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Manis Lydekkeri
''Manis'' (" spirit") is a genus of South Asian and East Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...n pangolins, the Asiatic pangolins, from subfamily Maninae, within family Manidae. Taxonomy * Subfamily: Maninae (Asiatic pangolins) ** Genus: ''Manis'' (Asiatic pangolins) *** '' Manis crassicaudata'' (Indian pangolin) *** '' Manis pentadactyla'' (Chinese pangolin) *** '' Manis aurita'' (= ''Manis indoburmanica'')Wangmo, L. K., Ghosh, A., Dolker, S., Joshi, B. D., Sharma, L. K., & Thakur, M. (2025)Indo-Burmese pangolin (''Manis indoburmanica''): a novel phylogenetic species of pangolin evolved in Asia.Mammalian Biology, 1-8. *** '' Manis mysteria'' (Cryptic pangolin)https://www.sci.news/biology/manis-mysteria-12316.html *** †'' Manis hungarica'' *** †'' Manis lyd ...
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Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene followed the Oligocene and preceded the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by distinct global events but by regionally defined transitions from the warmer Oligocene to the cooler Pliocene Epoch. During the Early Miocene, Afro-Arabia collided with Eurasia, severing the connection between the Mediterranean and Indian Oceans, and allowing the interchange of fauna between Eurasia and Africa, including the dispersal of proboscideans and Ape, hominoids into Eurasia. During the late Miocene, the conn ...
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Manidae
Manidae (" spirits") is the only extant family of pangolins. This family comprises three genera ('' Manis'' from subfamily Maninae, '' Phataginus'' from subfamily Phatagininae, and '' Smutsia'' from subfamily Smutsiinae), as well as the extinct Fayum pangolin. Classification and phylogeny History of classification All species of living pangolin had been assigned to the genus ''Manis'' until the late 2000s, when research prompted the splitting of extant pangolins into three genera: '' Manis'', '' Phataginus'', and '' Smutsia''. Taxonomy * Family: Manidae (pangolins) ** Subfamily: Maninae (Gray, 1821) (Asian pangolins) *** Genus: '' Manis'' (Linnaeus, 1758) (Asian pangolin) **** (unranked): northern Asian clade ***** '' Manis pentadactyla'' (Linnaeus, 1758) (Chinese pangolin) ***** †'' Manis hungarica'' (Kormos, 1934) **** (unranked): southern Asian clade ***** '' Manis crassicaudata'' (Gray, 1827) (Indian pangolin) ***** ''Manis'' sp. (''Scale_H4'' & ''Scale_H8'')Jingyan ...
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Phataginus
African tree pangolin (''Phataginus'') is a genus of African pangolins from subfamily small African pangolins (Phatagininae), within family Manidae. Its members are the more arboreal of the African pangolins. From 2010 to 2019, at least 895,000 pangolins from the genus ''Phataginus'' were illegally trafficked. The animal is hunted and poached for its scales and meat, and is often used in making traditional medicine in places such as China and Vietnam. Attempts to protect these mammals from trafficking and extinction are ongoing; unfortunately their slow reproduction rate stymies population recovery. Currently the tree pangolin is listed as vulnerable. All pangolin species have been listed as vulnerable, endangered, or critically endangered. Etymology Constantine Rafinesque (1821) formed the Neo-Latin generic name ''Phataginus'' from the French term ''phatagin'', adopted by Count Buffon (1763) after the reported local name ''phatagin'' or ''phatagen'' used in the East Indies. ...
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Necromanis
''Necromanis'' ("extinct pangolin") is an extinct genus of pangolin from superfamily Manoidea. It lived from the middle Oligocene to middle Miocene in Europe. It was originally placed within family Manidae, but was eventually removed from it as more fossil pholidotids from outside that family were found and studied more extensively (i.e., with the discovery and study of ''Eomanis'' and ''Patriomanis'').Gaudin, Timothy J., Robert J. Emry, and Brandon Pogue. "A new genus and species of pangolin (Mammalia, Pholidota) from the late Eocene of Inner Mongolia, China." Journal of Vertebrate Paleontology 26.1 (2006): 146-159. Currently, ''Necromanis'' is placed as ''incertae sedis'' within the pholidotid superfamily Manoidea, together with the families Manidae and Patriomanidae. ''N. quercyi'' was originally placed within ''Teutomanis'' by Ameghino in 1905, but was later subsumed into ''Necromanis''. A new fossil humerus attributed to ''N. franconica'' from Quercy, France lead researchers ...
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Patriomanidae
Patriomanidae ("fathers of pangolins") is an extinct family of pangolins from superfamily Manoidea that includes two extinct genera '' Patriomanis'' and '' Cryptomanis''. Their fossils are found in Asia and North America. Taxonomy * Family: †Patriomanidae ** Genus: †'' Cryptomanis'' *** †''Cryptomanis gobiensis'' ** Genus: †'' Patriomanis'' *** †''Patriomanis americana'' Phylogeny Phylogenetic position of family Patriomanidae within superfamily Manoidea. See also * Mammal classification * Manoidea Manoidea ("Manes, spirits") is a superfamily of pangolins from suborder Eupholidota that includes extant family Manidae, extinct family Patriomanidae and extinct genus ''Necromanis''. Taxonomy * Superfamily: Manoidea (pangolins) ** Family: Ma ... References Prehistoric pangolins Prehistoric mammal families {{Paleo-mammal-stub ...
