Lower Ordovician
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The Early Ordovician is the first
epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided b ...
of the
Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years f ...
period, corresponding to the Lower Ordovician
series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used i ...
of the Ordovician system. It began after the Age 10 of the
Furongian The Furongian is the fourth and final epoch and series of the Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding ...
epoch of the
Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ...
and lasted from to million years ago, until the Dapingian age of the
Middle Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era, and the second of twelve periods of the Phanerozoic Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period Ma (milli ...
. It includes
Tremadocian The Tremadocian is the lowest stage of Ordovician. Together with the later Floian Stage it forms the Lower Ordovician Epoch. The Tremadocian lasted from to million years ago. The base of the Tremadocian is defined as the First appearance datum ...
and
Floian The Floian is the second stage of the Ordovician Period. It succeeds the Tremadocian with which it forms the Lower Ordovician series. It precedes the Dapingian Stage of the Middle Ordovician. The Floian extended from to million years ago. The ...
ages.


History

International Commission on Stratigraphy The International Commission on Stratigraphy (ICS), sometimes unofficially referred to as the International Stratigraphic Commission, is a daughter or major subcommittee grade scientific organization that concerns itself with stratigraphy, strati ...
(ICS) appointed working groups on the boundaries of the Ordovician subdivisions in 1974. The boundaries were established by the fauna of
conodonts Conodonts, are an extinct group of marine jawless vertebrates belonging to the Class (biology), class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning "cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known ...
and/or
graptolites Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian (Miaolingian, Wuliuan) through th ...
. In 1995, the Subcommission on Ordovician Stratigraphy, with the support of 90% majority of voting members, adopted the division of the Ordovician system into three series: Lower,
Middle Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek ...
and Upper. In the same year, it was decided to divide each of the three series into two global stages. ''Tetragraptus approximatus'' zone was chosen as the base of the upper stage of the Lower Ordovician by 95% of the votes. Before the adoption of the international standard, the series from the chart of Britain, the type locality of the Ordovician system, were accepted by default as Ordovician subdivisions. In accordance with this scale, the Ordovician was divided into six series, of which the lower one, the
Tremadocian The Tremadocian is the lowest stage of Ordovician. Together with the later Floian Stage it forms the Lower Ordovician Epoch. The Tremadocian lasted from to million years ago. The base of the Tremadocian is defined as the First appearance datum ...
, passed into the
International Stratigraphic Chart The International Commission on Stratigraphy (ICS), sometimes unofficially referred to as the International Stratigraphic Commission, is a daughter or major subcommittee grade scientific organization that concerns itself with stratigraphical, ge ...
(ISC) as the stage of the same name. The upper
Arenig In geology, the Arenig (or Arenigian) is a time interval during the Ordovician period and also the suite of rocks which were deposited during this interval. History The term was first used by Adam Sedgwick in 1847 with reference to the "Areni ...
series corresponds to the upper part of the Lower and lower part of the Middle Ordovician global series. On the British chart, the boundaries of Ordovician subdivisions are determined by local stratigraphic and paleontological features that are poorly defined in the rest of the world. As a result, local series and stages, very different from the British standard, were identified on different paleoplates or modern continents. In 2008, the traditional British Ordovician subdivisions were replaced by seven new stages with defined GSSPs. Tremadocian and Floian stages were included in the Lower Ordovician series. In 2011, the Russian regional stratigraphic chart was changed from the British to the international standard division of the Ordovician system.


