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Nothia (plant)
''Nothia'' was a genus of Early Devonian vascular plants whose fossils were found in the Rhynie chert in Scotland. It had branching horizontal underground stems (rhizomes) and leafless aerial stems (axes) bearing lateral and terminal spore-forming organs (sporangia). Its aerial stems were covered with small 'bumps' (emergences), each bearing a stoma. It is one of the best described early land plants. Its classification remains uncertain, although it has been treated as a zosterophyll. There is one species, ''Nothia aphylla''. History of discovery Fossilized remains, including bare stems (axes) and detached spore-forming organs (sporangia), were first described by Kidston and Lang in 1920 from the Rhynie chert of Aberdeenshire, Scotland – rocks which are of Pragian age (). The fragments were considered to be parts of '' Asteroxylon mackiei''. In 1964 Lyon described sporangia belonging to ''Asteroxylon mackiei'' and suggested that Kidston and Lang's specimens were a new specie ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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Protostele
In a vascular plant, the stele is the central part of the root or stem containing the tissues derived from the procambium. These include vascular tissue, in some cases ground tissue (pith) and a pericycle, which, if present, defines the outermost boundary of the stele. Outside the stele lies the endodermis, which is the innermost cell layer of the cortex. The concept of the stele was developed in the late 19th century by French botanists P. E. L. van Tieghem and H. Doultion as a model for understanding the relationship between the shoot and root, and for discussing the evolution of vascular plant morphology. Now, at the beginning of the 21st century, plant molecular biologists are coming to understand the genetics and developmental pathways that govern tissue patterns in the stele. Moreover, physiologists are examining how the anatomy (sizes and shapes) of different steles affect the function of organs. Protostele The earliest vascular plants had stems with a central core of v ...
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Lycophyte
The lycophytes, when broadly circumscribed, are a group of vascular plants that include the clubmosses. They are sometimes placed in a division Lycopodiophyta or Lycophyta or in a subdivision Lycopodiophytina. They are one of the oldest lineages of extant (living) vascular plants; the group contains extinct plants that have been dated from the Silurian (ca. 425 million years ago). Lycophytes were some of the dominating plant species of the Carboniferous period, and included the tree-like Lepidodendrales, some of which grew over in height, although extant lycophytes are relatively small plants. The scientific names and the informal English names used for this group of plants are ambiguous. For example, "Lycopodiophyta" and the shorter "Lycophyta" as well as the informal "lycophyte" may be used to include the extinct zosterophylls or to exclude them. Description Lycophytes reproduce by spores and have alternation of generations in which (like other vascular plants) the sporop ...
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Stem Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown ...
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Paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor exc ...
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Rhyniophyte
The rhyniophytes are a group of extinct early vascular plants that are considered to be similar to the genus ''Rhynia'', found in the Early Devonian (around ). Sources vary in the name and Taxonomic rank#Ranks in botany, rank used for this group, some treating it as the class Rhyniopsida, others as the subdivision Rhyniophytina or the division Rhyniophyta. The first definition of the group, under the name Rhyniophytina, was by Harlan Parker Banks, Banks, since when there have been many redefinitions, including by Banks himself. "As a result, the Rhyniophytina have slowly dissolved into a heterogeneous collection of plants ... the group contains only one species on which all authors agree: the type species ''Rhynia gwynne-vaughanii''". When defined very broadly, the group consists of plants with dichotomously branched, naked aerial axes ("stems") with terminal spore-bearing structures (sporangia). The rhyniophytes are considered to be stem group tracheophytes (vascular plants). Def ...
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Author Citation (botany)
In botanical nomenclature, author citation is the way of citing the person or group of people who validly published a botanical name, i.e. who first published the name while fulfilling the formal requirements as specified by the ''International Code of Nomenclature for algae, fungi, and plants'' (''ICN''). In cases where a species is no longer in its original generic placement (i.e. a new combination of genus and specific epithet), both the authority for the original genus placement and that for the new combination are given (the former in parentheses). In botany, it is customary (though not obligatory) to abbreviate author names according to a recognised List of botanists by author abbreviation, list of standard abbreviations. There are differences between the botanical code and the normal Author citation (zoology), practice in zoology. In zoology, the publication year is given following the author names and the authorship of a new combination is normally omitted. A small number o ...
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Nomen Nudum
In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate description. This makes it a "bare" or "naked" name, which cannot be accepted as it stands. A largely equivalent but much less frequently used term is ''nomen tantum'' ("name only"). Sometimes, "''nomina nuda''" is erroneously considered a synonym for the term "unavailable names". However, not all unavailable names are ''nomina nuda'' which applies to published names, ''i.e.'' any published name that does not fulfill the requirements of Article 12 (if published before 1931) or Article 13 (if published after 1930). In zoology According to the rules of zoological nomenclature a ''nomen nudum'' is unavailable name, unavailable; the glossary of the ''International Code of Zoological Nomenclature'' gives this definition: And ...
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Gametophyte
A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the sexual phase in the life cycle of plants and algae. It develops sex organs that produce gametes, haploid sex cells that participate in fertilization to form a diploid zygote which has a double set of chromosomes. Cell division of the zygote results in a new diploid multicellular organism, the second stage in the life cycle known as the sporophyte. The sporophyte can produce haploid spores by meiosis that on germination produce a new generation of gametophytes. Algae In some multicellular green algae ('' Ulva lactuca'' is one example), red algae and brown algae, sporophytes and gametophytes may be externally indistinguishable (isomorphic). In ''Ulva'', the gametes are isogamous, all of one size, shape and general morphology. Land plant ...
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Geophyte
A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have an underground storage organ are called geophytes in the Raunkiær plant life-form classification system.; reprinted (1977) in ''History of ecology'' series, New York: Arno Press, Storage organs often, but not always, act as perennating organs which enable plants to survive adverse conditions (such as cold, excessive heat, lack of light or drought). Relationship to perennating organ Storage organs may act as perennating organs ('perennating' as in perennial, meaning "through the year", used in the sense of continuing beyond the year and in due course lasting for multiple years). These are used by plants to survive adverse periods in the plant's life-cycle (e.g. caused by cold, excessive heat, lack of light or drought). During these per ...
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Trimerophyte
Trimerophytopsida (or Trimeropsida) is a class of early vascular plants from the Devonian, informally called trimerophytes. It contains genera such as '' Psilophyton''. This group is probably paraphyletic, and is believed to be the ancestral group from which both the ferns and seed plants evolved. Different authors have treated the group at different taxonomic ranks using the names Trimerophyta, Trimerophytophyta, Trimerophytina, Trimerophytophytina and Trimerophytales. Taxonomy At first most of the early land plants other than the bryophytes (i.e. the polysporangiophytes) were placed in a single class Psilophyta, established in 1917 by Kidston and Lang. As additional fossils were discovered and described, it became apparent that the Psilophyta were not a homogeneous group of plants. In 1968 Banks first proposed splitting this taxon into three groups, which he put at the rank of subdivision; he clarified his proposal in 1975. One of the three groups was the Trimerophytina. The su ...
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