Notocupes
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Notocupes
''Notocupes'' is an extinct genus of medium-sized archostematan beetles from the Mesozoic Era of Eurasia, including over 50 described species. Historically, the genus was classified as a member of the family Ommatidae, but the presence of characters such as the horizontal mandibular cutting edge, separated procoxae and overlapping abdominal sternites indicate that the genus may have a closer affinity with the family Cupedidae. ''Notocupes'' is considered to be a junior synonym of '' Zygadenia'' by Kirejtshuk (2020), but other researchers suggest to reserve the genus ''Zygadenia'' as a form taxon for isolated elytra that probably belong to the genus ''Notocupes'', while retaining ''Notocupes'' as a valid genus for complete body fossils. Most species of ''Notocupes'' were described from compression fossils. An additional three species were described from Cenomanian-aged Burmese amber, which were treated as a separate genus, '' Echinocups'', by Kirejtshuk (2020), but Li ''et al.'' (2 ...
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Ommatidae
The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: '' Omma,'' '' Tetraphalerus'' and '' Beutelius''. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed ...
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Zygadenia
''Zygadenia'' is an extinct genus of archostematan beetles from the Early Jurassic to Early Cretaceous. It is considered to be a senior synonym of ''Notocupes'' by Kirejtshuk (2020), but other researchers suggest to reserve the genus ''Zygadenia'' as a form taxon for isolated elytra that probably belong to the genus ''Notocupes'', while retaining ''Notocupes'' as a valid genus for complete body fossils. Eleven species are included in the genus, after Strelnikova and Yan (2021, 2023). Kirejtshuk (2020) notes that the two Lower Jurassic Australian species ''Z. westraliensis'' and ''Z. martinae'' possibly merit placement in their own separate genus. Jurassic species Hettangian-Toarcian (201.4 Ma to 174.7 Ma) Xiaomeigou Formation, China * ''Z. haixiensis'' Song et al., 2024 Sinemurian-Toarcian (199.5 Ma to 174.1 Ma) ; Cattamarra Coal Measures, Australia * ''Z. martinae'' * ''Z. westraliensis'' Tithonian (152.1 Ma To 145.0 Ma) ; Shar Teeg, Mongolia * ''Z. giebeli'' * ''Z. h ...
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Echinocups
''Echinocups'' is an extinct genus of ommatid beetle. It was created in 2020 to house three species originally assigned to ''Notocupes'', ''E. denticollis'', ''E. neli'' and ''E. ohmkuhnlei'' The genus name refers to the sharp spikes present on the elytra. All three species are known from the Cenomanian aged Burmese amber of Myanmar. The status of ''Echinocups'' as a distinct genus was contested by Li ''et al.'' (2023), who considered the genus ''Echinocups'' to be a junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ... of the genus ''Notocupes''. References Burmese amber Ommatidae Fossil taxa described in 2020 Fossil beetle genera {{paleo-beetle-stub ...
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Rhabdocupes
''Rhabdocupes'' is an extinct genus of beetles in the family Ommatidae, known from the Late Triassic Madygen Formation of Kyrgyzstan, containing the following species: * ''Rhabdocupes baculatus'' Ponomarenko, 1969 * ''Rhabdocupes longus'' Ponomarenko, 1966 * ''Rhabdocupes minor'' Ponomarenko, 1966 ''Rhabdocupes'' may be closely related to the genus ''Notocupes ''Notocupes'' is an extinct genus of medium-sized archostematan beetles from the Mesozoic Era of Eurasia, including over 50 described species. Historically, the genus was classified as a member of the family Ommatidae, but the presence of charact ...''. References Archostemata Carnian genera Fossils of Kyrgyzstan Madygen Formation Fossil beetle genera {{paleo-beetle-stub ...
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Archostemata
The Archostemata are the smallest suborder of beetles, consisting of 50 living species in five families and over 200 described fossil species. They are an ancient lineage with a number of primitive characteristics. Antenna (biology), Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are Neoteny, paedogenic, ''Micromalthus debilis''. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic. The term "Archostemata" is used more broadly by some authors to include both modern archostematans as well as stem-group beetles like "protocoleopterans", which some modern archostematans closely resemble to due to their Plesiomorphy and symplesiomorphy, plesiomorphic morphology. Genetic research suggests that modern archostematans are a monophyletic group. Some genetic studies have recovered archostematans as the sister group of Myxophaga. A 2009 paper argued that th ...
