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Ruyangosaurus Scale
''Ruyangosaurus'' (Ruyang County lizard) is a genus of titanosauriform sauropod dinosaur recovered from the Early Cretaceous Haoling Formation of China. The type species is ''R. giganteus'', described in 2009 by Lü Junchang ''et al''. Description Along with '' Huanghetitan'' and '' Daxiatitan'', ''Ruyangosaurus'' is among the largest dinosaurs discovered in Cretaceous Asia. In 2016 Gregory S. Paul gave a length of 30 meters (100 ft) and a weight of 50+ tonnes (55 short tons) - making it a 'mega-sauropod'. ''Ruyangosaurus'' was probably about 35 meters (115 ft) long, as evidenced by its 207 cm long femur and 127 cm long right tibia. Classification The describers of ''Ruyangosaurus'' assigned it to Andesauridae. However, Andesauridae is not monophyletic and, as such, is no longer used. Comprehensive phylogenetic analyses of Titanosauriformes conducted by Philip Mannion and colleagues in 2013 found ''Ruyangosaurus'' to be in a polytomy with '' Andesaurus'' and other ...
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Early Cretaceous
The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java- Manihiki- Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kergue ...
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Ruyangosaurus Tibia
''Ruyangosaurus'' (Ruyang County lizard) is a genus of titanosauriform sauropod dinosaur recovered from the Early Cretaceous Haoling Formation of China. The type species is ''R. giganteus'', described in 2009 by Lü Junchang ''et al''. Description Along with '' Huanghetitan'' and '' Daxiatitan'', ''Ruyangosaurus'' is among the largest dinosaurs discovered in Cretaceous Asia. In 2016 Gregory S. Paul gave a length of 30 meters (100 ft) and a weight of 50+ tonnes (55 short tons) - making it a 'mega-sauropod'. ''Ruyangosaurus'' was probably about 35 meters (115 ft) long, as evidenced by its 207 cm long femur and 127 cm long right tibia. Classification The describers of ''Ruyangosaurus'' assigned it to Andesauridae. However, Andesauridae is not monophyletic and, as such, is no longer used. Comprehensive phylogenetic analyses of Titanosauriformes conducted by Philip Mannion and colleagues in 2013 found ''Ruyangosaurus'' to be in a polytomy with '' Andesaurus'' and other ...
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Euhelopus
''Euhelopus'' is a genus of sauropod dinosaur that lived between 145 and 133 million years ago during the Berriasian and Valanginian stages of the Early Cretaceous in what is now Shandong Province in China. It was a large quadrupedal herbivore. Unlike most other sauropods, ''Euhelopus'' had longer forelegs than hind legs. This discovery was paleontologically significant because it represented the first dinosaur scientifically investigated from China: seen in 1913, rediscovered in 1922, and excavated in 1923 and studied by T'an during the same year.T'an, H. C. (1923). New research on the Mesozoic and early Tertiary geology in Shantung. Geological Survey of China Bulletin 5:95-135 Unlike most sauropod specimens, it has a relatively complete skull. ''Euhelopus'' was a long-necked sauropod similar to '' Mamenchisaurus'', but its affinities are controversial. Most studies favor a close relationship between ''Euhelopus'' and titanosaurs, rather than mamenchisaurids. Description Si ...
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Qiaowanlong
''Qiaowanlong'' (meaning "Qiaowan dragon") is a genus of sauropod dinosaur. Fossils belonging to the genus were found in 2007 from the Yujinzi Basin of Gansu, China, and were described in 2009 in the ''Proceedings of the Royal Society'' B. The remains come from a geological formation called the Xiagou Formation in the Xinminpu Group, dating to the Early Cretaceous (late Aptian stage). The only known specimen consists of articulated cervical (neck) vertebrae and a right pelvic girdle, as well as several unidentified bone fragments. ''Qiaowanlong'' was initially reported as the first brachiosaurid to have been found from China. However, later analysis found that it was more closely related to titanosauriformes like ''Euhelopus'' and ''Erketu''. It is estimated to have had a length of around and would have weighed around 6 tonnes.Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 204 The type species In zoological nomenclature, a typ ...
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Tangvayosaurus
''Tangvayosaurus'' (meaning "Tang Vay lizard") is a genus of sauropod dinosaur from the Aptian-Albian age Lower Cretaceous Gres superieurs Formation of Savannakhet Province, Laos. It was a basal somphospondylan, about 15 m long, and is known from the remains of two or three individuals. Description It is based on TV4-1 to TV4-36, consisting of a partial pelvis, several back vertebrae and a tail vertebra, ribs, and an upper arm bone (humerus). Another skeleton includes 38 tail vertebrae, a neck vertebra, and most of a hind limb. The type species, ''Tangvayosaurus hoffeti'', was described by a group of a dozen scientists led by Ronan Allain in 1999 and the species name honours French palaeontologist Joshua Hoffet (1901-disappeared 1945). Allain ''et al.'' also referred the old species ''"Titanosaurus" falloti'' (Hoffet, 1942),J.-H. Hoffet. (1942). Description de quelques ossements de Titanosauriens du Sénonien du Bas-Laos escription of some titanosaurian bones fro ...
