australians within each of the Aboriginal Australian languages, language groups across Australia in their Aboriginal c ...
and
Australian folklore Australian folklore refers to the folklore and urban legends that have evolved in Australia from Aboriginal Australian myths to colonial and Contemporary literature, contemporary folklore including Australians, people, places and events, that ha ...
. Being
predatory Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
lizards, goannas are often quite large with sharp teeth and claws. The largest is the
perentie The perentie (''Varanus giganteus'') is a species of monitor lizard. It is one of the largest living lizards on earth, after the Komodo dragon, Asian water monitor, and the crocodile monitor. Found west of the Great Dividing Range in the arid ar ...
(''V. giganteus''), which can grow over in length. Not all goannas are so large; pygmy goannas may be smaller than the arm of an adult human. The smallest of these, the
short-tailed monitor The short-tailed pygmy monitor (''Varanus brevicauda'') is the second smallest living monitor lizard in the world with a maximum length of 25 cm. Distribution The short-tailed pygmy monitor ranges throughout central Australia, from the coas ...
(''V. brevicauda''), reaches only in length. They survive on smaller prey, such as insects and mice. Goannas combine predatory and scavenging behaviours. They prey on any animal they can catch that is small enough to eat whole. They have been blamed by farmers for the death of
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
, though most likely erroneously, as goannas are also eaters of
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
and are attracted to rotting meat. Most goannas are dark-coloured, with greys, browns, blacks, and greens featuring prominently; however, white is also common. Many desert-dwelling species also feature yellow-red tones.
Camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
ranges from bands and stripes to splotches, speckles, and circles, and can change as the creature matures, with juveniles sometimes being brighter than adults. Like most lizards, goannas lay
eggs An egg is an organic vessel in which an embryo begins to develop. Egg, EGG or eggs may also refer to: Biology * Egg cell, the female reproductive cell (gamete) in oogamous organisms Food * Eggs as food Places * Egg, Austria * Egg, Switzerland ...
. Most lay eggs in a nest or burrow, but some species lay their eggs inside
termite Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...
mounds. This offers protection and incubation; additionally, the termites may provide a meal for the young as they hatch. Unlike some other species of lizards, goannas do not have the ability to regrow limbs or tails.


Etymology

The name ''goanna'' is derived from ''iguana''. Early European bush settlers in Australia likened goannas to the South American lizards. Over time, the initial vowel sound was dropped. A similar explanation is used to link
possum Possum may refer to: Animals * Didelphimorphia, or (o)possums, an order of marsupials native to the Americas ** Didelphis, a genus of marsupials within Didelphimorphia *** Common opossum, native to Central and South America *** Virginia opossum ...
s to the American
opossum Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North A ...
. The South African term for a monitor lizard is ''
leguaan The Nile monitor (''Varanus niloticus'') is a large member of the Monitor lizard, monitor family (Varanidae) found throughout most of Sub-Saharan Africa, particularly in drier regions, and along the Nile River and its tributaries in East Africa. ...
'' and this word is also from Spanish 'la iguana'.


