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Multituberculates
Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, and reached a peak diversity during the Late Cretaceous and Paleocene. They eventually declined from the mid-Paleocene onwards, disappearing from the known fossil record in the late Eocene. They are the most diverse order of Mesozoic mammals with more than 200 species known, ranging from mouse-sized to beaver-sized. These species occupied a diversity of ecological niches, ranging from burrow-dwelling to squirrel-like arborealism to jerboa-like hoppers. Multituberculates are usually placed as crown mammals outside either of the two main groups of living mammals, Theria — placentals and marsupials — and Monotremata,Agustí-Antón 2002, pp 3-4 but usually as closer to Theria than to monotremes. They are considered to be closely rel ...
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Catopsbaatar
''Catopsbaatar'' is a genus of multituberculate, an extinct Order (biology), order of rodent-like mammals. It lived in what is now Mongolia during the late Campanian age of the Late Cretaceous epoch (geology), epoch, about 72 million years ago. The first fossils were collected in the early 1970s, and the animal was named as a new species of the genus ''Djadochtatherium'' in 1974, ''D. catopsaloides''. The Specific name (zoology), specific name refers to the animal's similarity to the genus ''Catopsalis''. The species was moved to the genus ''Catopsalis'' in 1979, and received its own genus (''Catopsbaatar'', Greek and Mongolian for 'visible hero') in 1994. Five skulls, one molar, and one skeleton with a skull are known; the last is the genus' most complete specimen. ''Catopsbaatar'' was a member of the family Djadochtatheriidae. The skull of ''Catopsbaatar'' was up to long and, as in other multituberculates, proportionally large. The external appearance of these animals' h ...
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Gondwanatheria
Gondwanatheria is an extinct group of mammaliaforms that lived in parts of Gondwana, including Madagascar, India, South America, Africa, and Antarctica during the Upper Cretaceous through the Miocene (and possibly much earlier, if '' Allostaffia'' is a member of this group). Until recently, they were known only from fragmentary remains. They are generally considered to be closely related to the multituberculates and likely the euharamiyidians, well known from the Northern Hemisphere, with which they form the clade Allotheria. Classification For several decades the affinities of the group were not clear, being first interpreted as early xenarthrans, or "toothless" mammals similar to the modern anteater. A variety of studies have placed them as allotheres related to multituberculates, possibly even true multituberculates, closer to cimolodonts than "plagiaulacidans" are. However, a more recent study recovered them as nested among haramiyidans, rendering them as non-mammalian ...
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Plagiaulacoid
A plagiaulacoid is a type of blade-like, most often serrated, tooth present in various mammal groups, usually a premolar. Among modern species it is present chiefly on diprotodontian marsupials (specifically, Potoroidae, ''Bettongia'' and '' Burramys''), which have both the upper and lower first premolars converted into serrated blades. However, various other extinct groups also possessed plagiaulacoids. These would be multituberculates, some "Plesiadapiformes" such as ''Carpolestes'' and various metatherians such as '' Epidolops'' and various early diprotodontians. In many of these only a lower premolar became converted into a blade, while the upper premolars showed less specialisation. The various independent development of these teeth is considered a good example of convergent evolution. Multituberculates in particular probably own the most specialised of all plagiaulacoids. In early taxa, all lower premolars became plagiaulacoids, forming a "saw"-like arrangement. However, in Ci ...
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Zofia Kielan-Jaworowska
Zofia Emilia Kielan-Jaworowska (25 April 1925 – 13 March 2015) was a Polish paleobiologist. In the mid-1960s, she led a series of Polish-Mongolian paleontological expeditions to the Gobi Desert. She was the first woman to serve on the executive committee of the International Union of Geological Sciences. The most notable dinosaur species she discovered include: '' Deinocheirus'' and '' Gallimimus'' while '' Kielanodon'' and '' Zofiabaatar'' were named in her honour. In her obituary in ''Nature'', Richard L. Cifelli wrote that "Much of what we know about the origin and early evolution of mammals stems, directly or indirectly, from er work. Early life and education Zofia Kielan-Jaworowska was born in Sokołów Podlaski, Poland, on April 25, 1925. In 1928, her father, Franciszek Kielan, was offered a job for the Association of Agriculture and Trade Cooperatives in Warsaw, to which her family moved for five years. Zofia and family returned to Warsaw in 1934 and lived in Żol ...
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Mesozoic
The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian reptiles such as the dinosaurs, and of Gymnosperm, gymnosperms such as cycads, ginkgoaceae and Araucariaceae, araucarian conifers; a hot Greenhouse and icehouse earth, greenhouse climate; and the tectonic break-up of Pangaea. The Mesozoic is the middle of the three eras since Cambrian explosion, complex life evolved: the Paleozoic, the Mesozoic, and the Cenozoic. The era began in the wake of the Permian–Triassic extinction event, the largest mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event, another mass extinction whose victims included the non-avian dinosaurs, Pterosaur, pterosaurs, Mosasaur, mosasaurs, and Plesiosaur, plesiosaurs. The Mesozoic was a time of significant tectonic, climatic, an ...