Definition

The
Global Boundary Stratotype Section and Point A Global Boundary Stratotype Section and Point (GSSP), sometimes referred to as a golden spike, is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. ...
(GSSP) of the Lower Ordovician, which is also a GSSP of the
Tremadocian The Tremadocian is the lowest stage of Ordovician. Together with the later Floian Stage it forms the Lower Ordovician Epoch. The Tremadocian lasted from to million years ago. The base of the Tremadocian is defined as the First appearance datum ...
stage Stage, stages, or staging may refer to: Arts and media Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly Brit ...
and the whole Ordovician system, is established in the Green Point section in the west of the
Newfoundland Newfoundland and Labrador is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region of Labrador, having a total size of . As of 2025 the population ...
,
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
() and corresponds to the
first appearance datum First appearance datum (FAD) is a term used by geologists and paleontologists to designate the first appearance of a species in the geologic record. FADs are determined by identifying the geologically oldest fossil discovered, to date, of a partic ...
(FAD) of the
conodont Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
''
Iapetognathus fluctivagus ''Iapetognathus fluctivagus'' is a species of denticulate cordylodan conodonts belonging to the genus '' Iapetognathus''. It existed during the Tremadocian Age ( million years ago) of the Ordovician. It is an important index fossil in biostrat ...
'' at the 101.8 m above the base of the outcrop. Doubts have been expressed about the identification of the index taxon, but it occurs below the '' Rhabdinopora'' and related graptolites. The GSSP of the
Floian The Floian is the second stage of the Ordovician Period. It succeeds the Tremadocian with which it forms the Lower Ordovician series. It precedes the Dapingian Stage of the Middle Ordovician. The Floian extended from to million years ago. The ...
, the second and last stage of the Lower Ordovician, is established in the Diabasbrottet Quarry,
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
, () and defined by FAD of the
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian ( Miaolingian, Wuliuan) through t ...
'' Tetragraptus approximatus''. Two Auxiliary boundary Stratotype Sections and Points (ASSPs) were also established for the Lower Ordovician/Tremadocian: the first one in the Lawson Cove section in
Millard County, Utah Millard County ( ) is a county in the U.S. state of Utah. As of the 2020 United States census, the population was 12,975. Its county seat is Fillmore, and the largest city is Delta. History The Utah Territory legislature created the county ...
, USA, and the second one in the Xiaoyangqiao section,
North China North China () is a list of regions of China, geographical region of the People's Republic of China, consisting of five province-level divisions of China, provincial-level administrative divisions, namely the direct-administered municipalities ...
. Both ASSPs were approved by the Subcommission on Ordovician Stratigraphy in 2016 and 2019, respectively, but, in 2021, the
International Union of Geological Sciences The International Union of Geological Sciences (IUGS) is an international non-governmental organization devoted to global cooperation in the field of geology. As of 2023, it represents more than 1 million geoscientists around the world. About Fo ...
(IUGS) proposed to deny the use of specific points and replace them by Standard Auxiliary Boundary Stratotypes (SABS) for more "flexible" correlations with GSSPs.


Biostratigraphy

The global stages of the Lower Ordovician, as well as of other series, are also subdivided into stage slices (time slices), which are corresponding to the base of conodont or graptolite zones. In addition to the global stages, the ''Tetragraptus approximatus'' Zone is also defined in the regional subdivisions of the UK and
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
, and the ''Didymograptus protobifidus'' Zone in those of
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, Australia and Baltoscandia.


North America

The following conodont zones are distinguished in Early Ordovician deposits of
Boothia Peninsula Boothia Peninsula (; formerly ''Boothia Felix'', Inuktitut ''Kingngailap Nunanga'') is a large peninsula in Nunavut's northern Canadian Arctic, south of Somerset Island. The northern part, Murchison Promontory, is the northernmost point of ...
, Canada (from upper to lower): * ''Oepikodus communis'' and ''Reutterodus andinus'' ( Ship Point Formation, Floian); * ''Rossodus manitouensis'' and ''Acodus deltatus/Oneotodus costatus'' (upper part of Turner Cliffs Formation, Tremadocian); * ''Cordylodus angulatus'' (lower part of Turner Cliffs, Tremadocian).


Asia

Graptolite (g) and conodont (c) zonal subdivisions of southern
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
(from upper to lower): * ''Ps. angustifolius elongatus/E. broggeri'' (g), ''D. protobifidus'' (g), ''Ph. densus'' (g), upper part of ''Oepikodus evae'' (c), plus chitinozoan ''Conochitina raymondi'' Zone (Floian); * Lower part of ''Ph. densus'' (g), ''Ac. balticus'' (g), lower part of ''Oepikodus evae'' (c) (Floian); * ''T. approximatus'' (g), upper part of ''Paroistodus proteus'' (c) (Floian); * Lower part of ''Paroistodus proteus'' (c) (Tremadocian); * ''K. kiaeri/Ad. tenellus'' (g), ''B. ramosus/Tr. osloensis/Al. hyperboreus'' (g) (Tremadocian); * ''Iapetognathus fluctivagus'' (c) (Tremadocian).


Australia

On the Australian scale, the global Lower Ordovician roughly corresponds to the Lancefieldian, Bendigonian, Chewtonian and lower Castlemainian stages.