Species

For a list of all monitor lizards of the genus, see Complete list of genus ''Varanus''. The following are found in Australia. For the most part, in common names, "goanna" and "monitor" are interchangeable. *
Perentie The perentie (''Varanus giganteus'') is a species of monitor lizard. It is one of the largest living lizards on earth, after the Komodo dragon, Asian water monitor, and the crocodile monitor. Found west of the Great Dividing Range in the arid ar ...
– ''V. giganteus'' * Lace monitor – ''V. varius'' *
Sand goanna The sand goanna (''Varanus gouldii)'', also known commonly as Gould's monitor, the racehorse goanna, and the sand monitor, is a species of large Australian monitor lizard in the family Varanidae. Taxonomy John Edward Gray described the specie ...
– ''V. gouldii'' (also Gould's goanna or ground goanna) *
Mertens' water monitor Mertens' water monitor (''Varanus mertensi''), often misspelled Mertin's water monitor, is a species of monitor lizard. The species is Endemism, endemic to northern Australia, and is a wide-ranging, actively foraging, opportunistic predator of aq ...
– ''V. mertensi'' *
Spiny-tailed monitor The spiny-tailed monitor (''Varanus acanthurus''), also known as the Australian spiny-tailed monitor, the ridge-tailed monitor the Ackie dwarf monitor, and Colloquialism, colloquially simply ackie monitor, is an Australia, Australian species of li ...
– ''V. acanthurus'' (also ridge-tailed monitor) *
Mangrove monitor The mangrove monitor, mangrove goanna, or Western Pacific monitor lizard (''Varanus indicus'') is a member of the monitor lizard family with a large distribution from northern Australia and New Guinea to the Moluccas and Solomon Islands. It grows ...
– ''V. semiremex'' *
Black-headed monitor The black-headed monitor or black-tailed monitor (''Varanus tristis'') is a relatively small species of monitor lizard Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Afric ...
– ''V. tristis'' *
Short-tailed monitor The short-tailed pygmy monitor (''Varanus brevicauda'') is the second smallest living monitor lizard in the world with a maximum length of 25 cm. Distribution The short-tailed pygmy monitor ranges throughout central Australia, from the coas ...
– ''V. brevicauda'' * Argus monitor – ''V. panoptes'' (also yellow-spotted monitor) *
Rosenberg's monitor The Rosenberg's monitor (''Varanus rosenbergi'') is an Australian species of varanid reptile found in southern regions of the continent. They are large and fast predators with rugged bodies and long tails, having a combined length up to 1.5 metr ...
– ''V. rosenbergi'' * Spencer's goanna – ''V. spenceri'' *
Storr's monitor There are two species of lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as ...
– ''V. storri'' *
Dampier Peninsula monitor The Dampier Peninsula monitor or Dampier Peninsula goanna (''Varanus sparnus''), described in 2014, is the smallest known species of monitor lizard, growing up to 16.3 grams with a length of almost 23 cm and a SVL (snout to vent length) of 1 ...
– ''V. sparnus'' * Mitchell's water monitor – ''V. mitchelli'' * Kings' monitor – ''V. kingorum'' * Southern Pilbara rock goanna – ''V. hamersleyensis'' *
Black-palmed rock monitor The black-palmed rock monitor (''Varanus glebopalma'') is a member of the Varanidae family found in Australia. Also known as the twilight monitor or the long-tailed rock monitor, it is a member of the subgenus ''Odatria'', and is found in the nor ...
– ''V. glebopalma'' *
Kimberley rock monitor The Kimberley rock monitor (''Varanus glauerti)'' is a medium-sized species of monitor lizard
– ''V. glauerti'' * Pygmy mulga monitor – ''V. gilleni'' * Rusty desert monitor – ''V. eremius'' * Stripe-tailed goanna – ''V. caudolineatus'' * Pilbara monitor – ''V. bushi'' * Black-spotted ridge-tailed monitor – ''V. baritji'' *
Emerald tree monitor The emerald tree monitor (''Varanus prasinus'') or green tree monitor, is a small to medium-sized arboreal monitor lizard. It is known for its unusual coloration, which consists of shades from green to turquoise, topped with dark, transverse do ...
– ''V. prasinus'' *
Canopy goanna ''Varanus keithhornei'', commonly known as the canopy goanna, Keith Horne's monitor, blue-nosed tree monitor, or Nesbit River monitor, is a species of monitor lizards native to northeast Australia. It is a member of the '' Varanus prasinus'' sp ...
– ''V. keithhornei''


Habitats

Goannas are found throughout most of Australia, except for
Tasmania Tasmania (; palawa kani: ''Lutruwita'') is an island States and territories of Australia, state of Australia. It is located to the south of the Mainland Australia, Australian mainland, and is separated from it by the Bass Strait. The sta ...
, and manage to persist in a variety of environments. Most species are known to climb trees or outcrops; several primarily
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
species are known. The lace monitor (''V. varius'') is probably the best-known among these, but is not the most common. The lace monitor is the second-largest of all goannas, reaching lengths up to . Other more common tree goannas, such as the Timor tree monitor (''V. timorensis'') and mournful tree monitor (''V. tristis'',) do not grow to quite such lengths, typically a maximum of 61 cm, nose-to-tail. Other goannas are adapted to swampy coastal environments, such as the
mangrove goanna The mangrove monitor, mangrove goanna, or Western Pacific monitor lizard (''Varanus indicus'') is a member of the monitor lizard family with a large distribution from northern Australia and New Guinea to the Moluccas and Solomon Islands. It grows ...
(''V. semiremex''). Further still,
Mertens' water monitor Mertens' water monitor (''Varanus mertensi''), often misspelled Mertin's water monitor, is a species of monitor lizard. The species is Endemism, endemic to northern Australia, and is a wide-ranging, actively foraging, opportunistic predator of aq ...
(water goanna – ''V. mertensi''), found in lagoons and rivers across northern Australia, is streamlined for swimming, using its tail as a paddle. Most other goannas are good swimmers, but tend not to voluntarily venture into water.


