The Cupedidae are a small
family
Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
beetle
Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s, notable for the square pattern of "windows" on their
elytra
An elytron (; ; : elytra, ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometime ...
(hard forewings), which give the family their common name of reticulated beetles.
The family consists of about 30 species in 9 genera, with a
worldwide distribution. Many more extinct species are known, dating as far back as the
Triassic
The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
.
[ The family Ommatidae is considered a subfamily of Cupedidae by some authors, but ommatids have been found to more closely related to Micromalthidae in genomic analysis studies.
These beetles tend to be elongated with a parallel-sided body, ranging in length from , with colors brownish, blackish, or gray. The larvae are wood-borers, typically living in ]fungus
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
-infested wood, and sometimes found in wood construction.[ The larvae eat the fungus-infested dead wood or tree roots while the adults are believed to subsist on pollen and plant sap.
Males of '' Priacma serrata'' (western ]North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
) are notable for being strongly attracted to common household bleach
Bleach is the generic name for any chemical product that is used industrially or domestically to remove color from (i.e. to whiten) fabric or fiber (in a process called bleaching) or to disinfect after cleaning. It often refers specifically t ...
. This suggests that compounds in bleach may resemble attractive compounds found by the beetle in nature.[
]
Taxonomy
* '' Adinolepis'' – Australia
* '' Ascioplaga'' – New Caledonia, Australia
* '' Cupes'' – Europe, China, Paleogene
The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
; North America, Recent
* '' Distocupes'' – Australia
* '' Paracupes'' – South America
* '' Priacma'' – Myanmar, Cretaceous; North America, Recent
* '' Prolixocupes'' – North America, South America
* '' Rhipsideigma'' – Madagascar, East Africa
* '' Tenomerga'' – East and Southeast Asia, New Guinea, North America, South Africa
Fossil genera
After Kirejtshuk, Nel and Kirejtshuk (2016).
* †'' Anaglyphites'' – Karabastau Formation
The Karabastau Formation () is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that has been studied since ...
, Kazakhstan, Late Jurassic ( Oxfordian/Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
), Shar-Teg, Mongolia, Late Jurassic (Tithonian
In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
), Hengshan Formation, China, Dzun-Bain Formation, Mongolia, Zaza Formation
The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous. The age of the formation is disputed, and is considered likely to be Valanginian-Hauterivian, or Aptian in age. It comprises sandstones, ...
, Russia, Early Cretaceous (Aptian
The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
)
* †''"Anaglyphites" pluricavus'' – La Pedrera de Rúbies Formation, Spain, Early Cretaceous (Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
)
* †'' Apriacma'' – Yixian Formation
The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
, China, Aptian
* †'' Asimma'' – Madygen Formation
The Madygen Formation (Russian language, Russian: Madygen Svita) is a Middle Triassic, Middle–Late Triassic (Ladinian–Carnian) geologic formation and lagerstätte in the Batken Region, Batken and Osh Regions of western Kyrgyzstan, with minor ...
, Kyrgyzstan, Late Triassic
* †'' Barbaticupes'' – Burmese amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar, Late Cretaceous (Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
)
* †'' Cainomerga'' – Menat Formation
The Menat Formation is a geologic formation in France. It preserves fossils dating back to the Paleocene. It is a maar deposit located on top of an ancient volcano, the extent is very localised with the outcropping area being around 600 by 1000 me ...
, France, Paleocene
The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
* †'' Cretomerga'' – Yixian Formation, China, Aptian
* †'' Cupopsis'' – New Jersey amber
New Jersey Amber, sometimes called Raritan amber, is amber found in the Raritan Formation, Raritan and Magothy Formations of the Mid-Atlantic states, Central Atlantic (Eastern) coast of the United States. It is dated to the Late Cretaceous, Turoni ...
, Late Cretaceous (Turonian
The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
)
* †'' Cupidium'' – Karabastau Formation, Kazakhstan, Oxfordian/Kimmeridgian
* †'' Ensicupes'' – Dalazi Formation, Guyang Formation, China, Aptian
* †'' Furcicupes'' – Yixian Formation, China, Aptian
* †'' Gracilicupes'' – Daohugou, China, Middle Jurassic (Callovian
In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
)
* †'' Kirghizocupes'' – Madygen Formation, Kyrgyzstan, Late Triassic
* †'' Latocupes'' – Yixian Formation, China, Aptian
* †'' Mallecupes'' – Burmese amber, Myanmar, Cenomanian
* †'' Menatops'' – Menat Formation, France, Paleocene
* †'' Mesocupes'' – Karabastau Formation, Kazakhstan, Oxfordian, Las Hoyas, Spain, Barremian
* †'' Miocupes'' – Cypris Formation, Czech Republic, Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
* †''"Platycupes" sogdianus'' – Madygen Formation, Kyrgyzstan, Late Triassic
* †''"Priacma" sanzii'' – Las Hoyas, Spain, Barremian
* †'' Priacmopsis'' – Zaza Formation, Russia, Aptian
* †'' Pterocupes'' – Madygen Formation, Kyrgyzstan, Late Triassic
* †'' Taxopsis'' – Baltic amber
Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...
, Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
'' Notocupes'', traditionally considered a member of Ommatidae, has been suggested to be more closely related to Cupedidae via cladistic analysis.
References
External links
Tree of Life
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Beetle families
{{Cupedidae-stub