Paleogeography

The
Iapetus Ocean The Iapetus Ocean (; ) existed in the late Neoproterozoic and early Paleozoic eras of the geologic timescale (between 600 and 400 million years ago). It was in the southern hemisphere, between the paleocontinents of Laurentia, Baltica and Avalon ...
, located between
Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...
,
Laurentia Laurentia or the North American craton is a large continental craton that forms the Geology of North America, ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of ...
and
Baltica Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, i ...
, reached over 4000 km in width at the beginning of the Ordovician. In the north, between eastern Laurentia and Siberia, it connected with
Panthalassa Panthalassa, also known as the Panthalassic Ocean or Panthalassan Ocean (from Greek "all" and "sea"), was the vast superocean that encompassed planet Earth and surrounded the supercontinent Pangaea, the latest in a series of supercontinent ...
, which covered about half the planet. Formed in late Cambrian,
Rheic Ocean The Rheic Ocean (; ) was an ocean which separated two major paleocontinents, Gondwana and Laurussia ( Laurentia- Baltica- Avalonia). One of the principal oceans of the Paleozoic, its sutures today stretch from Mexico to Turkey and its closure r ...
stretched between Gondwana and the microcontinent of
Avalonia Avalonia was a microcontinent in the Paleozoic era. Crustal fragments of this former microcontinent are terranes in parts of the eastern coast of North America: Atlantic Canada, and parts of the East Coast of the United States, East Coast of the ...
. Cuyania (northwest of today's
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
) was located at low latitudes. Gondwana, already a large continent, consisted of today's
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
,
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, the
Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...
, India, eastern Australia and eastern
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
. Laurentia included the territories of
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, the USA,
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
,
Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
,
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
and partly
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
. Laurentia was located in the tropical latitudes of the southern hemisphere; today's central
Nevada Nevada ( ; ) is a landlocked state in the Western United States. It borders Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the seventh-most extensive, th ...
and western
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
were covered by sea waters at the end of the Early Ordovician.


Climate

In 2007, Bassett et al. analyzed the oxygen isotope values of Early Ordovician strata of the Lange Ranch section in central
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
and concluded that tropical sea temperatures at that time could have reached 37°C or 42°C. Similar results were recovered by Trotter et al. in 2008, after oxygen
isotope analysis Isotope analysis is the identification of isotopic signature, abundance of certain stable isotopes of chemical elements within organic and inorganic compounds. Isotopic analysis can be used to understand the flow of energy through a food we ...
of conodonts from the four paleoplates located in the Early Ordovician at low latitudes. Authors of the 2021 article give values in accordance with the study by Song et al. (2019): from 43.9°C 485 million years ago to 37.1°C 470 million years ago. High temperatures that persisted throughout the Early Ordovician affected the biodiversification later, in the Middle Ordovician. CO2 level in Early Ordovician atmosphere was high while
oxygen Oxygen is a chemical element; it has chemical symbol, symbol O and atomic number 8. It is a member of the chalcogen group (periodic table), group in the periodic table, a highly reactivity (chemistry), reactive nonmetal (chemistry), non ...
levels varied from approximately 10% to 13%. Studies of
oxygen isotopes There are three known stable isotopes of oxygen (8O): , , and . Radioactive isotopes ranging from to have also been characterized, all short-lived. The longest-lived radioisotope is with a half-life of , while the shortest-lived isotope is ...
from
apatite Apatite is a group of phosphate minerals, usually hydroxyapatite, fluorapatite and chlorapatite, with high concentrations of Hydroxide, OH−, Fluoride, F− and Chloride, Cl− ion, respectively, in the crystal. The formula of the admixture of ...
s (particularly conodont apatites from Nevada and Utah) show that at the end of the Early Ordovician the climate began to change from greenhouse to icehouse. These changes occurred gradually and were accompanied by fluctuations in sea surface temperature with an approximate duration of 104 to 107 years. By the end of the
Middle Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era, and the second of twelve periods of the Phanerozoic Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period Ma (milli ...
, the surface of the low-latitude seas had cooled to temperatures comparable to today's equatorial ones.