Diet

The diets of goannas vary greatly depending on the species and the habitat. Prey can include all manner of small animals: insects, smaller lizards, snakes, mammals, birds, and eggs. Meals are often eaten whole, thus the size of their meals may depend on the size of the animals. Many of the small species feed mostly on insects, with some being small lizard experts. Many of the medium to large species feed on whatever prey they can catch. This includes eggs, fish (''V. mertensi''), birds, snails, smaller lizards, snakes, marsupials, and other small mammals, such as rodents. The giant perentie has been observed killing a young
kangaroo Kangaroos are marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use, the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern gre ...
, and then biting out chunks of flesh like a dog. All species are carrion eaters, so feed on the carcasses of dead animals, including livestock and other large creatures. The smell of rotting meat also attracts these lizards.


Goannas and humans


Confrontations

Like most native fauna, goannas are rather wary of human intrusions into their habitat, and most likely run away (into the scrub, up a tree, or into the water, depending on the species). A goanna is a rather swift mover, and when pressed, sprints short distances on its hind legs. Goannas also rear up when threatened, either chased or cornered, and also inflate flaps of skin around their throats and emit harsh hissing noises. Some goannas lose their initial fear of humans, especially when food is involved (or has been previously involved). The wildlife authority recommends not feeding animals while in their territory. An attack can cause serious injury in exceptional cases, but most authorities doubt that a goanna will direct an intentional attack at a human unless the human has attempted to attack it (or grasp at it) first.
Indigenous Australians Indigenous Australians are people with familial heritage from, or recognised membership of, the various ethnic groups living within the territory of contemporary Australia prior to History of Australia (1788–1850), British colonisation. The ...
who hunt goannas for food consider the perentie to be a high-risk (but tasty) quarry. Debate is growing as to whether goannas are venomous. The incessant bleeding caused by goanna bites had been thought to be the result of
bacterial infection Pathogenic bacteria are bacteria that can cause disease. This article focuses on the bacteria that are pathogenic to humans. Most species of bacteria are harmless and many are beneficial but others can cause infectious diseases. The number of t ...
, but a 2005 study suggested monitor lizards (including goannas) are venomous and have oral toxin-producing glands. The goanna's hefty tail can be dangerous when swung, much like a crocodile's tail; small children and dogs have been knocked down by such attacks. Often victims in goanna attacks are bystanders, watching the person antagonising the goanna. Alarmed goannas can mistake standing humans for trees and attempt to climb them to safety, which is painful and can be distressing for both human and goanna.


Conservation status

Goannas are protected species throughout Australia.


