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Tyrannosaurini
Tyrannosaurinae (or tyrannosaurines) is one of the two extinct subfamilies of Tyrannosauridae, a family of coelurosaurian theropods that consists of at least three tribes and several genera. All fossils of these genera have been found in the Late Cretaceous deposits of western North America and east Asia. Compared to the related subfamily Albertosaurinae, tyrannosaurines overall are more robust and larger though the alioramins were gracile by comparison. This subfamily also includes '' Lythronax'', one of the oldest known tyrannosaurid genera, as well as the youngest and most famous member of the group, ''Tyrannosaurus rex''. History of discovery The first remains of tyrannosaurids were uncovered during expeditions led by the Geological Survey of Canada, which located numerous scattered teeth. These distinctive dinosaur teeth were given the name '' Deinodon'' ("terrible tooth") by Joseph Leidy in 1856. In 1892 Edward Drinker Cope described more tyrannosaur material in the form o ...
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Daspletosaurus
''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 77 and 74.4 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three named species. Fossils of the earlier type species, ''D. torosus'', have been found in Alberta, while fossils of a later species, ''D. horneri'', have been found only in Montana. ''D. wilsoni'' has been suggested as an intermediate species between ''D. torosus'' and ''D. horneri'' that evolved through anagenesis, though further research may be required to definitively support the theory. There are also multiple specimens which may represent new species of ''Daspletosaurus'' from Alberta and Montana, but these have not been formally described. The taxon '' Thanatotheristes'' has been suggested to represent a species of ''Daspletosaurus'', ''D. degrootorum'', but this has not been widely supported. ''Daspletosaurus'' is closely related to the much ...
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Tyrannosaurus Rex
''Tyrannosaurus'' () is a genus of large theropoda, theropod dinosaur. The type species ''Tyrannosaurus rex'' ( meaning 'king' in Latin), often shortened to ''T. rex'' or colloquially t-rex, is one of the best represented theropods. It lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other Tyrannosauridae, tyrannosaurids. Fossils are found in a variety of geological formations dating to the latest Campanian-Maastrichtian Age (geology), ages of the late Cretaceous Period (geology), period, 72.7 to 66 mya (unit), million years ago, with isolated specimens possibly indicating an earlier origin in the middle Campanian. It was the last known member of the tyrannosaurids and among the last non-Bird, avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced ...
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Tyrannosaurus
''Tyrannosaurus'' () is a genus of large theropod dinosaur. The type species ''Tyrannosaurus rex'' ( meaning 'king' in Latin), often shortened to ''T. rex'' or colloquially t-rex, is one of the best represented theropods. It lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other tyrannosaurids. Fossils are found in a variety of geological formations dating to the latest Campanian-Maastrichtian ages of the late Cretaceous period, 72.7 to 66 million years ago, with isolated specimens possibly indicating an earlier origin in the middle Campanian. It was the last known member of the tyrannosaurids and among the last non- avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, ...
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Asiatyrannus
''Asiatyrannus'' (meaning "Asian tyrant") is an extinct genus of tyrannosaurine theropod dinosaurs from the Late Cretaceous Nanxiong Formation of China. The genus contains a single species, ''A. xui'', known from a single specimen consisting of a skull and partial postcranial skeleton. ''Asiatyrannus'' is notable for its deep-snouted skull and small body size, in contrast to the gracile snout and larger size of the contemporary ''Qianzhousaurus''. It represents the southernmost record of an Asian tyrannosaurid. Discovery and naming The ''Asiatyrannus'' holotype specimen, ZMNH M30360, was discovered in September 2017 in sediments of the Nanxiong Formation near Shahe Town in Nankang District of Ganzhou City, Jiangxi Province, China. The specimen consists of most of an Joint, articulated skull in addition to disarticulated parts of the postcrania, comprising much of the right and left legs and several caudal vertebrae. In 2024, Zheng et al. Species description, described ''A ...
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Alioramini
Alioramini is a clade of long-snouted tyrannosaurine tyrannosaurid dinosaurs from the Late Cretaceous epoch. It includes the genera '' Alioramus'' and '' Qianzhousaurus''. Although tyrannosaurids are known from a variety of places around the globe, alioramins are restricted to Asia in mostly Maastrichtian strata. Many of the fossils attributed to Alioramini are not from fully developed individuals. Description Alioramins are medium-sized tyrannosaurids, reaching around in length. They have a more gracile body plan as compared to most other tyrannosaurines. Alioramins have rather shallow snouts, a trait that is rather rare among tyrannosaurs but can be found in the early tyrannosauroid, '' Xiongguanlong''. Alioramins are unique when compared to contemporary tyrannosaurs from the same time, such as '' Tarbosaurus'' and ''Tyrannosaurus'', because most of the longer snouted tyrannosauroids, such as ''Xiongguanlong,'' were found in deposits dating to earlier times during the Cretace ...
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Bistahieversor
''Bistahieversor'' (meaning "Bistahi destroyer"), also known as the "Bisti Beast", is a genus of basal Eutyrannosauria, eutyrannosaurian Theropoda, theropod dinosaur. The genus contains only a single known species, ''B. sealeyi'', described in 2010, from the Late Cretaceous of New Mexico. The holotype and a juvenile were found in the Hunter Wash Member of the Kirtland Formation, while other specimens came from the underlying Fossil Forest member of the Fruitland Formation. This dates ''Bistahieversor'' approximately 75.5 to 74.5 million years ago during the Campanian age, found in sediments spanning a million years. Discovery and naming The first remains now attributed to ''Bistahieversor'', a partial skull and skeleton, were described in 1990 as a specimen of ''Aublysodon''. Additional remains, consisting of the incomplete skull and skeleton of a juvenile, were described in 1992. Another complete skull and partial skeleton were found in the Bisti/De-Na-Zin Wilderness, Bisti/De- ...