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Phuwiangosaurus
''Phuwiangosaurus'' (meaning "Phu Wiang lizard") is a genus of titanosaur dinosaur from the Early Cretaceous (Valanginian-Hauterivian) Sao Khua Formation of Thailand. The type species, ''P. sirindhornae'', was described by Martin, Buffetaut, and Suteethorn in a 1993 press release and was formally named in 1994. The species was named to honor Princess Maha Chakri Sirindhorn of Thailand, who was interested in the geology and palaeontology of Thailand, while the genus was named after the Phu Wiang area, where the fossil was discovered. In 2010 Paul gave a length of 19 meters (62 ft) and a weight of 17 tonnes (18.7 short tons). In 2012 Holtz gave a higher estimation of 25 meters (82 ft). ''Phuwiangosaurus'' was originally assigned to Titanosauria, but more recent studies have placed it in a more basal position within the Titanosauriformes. Phylogenetic analyses presented by D'Emic (2012), Mannion ''et al.'' (2013), and Mocho ''et al.'' (2014) resolve ''Phuwiangosaurus'' within ...
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Lithostrotia
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of '' Malawisaurus'' and '' Saltasaurus'' and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek , meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch ''et al.'' include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous. History of research In 1895, Richard Lydekker named the family Titanosauridae to summarize sauropods with procoelous (concave on the front) caudal vertebrae. The name Titanosauridae has since been widely used, and was defined by Salgado and colleagues (1997), Gonzalaz-Riga (2003), and Salgado (2003) as a node- ...
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Xianshanosaurus
''Xianshanosaurus'' () is a genus of sauropod dinosaur from the Early Cretaceous (Aptian-Albian) of the Ruyang Basin in Henan Province, China. Its type and only species is ''Xianshanosaurus shijiagouensis''. It was described in 2009 by a team of paleontologists led by Lü Junchang. ''Xianshanosaurus'' may be a titanosaur, and ''Daxiatitan'' may be its closest relative, but its evolutionary relationships remain controversial. Taxonomy ''Xianshanosaurus'' and its type species ''X. shijiagouensis'' were named by Lü Junchang, Xu Li, Jiang Xiaojun, Jia Songhai, Li Ming, Yuan Chongxi, Zhang Xingliao, and Ji Qiang in 2009. The type specimen, accessioned in the Henan Geological Museum, consists of ten caudal vertebrae, a coracoid, a femur, and several ribs. The genus name refers to Xian Mountain (), located near where the holotype was found. Phylogenetic relationships When ''Xianshanosaurus'' was first described, its discoverers noted that it had an unusual mixture of characteristic ...
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Titanosauria
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a ''nomen dubium)'' '' Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ...
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Huanghetitan Ruyangensis
''Huanghetitan'' (meaning "Yellow River titan"), is a genus of sauropod dinosaur from the early Cretaceous Period. It was a basal titanosauriform which lived in what is now Gansu, China. History The type species, ''Huanghetitan liujiaxiaensis'', was described by You ''et al.'' in 2006. It is known from fragmentary materials including two caudal vertebrae, an almost complete sacrum, rib fragments, and the left shoulder girdle, and was discovered in the eastern part of the Lanzhou Basin (Hekou Group) in the Gansu Province in 2004. A second species, ''H. ruyangensis'', was described in 2007 from the Aptian-Albian Haoling Formation of Ruyang County, China (Henan Province). A recent cladistic analysis has found that this species is unlikely to be closely related to ''H. liujiaxiaensis'' and requires a new genus name. Description ''H. liujiaxiaensis'' is a relatively small sauropod, measuring long and weighing . ''H. ruyangensis'' is known from a partial vertebral column a ...
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Yongjinglong
''Yongjinglong'' is an extinct genus of titanosauriform sauropod dinosaur known from the Early Cretaceous of Lanzhou-Minhe Basin of Gansu Province, China. It contains a single species, ''Yongjinglong datangi''. Discovery ''Yongjinglong'' was first described and named by Li-Guo Li, Da-Qing Li, Hai-Lu You and Peter Dodson in 2014 and the type species is ''Yongjinglong datangi''. The generic name is derived from the name of the historical Yongjing County, near where the holotype of ''Yongjinglong'' and numerous dinosaur track fossils were collected, and from ''long'', meaning " dragon" in Chinese. The specific name, ''datangi'', honors the Tang dynasty and also Mr. Zhi-Lu Tang from the IVPP, for his contributions to the study of dinosaurs. ''Yongjinglong'' is known solely from the holotype GSGM ZH(08)-04, a partial postcranial skeleton and three teeth, currently housed at the Gansu Geological Museum, Gansu Province. The postcranial remains include one fragmentary dorsal rib, th ...
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Euhelopodidae
Euhelopodidae is a family of sauropod dinosaurs of disputed membership and affinities, which contains ''Euhelopus'' and its close relatives. Most proposed euhelopodids are from East Asia. Euhelopodidae was first recognized by Carl Wiman in 1929, under the name Helopodidae, as ''Euhelopus'' was originally named ''Helopus''. However, the name had already been proposed for a bird, so in 1956 Alfred Sherwood Romer proposed the name ''Euhelopus'' and Euhelopodinae as replacements; Romer classified Euhelopodinae as a subfamily of Brachiosauridae, in which he also included Camarasaurinae and Cetiosaurinae, rather than as a family of its own. In addition to ''Euhelopus'' itself, Romer included ''Chiayusaurus'', ''Omeisaurus'', and ''Tienshanosaurus'' in Euhelopodinae. The taxonomic content of Euhelopodidae is uncertain, as a result of the unstable position of ''Euhelopus'' itself. Some studies have concluded that ''Euhelopus'' is a non-neosauropod closely related to ''Mamenchisaurus'', ...
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