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Pterygota
Pterygota ( ) is a subclass of insects that includes all winged insects and groups who lost them secondarily. Pterygota group comprises 99.9% of all insects. The orders not included are the Archaeognatha (jumping bristletails) and the Zygentoma ( silverfishes and firebrats), two primitively wingless insect orders. Unlike Archaeognatha and Zygentoma, the pterygotes do not have styli or vesicles on their abdomen (also absent in some zygentomans), and with the exception of the majority of mayflies, are also missing the median terminal filament which is present in the ancestrally wingless insects. The oldest known representatives of the group appeared during the mid-Carboniferous, around 328–324 million years ago, and the group subsequently underwent rapid diversification. Claims that they originated substantially earlier during the Silurian or Devonian based on molecular clock estimates are unlikely based on the fossil record, and are likely analytical artefacts. __TOC__ Sys ...
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Insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, Thorax (insect anatomy), thorax and abdomen (insect anatomy), abdomen), three pairs of jointed Arthropod leg, legs, compound eyes, and a pair of antenna (biology), antennae. Insects are the most diverse group of animals, with more than a million described species; they represent more than half of all animal species. The insect nervous system consists of a insect brain, brain and a ventral nerve cord. Most insects reproduce Oviparous, by laying eggs. Insects Respiratory system of insects, breathe air through a system of Spiracle (arthropods), paired openings along their sides, connected to Trachea#Invertebrates, small tubes that take air directly to the tissues. The blood therefore does not carry oxygen; it is only partly contained in ves ...
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Mayfly
Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order Ephemeroptera. This order is part of an ancient group of insects termed the Palaeoptera, which also contains dragonflies and damselflies. Over 3,000 species of mayfly are known worldwide, grouped into over 400 genera in 42 families. Mayflies have ancestral traits that were probably present in the first flying insects, such as long tails and wings that do not fold flat over the abdomen. Their immature stages are aquatic fresh water forms (called "naiads" or "nymphs"), whose presence indicates a clean, unpolluted and highly oxygenated aquatic environment. They are unique among insect orders in having a fully winged terrestrial preadult stage, the subimago, which moults into a sexually mature adult, the imago. Mayflies "hatch" (emerge ...
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Paleoptera
The name Palaeoptera (from Greek ( 'old') + ( 'wing')) has been traditionally applied to those ancestral groups of winged insects (most of them extinct) that lacked the ability to fold the wings back over the abdomen as characterizes the Neoptera. The Diaphanopterodea, which are palaeopteran insects, had independently and uniquely evolved a different wing-folding mechanism. Both mayflies and dragonflies lack any of the smell centers in their brain found in Neoptera. Disputed status The complexities of the wing-folding mechanism, as well as the mechanical operation of the wings in flight ( indirect flight muscles), are such that it indicates that Neoptera is a monophyletic lineage. The plesiomorphic absence of wing-folding does not necessarily mean the Palaeoptera form a natural group – they may be an assemblage containing all insects, closely related or not, that "are not Neoptera", an example of a wastebasket taxon. If the extinct lineages are taken into account, it is ...
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Neoptera
Neoptera (Ancient Greek ''néos'' ("new") + ''pterón'' ("wing")) is a classification group that includes most orders of the winged insects, specifically those that can flex their wings over their abdomens. This is in contrast with the more basal orders of winged insects (the " Palaeoptera" assemblage), which are unable to flex their wings in this way. Classification The taxon Neoptera was proposed by А.М. Martynov in 1923 and 1924, in the following classification: Pterygota *division Palaeoptera **order Odonata **order Agnatha (correct name: Ephemeroptera) **†order Dictyoneuridea **†order Megasecoptera **†order Meganisoptera **†order Protephemeroidea *division Neoptera **superorder Endopterygota ***order Coleoptera ***order Strepsiptera ***order Neuroptera ***order Raphidioptera ***order Megaloptera ***order Diptera ***order Mecoptera ***order Trichoptera ***order Lepidoptera ***order Hymenoptera **subdivision Polyneoptera ***superorder Orthopteroid ...
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Odonatoptera
The Odonatoptera are a superorder (sometimes treated as an order) of ancient winged insects, placed in the probably paraphyletic group Palaeoptera. The dragonflies and damselflies (which are placed in the subgroup Odonata) are the only living members of this group, which was far more diverse in the late Paleozoic and contained gigantic species, including the griffinflies (colloquially called "giant dragonflies", although they were not dragonflies in the strict sense) of the order Meganisoptera (formerly Protodonata). This lineage dates back at least to the Bashkirian, not quite 320 million years ago.  Systematics and taxonomy There is little consensus about the relationships of the Odonatoptera. What is certain is that they are a clade of winged insects that stands outside the Neoptera. But various authors' analyses have yielded any one of three mutually exclusive phylogenies, or some variant thereof: The least problematic (in a taxonomic sense) view is that the Odonatopte ...
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Zygentoma
Zygentoma are an order in the class Insecta, and consist of about 550 known species. The Zygentoma include the so-called silverfish or fishmoths, and the firebrats. A conspicuous feature of the order are the three long caudal filaments. The two lateral filaments are cerci, and the medial one is an epiproct or ''appendix dorsalis''. In this they resemble the Archaeognatha, although the cerci of Zygentoma, unlike in the latter order, are nearly as long as the epiproct. Until the late twentieth century the Zygentoma were regarded as a suborder of the Thysanura, until it was recognized that the order Thysanura was paraphyletic, thus raising the two suborders to the status of independent monophyletic orders, with Archaeognatha as sister group to the Dicondylia, including the Zygentoma. Etymology The name "Zygentoma" is derived from the Greek ('), in context meaning "yoke" or "bridge"; and ('), "insects" (literally meaning "cut into", in reference to the segmented anatomy o ...