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Labocania
''Labocania'' is a genus of carnivorous theropod dinosaur of disputed affinities from the Late Cretaceous of Mexico. Initially regarded as an indeterminate theropod, later studies mentioned possible affinities with tyrannosaurids, abelisaurids, or carcharodontosaurids. More recent research based on additional fossil material found support for a position within the tyrannosaurine clade Teratophoneini. Two species have been proposed: ''L. anomala'' (the type species), likely from the La Bocana Roja Formation, and ''L. aguillonae'' from the Cerro del Pueblo Formation. Discovery In the summer of 1970, the National Geographic Society and the Los Angeles County Museum of Natural History organized a joint paleontological expedition, led by geologist William J. Morris, to the Arroyo del Rosario in Baja California, Mexico. While prospecting, volunteer Harley J. Garbani discovered the fragmentary skeleton of a theropod north of Punta Baja near Cerro Rayado. The collected specimen, IG ...
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Teratophoneus
''Teratophoneus'' ("monstrous murderer"; Greek: ''teras'', "monster" and ''phoneus'', "murderer") is a genus of Tyrannosaurinae, tyrannosaurine Theropoda, theropod dinosaur that lived during the late Campanian age of the Late Cretaceous period, (about 76.5 to 75.5 million years ago) in what is now Utah. It contains a single known species, ''T. curriei''. It is known from an incomplete skull and postcranial skeleton recovered from the Kaiparowits Formation and was specific name (zoology), specifically named ''T. curriei'' in honor of famed paleontologist Philip J. Currie. Discovery and naming Fossils of ''Teratophoneus'' were first found in the Kaiparowits Formation of southern Utah. Later, fossils from the same formation were discovered and identified as the genus. Argon-argon dating, Argon-argon radiometric dating indicates that the Kaiparowits Formation was deposited between 76.1 and 74.0 million years ago, during the Campanian age of the Late Cretaceous period. This date mea ...
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Daspletosaurini
Daspletosaurini is an extinct clade (tribe) of tyrannosaurine dinosaurs that lived in Laramidia during the Late Cretaceous (Middle Campanian) period. It consists of two genera: ''Daspletosaurus'' and '' Thanatotheristes''. Four species have been described in the two genera, namely ''Daspletosaurus torosus'', ''Daspletosaurus horneri'', ''Daspletosaurus wilsoni'', and ''Thanatotheristes degrootorum''. At one point all of them were assigned as specimens of ''D. torosus'', but several papers since 2017 have found them to represent distinct species. Some researchers found anagenesis in the group, whether contained in a daspletosaurin clade or paraphyletic in respect to the lineage of tyrannosaurines leading up to ''Tyrannosaurus'', but a 2023 study refuted this theory on the basis of morphological and stratigraphical data. Description Daspletosaurins were large predators, reaching around in length. However, they were not the largest tyrannosaurids, as more derived taxa, such as ''Ty ...
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Zhuchengtyrannus
''Zhuchengtyrannus'' (meaning "Zhucheng tyrant") is a genus of Tyrannosauridae, tyrannosaurid theropod dinosaur known from the Campanian stage of the Late Cretaceous of Shandong Province, China. It belongs to the subfamily Tyrannosaurinae, and contains a single species, ''Zhuchengtyrannus magnus''. Discovery and naming ''Zhuchengtyrannus'' was first described and named by David W. E. Hone, Kebai Wang, Corwin Sullivan, Xijin Zhao, Shuqing Chen, Dunjin Li, Shuan Ji, Qiang Ji and Xing Xu in 2011 in paleontology, 2011 and the type species is ''Zhuchengtyrannus magnus''. The generic name is derived from the word ''Zhucheng'', which refers to the type locality, and ''tyrant'' in reference to its phylogenetic position as a tyrannosaurid. The specific name (zoology), specific name ''magnus'' meaning "great" in Latin refers to the relatively large size of ''Zhuchengtyrannus''. ''Zhuchengtyrannus'' is known solely from the holotype ZCDM V0031, a nearly complete right maxilla and associate ...
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Tyrannosauridae
Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to fifteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera is controversial, with some experts recognizing as few as three. All of these animals lived near the end of the Cretaceous Period and their fossils have been found only in North America and Asia. Although descended from smaller ancestors, tyrannosaurids were almost always the largest predators in their respective ecosystems, putting them at the apex of the food chain. The largest species was ''Tyrannosaurus rex'', the most massive known terrestrial predator, which measured over in length and according to most modern estimates up to in weight. Tyrannosaurids were bipedal carnivores with massive skulls filled with large teeth. Despite their large size, their legs were long and proportioned for fast movement. In contrast, their arms were ...
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Family (biology)
Family (, : ) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". The delineation of what constitutes a family—or whether a described family should be acknowledged—is established and decided upon by active taxonomists. There are not strict regulations for outlining or acknowledging a family, yet in the realm of plants, these classifications often rely on both the vegetative and reproductive characteristics of plant species. Taxonomists frequently hold varying perspectives on these descriptions, leading to a lack of widespread consensus within the scientific community ...
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