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Ottoia
''Ottoia'' is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to ''Ottoia'' on spurious grounds, the only clear ''Ottoia'' macrofossils come from the Burgess Shale of British Columbia, which was deposited . Microfossils extend the record of ''Ottoia'' throughout the Western Canada Sedimentary Basin, from the mid- to late- Cambrian. A few fossil finds are also known from China. Morphology ''Ottoia'' specimens are on average 8 centimeters in length. Both length and width show variation with contraction; shorter specimens often being wider than longer ones. The characteristic proboscis of priapulids is present at the anterior, attached to the trunk of the animal, proceeded by the "bursa" at the posterior. The organism's body is bilaterally symmetrical, however, its anterior displays external radial symmetry. Like some other modern invertebrates, a cuticle restricts the size of and prot ...
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Archaeopriapulida
Archaeopriapulida is a group of priapulid worms known from Cambrian lagerstätte. The group is closely related to, and very similar to, the modern Priapulids. It is unclear whether it is mono- or polyphyletic. Despite a remarkable morphological similarity to their modern cousins, they fall outside of the priapulid crown group, which is not unambiguously represented in the fossil record until the Carboniferous. In addition to well-preserved body fossils, remains of several archaeopriapulid taxa are known to have been preserved primarily as organic microfossils, such as isolated scalids and pharyngeal teeth. They are probably closely related or paraphyletic to the palaeoscolecids; the relationship between these basal worms is somewhat unresolved. Species * Genus '' Acosmia'' Chen & Zhoi 1997 (Chengjiang deposits) ** '' Acosmia maotiania'' Chen & Zhoi 1997 * Genus '' Archotuba'' Hou ''et al''. 1999 (possibly a Cnidarian) (Chengjiang deposits) ** '' Archotuba conoidalis'' Hou ' ...
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Priapulida
Priapulida (priapulid worms, from Gr. πριάπος, ''priāpos'' ' Priapus' + Lat. ''-ul-'', diminutive), sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis. They live in the mud, except for a few tropical meiobenthic species which live in medium- to coarse-grained sands, and are found in comparatively shallow waters to deep waters and no warmer than 12–13°C. Some species show a remarkable tolerance for hydrogen sulfide, anoxia and low salinity. ''Halicryptus spinulosus'' appears to prefer brackish shallow waters. They can be quite abundant in some areas. In an Alaskan bay as many as 85 adult individuals of '' Priapulus caudatus'' per square meter has been recorded, while the density of its larvae can be as high as 58,000 per square meter (5,390 per square foot). ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, are fossils that formed around 505 million years ago in the mid-Cambrian, Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's 1989 book ''Wonderful Life (book), Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were ...
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Paleobiota Of The Burgess Shale
This is a list of the Biota (ecology), biota of the Burgess Shale, a Cambrian lagerstätte located in Yoho National Park in Canada. The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old (Wuliuan, middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. During the Cambrian, the ecosystem of the Burgess Shale sat under 100 to 300 metres (330 to 1000 feet) of water at the base of a submarine canyon known as the Cathedral escarpment, Cathedral Escarpment, which today is a part of the Canadian Rockies. The ecosystem would have sat in dimly lit water, most likely at the edge, or in the Mesopelagic zone. The ecosystem was preserved by rapid mudslides that quickly buried organisms near, or on the seafloor, which helps explain the rarity of nektonic organisms at the site. The shale would have supported uniq ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wife, establishing a quarry on the flanks of Fossil Ridge. The significance of soft-bodied preservation, and the range of organisms he recognised as new to science, led him to return to the quarry almost every year until 1924. At that point, aged 74, he had amassed over 65,000 s ...
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Lower Ordovician
The Early Ordovician is the first epoch of the Ordovician period, corresponding to the Lower Ordovician series of the Ordovician system. It began after the Age 10 of the Furongian epoch of the Cambrian and lasted from to million years ago, until the Dapingian age of the Middle Ordovician. It includes Tremadocian and Floian ages. History International Commission on Stratigraphy (ICS) appointed working groups on the boundaries of the Ordovician subdivisions in 1974. The boundaries were established by the fauna of conodonts and/or graptolites. In 1995, the Subcommission on Ordovician Stratigraphy, with the support of 90% majority of voting members, adopted the division of the Ordovician system into three series: Lower, Middle and Upper. In the same year, it was decided to divide each of the three series into two global stages. ''Tetragraptus approximatus'' zone was chosen as the base of the upper stage of the Lower Ordovician by 95% of the votes. Before the adoption of the in ...
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Wheeler Shale
The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including ''Naraoia'', ''Wiwaxia'' and ''Hallucigenia'') and preservation style ( carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and t ...
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Gizzard
The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including archosaurs (birds and other dinosaurs, crocodiles, alligators, pterosaurs), earthworms, some gastropods, some fish, and some crustaceans. This specialized stomach constructed of thick muscular walls is used for grinding up food, often aided by particles of stone or grit. In certain insects and molluscs, the gizzard features chitinous plates or teeth. Etymology The word ''gizzard'' comes from the Middle English ''giser'', which derives from a similar word in Old French ''gésier'', which itself evolved from the Latin">-4; we might wonder whether there's a point at which it's appropriate to talk of the beginnings of French, that is, when it wa ... ''gésier'', which itself evolved from the Latin ''gigeria'', meaning giblets. Structure In birds Birds swallow food and store it in their crop if necessary. Then the food passes into thei ...
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