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Euharamiyida
Euharamiyida also known as Eleutherodontida, is clade of early mammals or mammal-like cynodonts from the Middle Jurassic to Early Cretaceous of Eurasia and possibly North America. The group is sometimes considered a sister group to Multituberculata, or part of an earlier divergence within the synapsid line. It is disputed whether or not they are related to the haramiyids from the Late Triassic, such as '' Haramiyavia''. The morphology of their teeth indicates that they were herbivorous or omnivorous. Some members of the group are known to be arboreal, including gliding forms similar to modern flying squirrels or colugos. Evolution The position of euharamyidans is contested. They are either considered crown group mammals as members of Allotheria, related to multituberculates, or stem-group mammals within Mammaliaformes. The position is often dependent on the relationships of euharamiyids to the Late Triassic haramiyids such as '' Haramiyavia'' and ''Thomasia''. In some studi ...
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Cimolodonta
Cimolodonta is a clade of Multituberculata, multituberculate mammals that lived from the Cretaceous to the Eocene. They probably lived something of a rodent-like existence until their ecological niche was assumed by true rodents. The more basal multituberculates are found in a different suborder, "Plagiaulacida", a Paraphyly, paraphyletic group containing all non cimolodontan multituberculates. Cimolodonta is apparently a natural (monophyletic) suborder. Remains have been identified from across the Northern Hemisphere. They first appeared during the Aptian extinction, Aptian, and completely replaced the more primitive plagiaulacidans by the early Late Cretaceous. The taxon is recognized as the informal Paracimexomys group and the superfamilies Djadochtatherioidea, Taeniolabidoidea, and Ptilodontoidea. Additionally, and of uncertain affinities, are the families Cimolomyidae, Boffius, Boffiidae, Eucosmodontidae, Kogaionidae, Microcosmodontidae and the two genera ''Uzbekbaatar'' and ...
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Mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three Evolution of mammalian auditory ossicles, middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors Genetic divergence, diverged in the Carboniferous Period over 300 million years ago. Around 6,640 Neontology#Extant taxon, extant species of mammals have been described and divided into 27 Order (biology), orders. The study of mammals is called mammalogy. The largest orders of mammals, by number of species, are the rodents, bats, and eulipotyphlans (including hedgehogs, Mole (animal), moles and shrews). The next three are the primates (including humans, monkeys and lemurs), the Artiodactyl, even-toed ungulates (including pigs, camels, and whales), and the Carnivora (including Felidae, ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ninth and longest geological period of the entire Phanerozoic. The name is derived from the Latin , 'chalk', which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation . The Cretaceous was a period with a relatively warm climate, resulting in high Sea level#Local and eustatic, eustatic sea levels that created numerous shallow Inland sea (geology), inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was largely ice-free, although there is some evidence of brief periods of glaciation during the cooler first half, and forests extended to the poles. Many of the dominant taxonomic gr ...
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Placental
Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus (biology), fetus is carried in the uterus of its mother to a relatively late stage of development. The name is something of a misnomer, considering that marsupials also nourish their fetuses via a placenta, though for a relatively briefer period, giving birth to less-developed young, which are then nurtured for a period inside the mother's pouch (marsupial), pouch. Placentalia represents the only living group within Eutheria, which contains all mammals that are more closely related to placentals than they are to marsupials. Anatomical features Placental mammals are anatomically distinguished from other mammals by: * a sufficiently wide opening at the bottom of the pelvis to allow the birt ...
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Allotheria
Allotheria (meaning "other beasts", from the Ancient Greek language, Greek , '–other and , '–wild animal) is an extinct clade of mammals known from the Mesozoic and early Cenozoic. Shared characteristics of the group are the presence of lower Molar (tooth), molariform Tooth (animal), teeth equipped with longitudinal rows of cusps and enlarged incisors. Typically, the canine teeth are also lost. Allotheria includes Multituberculata, Gondwanatheria (which may be part of Multituberculata, as the sister group to Cimolodonta), and probably Haramiyida, (sometimes only including Euharamiyida) although some studies have recovered haramiyidans to be basal mammaliaforms unrelated to multituberculates. Allotherians are often placed as crown group mammals, more closely related to living marsupials and placentals (Theria) than to monotremes or Eutriconodonta, eutriconodonts, though some studies place the entirety of Allotheria outside of crown Mammalia. References Further reading

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Taeniolabis NT Small
''Taeniolabis'' ("banded incisor") is a genus of extinct multituberculate mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ... from the Paleocene of North America. Description ''Taeniolabis'' is a member of the Taeniolabidoidea, a superfamily of multituberculates that are known for their highly derived teeth, and a short wide snout with a blocky head. The teeth modifications were likely an adaptation for herbivory that may have resulted from rapid diversification of angiosperms at the very end of the Cretaceous, which would thus have created opportunities for novel specialization in herbivores. ''T. taoensis'' possessed an elongate, gently curved cochlear canal and an enlarged vestibule. It is the largest known multituberculate, as well as the largest allotherian mammal, ...
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