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Cupedidae
The Cupedidae are a small family of beetles, notable for the square pattern of "windows" on their elytra (hard forewings), which give the family their common name of reticulated beetles. The family consists of about 30 species in 9 genera, with a worldwide distribution. Many more extinct species are known, dating as far back as the Triassic. The family Ommatidae is considered a subfamily of Cupedidae by some authors, but ommatids have been found to more closely related to Micromalthidae in genomic analysis studies. These beetles tend to be elongated with a parallel-sided body, ranging in length from , with colors brownish, blackish, or gray. The larvae are wood-borers, typically living in fungus-infested wood, and sometimes found in wood construction. The larvae eat the fungus-infested dead wood or tree roots while the adults are believed to subsist on pollen and plant sap. Males of '' Priacma serrata'' (western North America) are notable for being strongly attracted to commo ...
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Madygen Formation
The Madygen Formation (Russian language, Russian: Madygen Svita) is a Middle Triassic, Middle–Late Triassic (Ladinian–Carnian) geologic formation and lagerstätte in the Batken Region, Batken and Osh Regions of western Kyrgyzstan, with minor outcrops in neighboring Tajikistan and Uzbekistan. The conglomerate (geology), conglomerates, sandstones and mudstones of the thick formation were deposited in terrestrial lacustrine, alluvial, fluvial and river delta, deltaic depositional environment, environments. The formation, extending across the Fergana Valley and Fergana Range, is unique for Central Asia, as it represents one of the few known continental deposits and the Madygen Formation is renowned for the preservation of more than 20,000 fossil insects, making it one of the richest Triassic lagerstätten in the world. Other vertebrate fossils as fish, amphibians, reptiles and synapsids have been recovered from the formation too, as well as minor fossil flora. The lake sediments ...
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Toarcian
The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million years ago) and 174.7 ±0.8 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian Age began with the Toarcian Oceanic Anoxic Event, a major anoxic event associated with marine extinctions and increased global temperatures that sets its fossil faunas apart from the previous Pliensbachian age. It is believed to have ended with a global cooling event known as the Comptum Cooling Event, although whether it represented a worldwide event is controversial. Stratigraphic definitions The Toarcian takes its name from the city of Thouars, just south of Saumur in the Loire Valley of France. The stage was introduced by French palaeontologist Alcide d'Orbigny in 1842, after examining stratum, rock strata of this age in a quar ...
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Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the Oxfordian. Stratigraphic definitions The Callovian Stage was first described by French palaeontologist Alcide d'Orbigny in 1852. Its name derives from the latinized name for Kellaways Bridge, a small hamlet 3 km north-east of Chippenham, Wiltshire, England. The base of the Callovian is defined as the place in the stratigraphic column where the ammonite genus '' Kepplerites'' first appears, which is the base of the biozone of '' Macrocephalites herveyi''. A global reference profile (a GSSP) for the base had in 2009 not yet been assigned. The top of the Callovian (the base of the Oxfordian) is at the first appearance of ammonite species '' Brightia thuouxensis''. Subdivision The Callovian is often subdivided into three substages ...
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Bajocian
In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.9 ±0.8 Ma to around 168.2 ±1.2 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stratigraphic definitions The Bajocian Stage takes its name from the Latin name (Bajocae) of the town of Bayeux, in the region of Normandy in France. The stage was named and introduced in scientific literature by French palaeontologist Alcide d'Orbigny in 1842. The base of the Bajocian stage is defined as the place in the stratigraphic column where fossils of the ammonite genus '' Hyperlioceras'' first appear. A global reference profile (a GSSP) for the base is located at Murtinheira, close to Cabo Mondego in Portugal.The GSSP is described by Pavia & Enay (1997) The top of the Bajocian (the base of the Bathonian) is at the first appearance of ammonite species ''Parkinsonia convergens''. Subdivision The Bajocian is often divided ...
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