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Manoidea
Manoidea ("Manes, spirits") is a superfamily of pangolins from suborder Eupholidota that includes extant family Manidae, extinct family Patriomanidae and extinct genus ''Necromanis''. Taxonomy * Superfamily: Manoidea (pangolins) ** Family: Manidae (pangolins) ** Family: Extinction, †Patriomanidae ** ''Incertae sedis'' *** Genus: †''Necromanis'' Phylogeny Phylogenetic position of superfamily Manoidea within suborder Eupholidota. See also * Mammal classification * Eupholidota References

Pangolins Mammal superfamilies {{mammal-stub ...
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Eomanis
''Eomanis'' ("dawn pangolin") is the earliest known true (and scaled) pangolin from extinct family Eomanidae (and extinct superfamily Eomanoidea) within suborder Eupholidota. It lived during the Eocene in Europe. ''Eomanis'' fossils found in the Messel Pit in Germany are very similar in size and anatomy to living pangolins of the genus ''Manis'', indicating that pangolins have remained largely unchanged in morphology and behavior for 50 million years. However, unlike modern pangolins, its tail and legs did not bear scales. According to the stomach contents of the excellently preserved Messel specimens, ''Eomanis’'' diet consisted of both insects and plants. Another early mammal discovered in the Messel Pit that was similar in anatomy and likely also had an insectivorous diet was '' Eurotamandua''. Despite its name, ''Eurotamandua'' was almost certainly not a xenarthran because it lacked the characteristic xenarthran joints present in all living and extinct xenarthrans. In add ...
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Eupholidota
Eupholidota ("true pangolins") is a suborder of pangolins that includes two superfamilies: extant Manoidea and extinct Eomanoidea. Taxonomy * Suborder: Eupholidota (true pangolins) ** Superfamily: Manoidea (pangolins) *** Family: Manidae (pangolins) *** Family: †Patriomanidae *** ''Incertae sedis'' **** Genus: †''Necromanis'' ** Superfamily: † Eomanoidea *** Family: † Eomanidae Phylogeny Phylogenetic position of suborder Eupholidota within order Pholidota. See also * Mammal classification * Pangolin Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (). The one extant family, the Manidae, has three genera: '' Manis'', '' Phataginus'', and '' Smutsia''. ''Manis'' comprises four species found in Asia, while ' ... References Pangolins {{mammal-stub ...
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Eurotamandua
''Eurotamandua'' ("european ''Tamandua''") is an extinct genus of mammal from extinct family Eurotamanduidae that lived during the middle Eocene. A single fossil is known, coming from the Messel Pit in southwestern Germany. ''Eurotamandua'' was about long. Most palaeontologists now classify ''Eurotamandua'' as a pangolin. When its fossils were first discovered, ''Eurotamandua'' was originally thought to be an anteater, as it lacked the characteristic fused-hair scales of other pangolins. ''Eurotamandua'' placement within the pangolins was made primarily because of a lack of the characteristic "xenarthran" joints found in all xenarthrans, including tamanduas. There is still much ambiguity in the taxonomy of all mammals prior to the Eocene, so there is the possibility that ''Eurotamandua'' was a primitive xenarthran. However, this is highly unlikely because all known fossil evidence indicates that xenarthrans existed exclusively in South America from the beginning of the Cenozoic ...
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Euromanis
''Euromanis'' ("european pangolin") is one of the earliest known pangolin genera. It lived during the middle Eocene in Europe. ''Euromanis'' fossils found in the Messel Pit in Germany. Unlike modern pangolins, it did not bear scales on its body. Phylogeny Phylogenetic position of genus ''Euromanis'' within order Pholidota. See also * Mammal classification * Pangolin Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (). The one extant family, the Manidae, has three genera: '' Manis'', '' Phataginus'', and '' Smutsia''. ''Manis'' comprises four species found in Asia, while ' ... References Prehistoric pangolins Eocene mammals Myrmecophagous mammals Prehistoric placental genera Cenozoic mammals of Europe Fossil taxa described in 1994 {{paleo-mammal-stub ...
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Pangolin
Pangolins, sometimes known as scaly anteaters, are mammals of the order Pholidota (). The one extant family, the Manidae, has three genera: '' Manis'', '' Phataginus'', and '' Smutsia''. ''Manis'' comprises four species found in Asia, while ''Phataginus'' and ''Smutsia'' include two species each, all found in sub-Saharan Africa. These species range in size from . Several extinct pangolin species are also known. In September 2023, nine species were reported. Pangolins have large, protective keratin scales, similar in material to fingernails and toenails, covering their skin; they are the only known mammals with this feature. Depending on the species, they live in hollow trees or burrows. Pangolins are nocturnal, and their diet consists of mainly ants and termites, which they capture using their long tongues. They tend to be solitary animals, meeting only to mate and produce a litter of one to three offspring, which they raise for about two years. Pangolins superficially resemb ...
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