Culture and folklore

Goannas have a prominent place in the culture of indigenous Australians. This includes totemic relationships, anthropomorphic representations in
dreamtime The Dreaming, also referred to as Dreamtime, is a term devised by early anthropologists to refer to a religio-cultural worldview attributed to Australian Aboriginal religion and mythology, Australian Aboriginal mythology. It was originally u ...
stories, and as a food source. Representations of goannas are common in indigenous artwork, not just as food, but also as a symbolic spiritual motif. Smaller goannas and the mighty perentie are often considered two different animals when appearing in aboriginal works, as in the story "How the Goanna and Perentie Got Their Colours". European settlers perpetuated several
old wives' tale An "old wives' tale" is a colloquial expression referring to spurious or superstitious claims. They can be said sometimes to be a type of urban legend, said to be passed down by older women to a younger generation. Such tales are considered super ...
s about goanna habits and abilities; some of these have persisted in modern folklore among campers and bushmen. This includes the above-mentioned exaggeration of goannas dragging off sheep from shepherds' flocks in the night. This might even be exaggerated into child-snatching, rivalling drop bears (attack
koala The koala (''Phascolarctos cinereus''), sometimes inaccurately called the koala bear, is an arboreal herbivorous marsupial native to Australia. It is the only Extant taxon, extant representative of the Family (biology), family ''Phascolar ...
s) as a tourist scarer, but probably more convincing due to the reptiles' carnivorous nature and fearsome appearance. A common tale was that the bite of a goanna was infused with a powerful, incurable
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
. Every year after the bite (or every seven years), the wound would flare up again. For many years, herpetologists generally believed goannas were nonvenomous, and lingering illness from their bites was due solely to
infection An infection is the invasion of tissue (biology), tissues by pathogens, their multiplication, and the reaction of host (biology), host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmis ...
and
septicaemia Sepsis is a potentially life-threatening condition that arises when the body's response to infection causes injury to its own tissues and organs. This initial stage of sepsis is followed by suppression of the immune system. Common signs and s ...
as a result of their saliva being rife with
bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...
from carrion and other food sources. However, in 2005, researchers at the
University of Melbourne The University of Melbourne (colloquially known as Melbourne University) is a public university, public research university located in Melbourne, Australia. Founded in 1853, it is Australia's second oldest university and the oldest in the state ...
announced that oral venom glands had been found in both goannas and iguanas.Goanna venom rocks the reptile record
, UniNews Vol. 14, No. 22 28 November - 12 December 2005, University of Melbourne, Retrieved 8 March 2006 Because the goanna regularly eats snakes (which may involve a fierce struggle), including venomous species, they are often said to be immune to
snake venom Snake venom is a highly toxic saliva containing zootoxins that facilitates in the immobilization and digestion of prey. This also provides defense against threats. Snake venom is usually injected by unique fangs during a Snakebite, bite, though ...
. However, no evidence found suggests an actual venom
immunity Immunity may refer to: Medicine * Immunity (medical), resistance of an organism to infection or disease * ''Immunity'' (journal), a scientific journal published by Cell Press Biology * Immune system Engineering * Radiofrequence immunity ...
. Other stories say that the lizard eats a legendary plant, or drinks from a healing
spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a he ...
, which neutralises the venom. (This idea is immortalised in
Banjo Paterson Andrew Barton "Banjo" Paterson, (17 February 18645 February 1941) was an Australian bush poet, journalist and author, widely considered one of the greatest writers of Australia's colonial period. Born in rural New South Wales, Paterson worke ...
's humorous poem "Johnson's Antidote".) Goanna fat or oil has been anecdotally imbued with mystical healing properties (possibly in connection with their supposed venom immunity). Aboriginal people traditionally used goanna oil as an important
bush medicine Bush medicine comprises traditional medicines used by Indigenous Australians, being Aboriginal and Torres Strait Islander people. Indigenous people have been using various components of native Australian flora and some fauna as medicine for t ...
, and it also became a common medicine among Caucasians shortly after British settlement in Australia. Said to be a cure-all, and possessing amazing powers of penetration (passing through metal as if it were not there), it was sold among early settlers like
snake oil Snake oil is a term used to describe False advertising, deceptive marketing, health care fraud, or a scam. Similarly, snake oil salesman is a common label used to describe someone who sells, promotes, or is a general proponent of some valueless ...
in the
Old West The American frontier, also known as the Old West, and popularly known as the Wild West, encompasses the geography, history, folklore, and culture associated with the forward wave of American expansion in mainland North America that bega ...
of North America. A goanna features as the heroic figure Mr Lizard in the Australian author
May Gibbs Cecilia May Gibbs Order of the British Empire, MBE (17 January 1877 – 27 November 1969) was an Australian children's author, illustrator, and cartoonist. She is best known for her gumnut babies (also known as "bush babies" or "bush fairies"), ...
’ children's books ''
Snugglepot and Cuddlepie ''Snugglepot and Cuddlepie'' is a series of books written by Australians, Australian author May Gibbs. The books chronicle the adventures of the eponymous Snugglepot and Cuddlepie. The central story arc concerns Snugglepot and Cuddlepie (who ar ...
''. A bronze statue of the goanna Mr Lizard has been placed outside the State Library of Victoria. The villain in the Disney film ''
The Rescuers Down Under ''The Rescuers Down Under'' is a 1990 American animated adventure film produced by Walt Disney Feature Animation and released by Walt Disney Pictures. It is the sequel to ''The Rescuers'' (1977). In ''The Rescuers Down Under'', Bernard and Bianc ...
'', Percival C. McLeach, has a pet goanna named Joanna.


References


Further reading

* Cogger, H. (1967). ''Australian Reptiles in Colour''. Sydney: A. H. & A. W. Reed, * King, Dennis & Green, Brian. 1999. ''Goannas: The Biology of Varanid Lizards''. University of New South Wales Press. * Underhill, D. (1993). ''Australia's Dangerous Creatures''. Sydney: Reader's Digest.


External links


Johnson's Antidote
by
Banjo Paterson Andrew Barton "Banjo" Paterson, (17 February 18645 February 1941) was an Australian bush poet, journalist and author, widely considered one of the greatest writers of Australia's colonial period. Born in rural New South Wales, Paterson worke ...
.
A Run-In With An Australian Desert Lizard
(
NPR National Public Radio (NPR) is an American public broadcasting organization headquartered in Washington, D.C., with its NPR West headquarters in Culver City, California. It serves as a national Radio syndication, syndicator to a network of more ...
)
Waitress removes goanna from restaurant
(includes video) {{Varanoidea Monitor lizards of Australia Bushfood Bush medicine Australian Aboriginal bushcraft Reptile common names