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Metapterygota
Metapterygota is a clade of winged insects containing order Odonata and Infraclass Neoptera. They share morphological characteristics of a loss of caudal filaments and a subimago stage. Genetically the clade is supported by similarities in the mitochondrial genomes. Classification * Superorder Odonatoptera ** Order Protodonata or Meganisoptera †(extinct) ** Order Protanisoptera †(extinct) ** Order Triadophlebioptera †(extinct) ** Order Protozygoptera or Archizygoptera †(extinct) ** Order Odonata (dragonflies and damselflies) * Infraclass Neoptera ** Polyneoptera *** Order Caloneurodea †(extinct) *** Order Glosselytrodea †(extinct) *** Order Titanoptera †(extinct) *** Order Protorthoptera †(extinct) *** Order Plecoptera (stoneflies) *** Order Embioptera (webspinners) *** Order Zoraptera (angel insects) *** Order Dermaptera (earwigs) *** Order Orthoptera (grasshoppers, etc.) *** Order Phasmatodea (stick insects) *** Superorder Notoptera **** Order Gryllobl ...
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Exopterygota
The Exopterygota (Ancient Greek ''ἔξω'' (éxō, "outside") + ''πτερόν'' (pterón, "wing") + Neo-Latin ''-ota'' ("having")), also known as Hemimetabola, are a superorder of insects of the subclass Pterygota in the infraclass Neoptera, in which the young resemble adults but have externally developing wings. They undergo a modest change between immature and adult, without going through a pupal stage. The nymphs develop gradually into adults through a process of moulting. The Exopterygota are a highly diverse insect superorder, with at least 130,000 living species divided between 15 orders. They include cockroaches, termites, grasshoppers, thrips, lice and stick insects, among many other types of insects. Exopterygota are distinguished from the Endopterygota (or Holometabola) by the way their wings develop. Endopterygota (meaning literally "internal winged forms") develop wings inside the body and undergo an elaborate metamorphosis involving a pupal stage. Exopterygota ...
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Infraclass
In biology, taxonomic rank (which some authors prefer to call nomenclatural rank because ranking is part of nomenclature rather than taxonomy proper, according to some definitions of these terms) is the relative or absolute level of a group of organisms (a ''taxon'') in a hierarchy that reflects evolutionary relationships. Thus, the most inclusive clades (such as Eukarya and Animalia) have the highest ranks, whereas the least inclusive ones (such as ''Homo sapiens'' or ''Bufo bufo'') have the lowest ranks. Ranks can be either relative and be denoted by an indented taxonomy in which the level of indentation reflects the rank, or absolute, in which various terms, such as species, genus, family, order, class, phylum, kingdom, and domain designate rank. This page emphasizes absolute ranks and the rank-based codes (the Zoological Code, the Botanical Code, the Code for Cultivated Plants, the Prokaryotic Code, and thCode for Viruses require them. However, absolute ranks are not req ...
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Palaeodictyopteroidea
The Palaeodictyopteroidea or Paleodictyopterida are an extinct superorder of Palaeozoic beaked insects, characterised by unique mouthparts consisting of 5 stylets. They represent the first important terrestrial herbivores, and the first major group of herbivorous insects. They appear during the Middle Carboniferous (late Serpukhovian or early Bashkirian) and continue through to the Lopingian, Late Permian. This large and diverse group includes 50% of all known Paleozoic insects. Palaeodictyopteroidea nymphs possessed movable wing pads and appear to have been able to perform simple flapping flight. References * * * External linksPaleodictyopteroideaat the Tree of Life project
by Nina Sinitshenkova, 2002 Palaeodictyopteroidea, Carboniferous insects Permian insects Pennsylvanian first appearances Lopingian extinctions Insect superorders Taxa named by Boris Rohdendorf Pterygota {{paleo-insect-stub ...
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Firebrat
The firebrat (''Thermobia domestica'') is a small insect (typically 1–1.5 cm) in the order Zygentoma. Habitat Firebrats prefer relatively warm temperatures (36–39 °C) and require some humidity. They are commonly found indoors near heat sources such as furnaces and boilers. They feed on a wide variety of carbohydrates and starches that are also protein sources such as dog food, flour and book bindings. They are distributed throughout most parts of the world and are normally found outdoors under rocks, plant litter, and in similar environments, but are also often found indoors where they are considered pests. They do not cause major damage, but they can contaminate food, damage paper goods, and stain clothing. Otherwise they are mostly harmless. Behavior Firebrats utilize pheromones to lead other firebrats to attract one another and congregate. To maintain a group, firebrats must remain in contact with one another. Breeding At 1.5 to 4.5 months of age t ...
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Class (biology)
In biological classification, class () is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are domain, kingdom, phylum, order, family, genus, and species, with class ranking between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name – and not just called a ''top-level genus'' ''(genus summum)'' – was first introduced by French botanist Joseph Pitton de Tournefort in the classification of plants that appeared in his '' Eléments de botanique'' of 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of construction, whic ...
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