Major events

The
Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE) was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced wi ...
(GOBE), manifested in an increase in the diversity of marine life, occurred in Early Ordovician. Along with the Cambrian explosion and radiation in the early
Mesozoic The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian r ...
, it is the most significant biodiversification in the
Phanerozoic The Phanerozoic is the current and the latest of the four eon (geology), geologic eons in the Earth's geologic time scale, covering the time period from 538.8 million years ago to the present. It is the eon during which abundant animal and ...
eon. This event occurred at different intervals depending on the groups of organisms and the geographical region, and, in fact, the GOBE refers to a whole complex of sequential and interconnected processes of
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
and migration of organisms. The first phase, associated with
planktonic Plankton are the diverse collection of organisms that drift in water (or air) but are unable to actively propel themselves against currents (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they pro ...
communities, probably began in the late
Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ...
and ended in the late Early Ordovician. The Early Ordovician marks the beginning of the second phase, which affected the inhabitants of
benthos Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone.Dumugol Formation,
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...
. This reefs, built by the microorganisms and
sponge Sponges or sea sponges are primarily marine invertebrates of the animal phylum Porifera (; meaning 'pore bearer'), a basal clade and a sister taxon of the diploblasts. They are sessile filter feeders that are bound to the seabed, and a ...
s of the genus '' Archaeoscyphia'', are different from the Cambrian and early Tremadocian and this discovery confirm that reef communities occupied deeper-sea habitats as early as in the middle Tremadocian. The radiation of marine life during the GOBE in the Early Ordovician was resulted by the transition from sulphidic to oxic conditions in the oceans of that time. Substantial older ("pre-GOBE") radiations are observed in the Early Ordovician sediments of
South China South China ( zh, s=, p=Huá'nán, j=jyut6 naam4) is a geographical and cultural region that covers the southernmost part of China. Its precise meaning varies with context. A notable feature of South China in comparison to the rest of China is ...
. However, it is questionable, were they global or not. From the
Furongian The Furongian is the fourth and final epoch and series of the Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding ...
to the end of the Early Ordovician, 495-470 Ma, the Ollo de Sapo magmatic event occurred on the northwestern territory of
Gondwana Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...
, which is now the
Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...
.


Paleontology

In Early Ordovician (Floian) strata of the San Juan Formation, Argentina, the oldest
microfossils A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, ...
, known as calcisphers or calcitarchs, are present. Early forms reached 80 to 250 μm in diameter. Some of these organisms are probably
algae Algae ( , ; : alga ) is an informal term for any organisms of a large and diverse group of photosynthesis, photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular ...
. Calcitarchs lived in the subtidal zone, from coastal waves to shoals and
reef A reef is a ridge or shoal of rock, coral, or similar relatively stable material lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic (non-living) processes such as deposition (geol ...
s. Algae of the genus '' Amsassia'' appeared. They already lived off the southern coast of
Laurentia Laurentia or the North American craton is a large continental craton that forms the Geology of North America, ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of ...
and Cuyania in the Early Ordovician, and later occupied an even wider range and disappeared only during the Ordovician-Silurian extinction. In Tremadocian, ''Amsassia'' inhabited small reefal mounds in shallow sea waters. Discovered in the Floian strata of
Newfoundland Newfoundland and Labrador is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region of Labrador, having a total size of . As of 2025 the population ...
, coral-like
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s of '' Reptamsassia divergens'' and ''Reptamsassia minuta'' allows to judge the level of development of reef ecosystems of the Early Ordovician, since they are the oldest example of
symbiotic Symbiosis (Ancient Greek : living with, companionship < : together; and ''bíōsis'': living) is any type of a close and long-term biolo ...
intergrowth of modular species.
Echinoderm An echinoderm () is any animal of the phylum Echinodermata (), which includes starfish, brittle stars, sea urchins, sand dollars and sea cucumbers, as well as the sessile sea lilies or "stone lilies". While bilaterally symmetrical as ...
diversity increased in the Early Ordovician: new classes appeared, including
asteroids An asteroid is a minor planet—an object larger than a meteoroid that is neither a planet nor an identified comet—that orbits within the Solar System#Inner Solar System, inner Solar System or is co-orbital with Jupiter (Trojan asteroids). As ...
, ophiuroids,
crinoid Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that remain attached to the sea floor by a stalk in their adult form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are ...
s and
diploporita Diploporita is an extinct group of blastozoans that ranged from the Ordovician to the Devonian. These echinoderms are identified by a specialized respiratory structure, called diplopores. Diplopores are a double pore system that sit within a depr ...
ns. Various Early Ordovician echinoderms are found in the Fillmore Formation in
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
, USA,
Fezouata Formation The Fezouata Formation or Fezouata Shale is a geological formation in Morocco which dates to the Ordovician, Early Ordovician.
in
Anti-Atlas The Anti-Atlas, also known as Lesser Atlas or Little Atlas, is a mountain range in Morocco, a part of the Atlas Mountains in the northwest of Africa. The Anti-Atlas extends from the Atlantic Ocean in the southwest toward the northeast, to the heig ...
, Morocco, and Saint-Chinian Formation in
Montagne Noire The Montagne Noire (; , known as the 'Black Mountain' in English) is a mountain range in central southern France. It is located at the southwestern end of the Massif Central at the juncture of the Tarn, Hérault and Aude departments. Its highe ...
, France.
Nektaspida Nektaspida (also called Naraoiida, Nektaspia and Nectaspida) is an extinct order of non- mineralised artiopodan arthropods. They are known from the lower-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficiall ...
have become less diverse after Cambrian. They most likely inhabited restricted or colder
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
seas in the Ordovician. '' Tariccoia tazagurtensis'' is a member of this arthropod clade from the Lower Ordovician of Morocco. In the same Fezouata Formation, ''
Aegirocassis ''Aegirocassis'' is an extinct genus of giant radiodont arthropod belonging to the family Hurdiidae that lived 480 million years ago during the early Ordovician in the Fezouata Formation of Morocco. It is known by a single species, ''Aegirocassis ...
'' was discovered. This
filter feeder Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a s ...
arthropod reached over 2 m in length and was the largest animal of its time. Marrellomorphs, appeared in Cambrian, continued to exist in Early Ordovician. Soft-bodied and shelly organisms, including Cambrian
relic In religion, a relic is an object or article of religious significance from the past. It usually consists of the physical remains or personal effects of a saint or other person preserved for the purpose of veneration as a tangible memorial. Reli ...
s as well as the new taxa of the Ordovician origin, constitute a fossil Liexi fauna, preserved in the Lower Ordovician Madaoyu Formation of
Hunan Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
,
South China South China ( zh, s=, p=Huá'nán, j=jyut6 naam4) is a geographical and cultural region that covers the southernmost part of China. Its precise meaning varies with context. A notable feature of South China in comparison to the rest of China is ...
. Fossil specimens include
bryozoa Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic animal, aquatic invertebrate animals, nearly all living in sedentary Colony (biology), colonies. Typically about long, they have a spe ...
ns,
sponge Sponges or sea sponges are primarily marine invertebrates of the animal phylum Porifera (; meaning 'pore bearer'), a basal clade and a sister taxon of the diploblasts. They are sessile filter feeders that are bound to the seabed, and a ...
s, echinoderms,
polychaete Polychaeta () is a paraphyletic class of generally marine Annelid, annelid worms, common name, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called c ...
s,
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian ( Miaolingian, Wuliuan) through t ...
s,
trilobite Trilobites (; meaning "three-lobed entities") are extinction, extinct marine arthropods that form the class (biology), class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most succ ...
s and
conodont Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
s.
Palaeoscolecid The palaeoscolecids are a group of extinct ecdysozoan worms resembling armoured priapulids. They are known from the Lower Cambrian to the lower Ludfordian (late Silurian); they are mainly found as disarticulated sclerites, but are also preserved ...
an worms, including '' Liexiscolex'' and a possible '' Ottoia'' specimen, are of great interest. Early Ordovician priapulids, similar to their Cambrian predecessors, are also known from northern China. In the deep-water sediments of the Early Ordovician (Floian) Al Rose Formation in the
Inyo Mountains The Inyo Mountains are a short mountain range east of the Sierra Nevada in eastern California in the United States. The range separates the Owens Valley to the west from Saline Valley to the east, extending for approximately south-southeast f ...
,
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
, the trilobite fauna have been discovered. Despite the low species diversity, this fauna is unique due to differences in the composition of families from more eastern complexes of the comparable age. Fossils identified as belonging to the '' Globampyx'', '' Protopresbynileus'', '' Carolinites'', '' Cloacaspis'', '' Geragnostus'' and '' Hintzeia'' genera. In the Early Ordovician, this territory was located near the paleocontinent of Laurentia.


Mineral resources

Oil An oil is any nonpolar chemical substance that is composed primarily of hydrocarbons and is hydrophobic (does not mix with water) and lipophilic (mixes with other oils). Oils are usually flammable and surface active. Most oils are unsaturate ...
and
gas Gas is a state of matter that has neither a fixed volume nor a fixed shape and is a compressible fluid. A ''pure gas'' is made up of individual atoms (e.g. a noble gas like neon) or molecules of either a single type of atom ( elements such as ...
exploration is underway in the Early Ordovician Tongzi and Meitan formations in Sichuan Basin and the Early Ordovician strata of the Tarim Basin, China. In the Lower Ordovician of the Tarim Basin near Tazhong, oil accumulations are found at depths of up to 9000 m.


References


Further reading

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External links

* * * {{Geological history, p, m * Geological epochs Ordovician geochronology